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<document id="7727C6FDF1F7FC19573DCED6C3D463C2" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850379EF691C5579BDFC7E68C2" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Berlandina artaxerxes Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="AA010096-605E-4BFD-A7D7-8797A2B89B34" docUuidSource="ZooBank" docVersion="8" lastPageNumber="76" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="74" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="48274396CB3691E25D784DD58222B9C8">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="0D6C6B383518851852E53283724DBDB0">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="BC9183343E0219B68C101DFC9E933C42" type="LSID">A21C0B82-E2D9-4402-9408-D6160843DAF4</mods:nameIdentifier>
<mods:affiliation id="598C44C9BC552A46C767FBB7EF5A45E9">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
<mods:nameIdentifier id="009995478DE94B2D56E3E75B41DE193B" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C4F50CC62BD7B22ACD7F09E77BD5777C">Chatzaki, Maria</mods:namePart>
<mods:nameIdentifier id="15B86CD5ACC633D32359BB07FC7D3B0F" type="LSID">2FBB48ED-84FE-4386-8F47-C958E9F8A2A1</mods:nameIdentifier>
<mods:nameIdentifier id="7386C6325C53821DB8E44043A1F5AC4C" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6590338029A77250D325FE1D6B4DE6CC">Esyunin, Sergei L.</mods:namePart>
<mods:nameIdentifier id="D69EDAAA363D03514A55571CA5E0CCB9" type="LSID">2B591857-5FAB-4B7D-B589-6C4977A3A493</mods:nameIdentifier>
<mods:affiliation id="3DBB9C01A5950AABF0D8A9451D445B4D">Sergei.</mods:affiliation>
<mods:nameIdentifier id="579731E8D13850D493005A064EDE0A45" type="email">Sergei.Esyunin@psu.ru</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="FA7B11E7E322D916CAE15D0BEF266463">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="C3146043FFA44A70BFA7DBCF76F6A9F2" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="789BCF09C57BE9E601EE71DB87761B9D">European Journal of Taxonomy</mods:title>
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<mods:date id="EBF6EE1D97154C5FBCC6483C36845274">2021</mods:date>
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<mods:number id="6B0669449264F8F6691FCB4576F0A466">2021-06-04</mods:number>
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<treatment id="03F987850379EF691C5579BDFC7E68C2" ID-DOI="http://doi.org/10.5281/zenodo.4906461" ID-GBIF-Taxon="182597831" ID-Zenodo-Dep="4906461" LSID="urn:lsid:zoobank.org:act:AA010096-605E-4BFD-A7D7-8797A2B89B34" httpUri="http://treatment.plazi.org/id/03F987850379EF691C5579BDFC7E68C2" lastPageId="8" lastPageNumber="76" pageId="6" pageNumber="74">
<subSubSection id="C34A65180379EF671C5579BDFC126BA8" box="[609,979,1742,1769]" pageId="6" pageNumber="74" type="nomenclature">
<paragraph id="8BEF36930379EF671C5579BDFC126BA8" blockId="6.[363,1224,1742,1840]" box="[609,979,1742,1769]" pageId="6" pageNumber="74">
<heading id="D0A781FF0379EF671C5579BDFC126BA8" box="[609,979,1742,1769]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="74" reason="2">
<taxonomicName id="4C504D100379EF671C5579BDFCB06BA8" ID-CoL="98YVX" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[609,881,1742,1769]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="74" phylum="Arthropoda" rank="species" species="artaxerxes" status="sp. nov.">
<emphasis id="B924EA810379EF671C5579BDFCB06BA8" bold="true" box="[609,881,1742,1769]" italics="true" pageId="6" pageNumber="74">Berlandina artaxerxes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0379EF671D4C79BCFC126BA8" box="[888,979,1743,1769]" pageId="6" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FFC13A910379EF671F5F7981FB096A4C" box="[363,1224,1778,1805]" pageId="6" pageNumber="74">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BEF36930379EF671CEF7865FC996A71" blockId="6.[363,1224,1742,1840]" box="[731,856,1814,1840]" pageId="6" pageNumber="74">
<figureCitation id="136B2A160379EF671CEF7865FC996A71" box="[731,856,1814,1840]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[221,1385,525,1854]" captionTargetId="figure-110@7.[202,1385,525,1854]" captionTargetPageId="7" captionText="Fig. 3. A–D. Berlandina artaxerxes sp. nov., holotype, ♂ (SMF). A. Habitus, dorsal view. B–D. Palp, ventral, proventral and retrolateral views. –E–G. Berlandina mesopotamica Al-Khazali,2020, ♀ (MHNG). E. Habitus, dorsal view. F–G. Epigyne, ventral and dorsal views. Abbreviations: Ao = apical outgrowth; Eb = embolar base; Po = pointed process. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906013" httpUri="https://zenodo.org/record/4906013/files/figure.png" pageId="6" pageNumber="74">Fig. 3A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A65180379EF661E897824FE8A6C95" lastPageId="7" lastPageNumber="75" pageId="6" pageNumber="74" type="diagnosis">
<paragraph id="8BEF36930379EF671E897824FEF76A30" blockId="6.[189,310,1879,1905]" box="[189,310,1879,1905]" pageId="6" pageNumber="74">
<heading id="D0A781FF0379EF671E897824FEF76A30" bold="true" box="[189,310,1879,1905]" fontSize="11" level="3" pageId="6" pageNumber="74" reason="3">
<emphasis id="B924EA810379EF671E897824FEF76A30" bold="true" box="[189,310,1879,1905]" pageId="6" pageNumber="74">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930379EF671E8978F4FAB76AA8" blockId="6.[189,1399,1927,2025]" pageId="6" pageNumber="74">
The new species is most similar to
<taxonomicName id="4C504D100379EF671C6178FBFB836AE0" authority="(O. Pickard-Cambridge, 1874)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1874" box="[597,1090,1927,1953]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="74" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B924EA810379EF671C6178FBFD146AE0" box="[597,725,1928,1953]" italics="true" pageId="6" pageNumber="74">B. venatrix</emphasis>
(O. Pickard-Cambridge, 1874)
</taxonomicName>
and
<taxonomicName id="4C504D100379EF671A4E78FBFDCE6A84" authority="Marusik, Fomichev &amp; Omelko, 2014" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="74" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="B924EA810379EF671A4E78FBFB4D6AE0" box="[1146,1164,1928,1953]" italics="true" pageId="6" pageNumber="74">B</emphasis>
.
<emphasis id="B924EA810379EF671AAF78F4FAC66AE0" box="[1179,1287,1927,1953]" italics="true" pageId="6" pageNumber="74">koponeni</emphasis>
<bibRefCitation id="EFC14B620379EF671B3978F4FDCE6A84" author="Marusik Y. M. &amp; Fomichev A. A. &amp; Omelko M. M." pageId="6" pageNumber="74" pagination="187 - 213" refId="ref17777" refString="Marusik Y. M., Fomichev A. A. &amp; Omelko M. M. 2014. A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa 3827 (2): 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014">Marusik, Fomichev &amp; Omelko, 2014</bibRefCitation>
</taxonomicName>
by having a broad embolar base (
<emphasis id="B924EA810379EF671D8C78D9FC186A85" box="[952,985,1962,1988]" italics="true" pageId="6" pageNumber="74">Eb</emphasis>
) with a similar sclerotized apical outgrowth (
<emphasis id="B924EA810379EF671F7C78BCFEA86AA8" box="[328,361,1999,2025]" italics="true" pageId="6" pageNumber="74">Ao</emphasis>
) parallel to a bent, long embolus and a basal process (
<emphasis id="B924EA810379EF671DC378BCFBD96AA9" box="[1015,1048,1999,2024]" italics="true" pageId="6" pageNumber="74">Po</emphasis>
). In the new species, the two
</paragraph>
<paragraph id="8BEF36930378EF661E897E7AFE8A6C95" blockId="7.[189,1399,265,468]" pageId="7" pageNumber="75">
elements of the embolar base are pointed (vs shallowly divided with the process small and curved in
<taxonomicName id="4C504D100378EF661E8B7E5EFE866C07" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" box="[191,327,300,327]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="B924EA810378EF661E8B7E5EFF0E6C07" box="[191,207,301,326]" italics="true" pageId="7" pageNumber="75">B</emphasis>
.
<emphasis id="B924EA810378EF661EE87E5FFE866C07" box="[220,327,300,326]" italics="true" pageId="7" pageNumber="75">koponeni</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFC14B620378EF661FB57E5FFDA76C07" author="Marusik Y. M. &amp; Fomichev A. A. &amp; Omelko M. M." box="[385,614,300,326]" pageId="7" pageNumber="75" pagination="187 - 213" refId="ref17777" refString="Marusik Y. M., Fomichev A. A. &amp; Omelko M. M. 2014. A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa 3827 (2): 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014">
Marusik
<emphasis id="B924EA810378EF661FD37E5EFDE16C07" box="[487,544,300,326]" italics="true" pageId="7" pageNumber="75">et al.</emphasis>
2014
</bibRefCitation>
: figs 32, 34, and the basal process bearing a very pointed sclerotized tooth in
<taxonomicName id="4C504D100378EF661F117E23FE696C28" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1874" box="[293,424,336,361]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B924EA810378EF661F117E23FE696C28" box="[293,424,336,361]" italics="true" pageId="7" pageNumber="75">B. venatrix</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFC14B620378EF661FDD7E23FDAE6C2B" author="Levy G." box="[489,623,335,362]" pageId="7" pageNumber="75" pagination="919 - 981" refId="ref17523" refString="Levy G. 1995. Revision of the spider subfamily Gnaphosinae in Israel (Araneae: Gnaphosidae). Journal of Natural History 29 (4): 919 - 981. https: // doi. org / 10.1080 / 00222939500770351" type="journal article" year="1995">Levy 1995</bibRefCitation>
: fig. 57, not present in
<taxonomicName id="4C504D100378EF661DBD7E23FBE86C28" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[905,1065,336,361]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="artaxerxes" status="sp. nov.">
<emphasis id="B924EA810378EF661DBD7E23FBE86C28" box="[905,1065,336,361]" italics="true" pageId="7" pageNumber="75">B. artaxerxes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0378EF661A067E23FB556C2B" box="[1074,1172,336,362]" pageId="7" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
). Also, the median apophysis in
<taxonomicName id="4C504D100378EF661F6F7E00FE386CCD" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[347,505,371,396]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="artaxerxes" status="sp. nov.">
<emphasis id="B924EA810378EF661F6F7E00FE386CCD" box="[347,505,371,396]" italics="true" pageId="7" pageNumber="75">B. artaxerxes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0378EF661C347E00FD9D6CCC" box="[512,604,371,397]" pageId="7" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
is bent prolaterally and located under the apical outgrowth’s curve (vs straight and longer, reaching the height of the embolus in
<taxonomicName id="4C504D100378EF661D9B7EE4FBFF6CF1" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" box="[943,1086,406,432]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="koponeni">
<emphasis id="B924EA810378EF661D9B7EE4FBFF6CF1" box="[943,1086,406,432]" italics="true" pageId="7" pageNumber="75">B. koponeni</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFC14B620378EF661A4B7EE5FAB36CF1" author="Marusik Y. M. &amp; Fomichev A. A. &amp; Omelko M. M." box="[1151,1394,406,432]" pageId="7" pageNumber="75" pagination="187 - 213" refId="ref17777" refString="Marusik Y. M., Fomichev A. A. &amp; Omelko M. M. 2014. A survey of East Palaearctic Gnaphosidae (Araneae). 1. On the Berlandina Dalmas, 1922 (Gnaphosinae) from Mongolia and adjacent regions. Zootaxa 3827 (2): 187 - 213. https: // doi. org / 10.11646 / zootaxa. 3827.2.4" type="journal article" year="2014">
Marusik
<emphasis id="B924EA810378EF661AD87EE4FAE86CF1" box="[1260,1321,406,432]" italics="true" pageId="7" pageNumber="75">et al.</emphasis>
2014
</bibRefCitation>
: figs 34–35).
</paragraph>
</subSubSection>
<caption id="DF2F661B0378EF661E897817FB726AA9" ID-DOI="http://doi.org/10.5281/zenodo.4906013" ID-Zenodo-Dep="4906013" httpUri="https://zenodo.org/record/4906013/files/figure.png" pageId="7" pageNumber="75" startId="7.[189,232,1892,1918]" targetBox="[221,1385,525,1854]" targetPageId="7">
<paragraph id="8BEF36930378EF661E897817FB726AA9" blockId="7.[189,1399,1892,2025]" pageId="7" pageNumber="75">
<emphasis id="B924EA810378EF661E897817FE8D6A3F" bold="true" box="[189,332,1892,1918]" pageId="7" pageNumber="75">Fig. 3. A–D</emphasis>
.
<taxonomicName id="4C504D100378EF661F687817FDA56A3F" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[348,612,1892,1918]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="artaxerxes" status="sp. nov.">
<emphasis id="B924EA810378EF661F687817FDA56A3F" box="[348,612,1892,1918]" italics="true" pageId="7" pageNumber="75">Berlandina artaxerxes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0378EF661C5A7816FD0A6A3E" box="[622,715,1893,1919]" pageId="7" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (SMF).
<emphasis id="B924EA810378EF661DF87817FC206A3F" bold="true" box="[972,993,1892,1918]" pageId="7" pageNumber="75">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B924EA810378EF661AC17816FAEA6A3E" bold="true" box="[1269,1323,1893,1919]" pageId="7" pageNumber="75">B–D</emphasis>
. Palp, ventral, proventral and retrolateral views. –
<emphasis id="B924EA810378EF661C9078F4FD1C6AE0" bold="true" box="[676,733,1927,1953]" pageId="7" pageNumber="75">E–G</emphasis>
.
<taxonomicName id="4C504D100378EF661CDE78FBFB1D6AE3" authority="Al-Khazali, 2020" authorityName="Al-Khazali" authorityYear="2020" box="[746,1244,1928,1954]" class="Arachnida" family="Gnaphosidae" genus="Berlandina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="mesopotamica">
<emphasis id="B924EA810378EF661CDE78FBFBD46AE0" box="[746,1045,1928,1954]" italics="true" pageId="7" pageNumber="75">Berlandina mesopotamica</emphasis>
<bibRefCitation id="EFC14B620378EF661A2D78FBFB1D6AE3" author="Al-Khazali A. M." box="[1049,1244,1928,1954]" pageId="7" pageNumber="75" pagination="444 - 446" refId="ref16885" refString="Al-Khazali A. M. 2020. Berlandina mesopotamica sp. nov. (Araneae: Gnaphosidae): new record of the genus in Iraq. Arachnology 18 (5): 444 - 446. https: // doi. org / 10.13156 / arac. 2020.18.5.444" type="journal article" year="2020">Al-Khazali,2020</bibRefCitation>
</taxonomicName>
, ♀ (MHNG).
<emphasis id="B924EA810378EF661E8978D8FF106A84" bold="true" box="[189,209,1963,1989]" pageId="7" pageNumber="75">E</emphasis>
. Habitus, dorsal view.
<emphasis id="B924EA810378EF661FED78D8FDD06A84" bold="true" box="[473,529,1963,1989]" pageId="7" pageNumber="75">F–G</emphasis>
. Epigyne, ventral and dorsal views. Abbreviations:
<emphasis id="B924EA810378EF661A5E78D8FB4A6A84" box="[1130,1163,1963,1989]" italics="true" pageId="7" pageNumber="75">Ao</emphasis>
= apical outgrowth;
<emphasis id="B924EA810378EF661E8978BDFF1F6AA9" box="[189,222,1998,2024]" italics="true" pageId="7" pageNumber="75">Eb</emphasis>
= embolar base;
<emphasis id="B924EA810378EF661F9C78BCFE086AA9" box="[424,457,1999,2024]" italics="true" pageId="7" pageNumber="75">Po</emphasis>
= pointed process. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<subSubSection id="C34A65180377EF691E897E7AFDF96C39" pageId="8" pageNumber="76" type="etymology">
<paragraph id="8BEF36930377EF691E897E7AFE856C62" blockId="8.[189,324,265,291]" box="[189,324,265,291]" pageId="8" pageNumber="76">
<heading id="D0A781FF0377EF691E897E7AFE856C62" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="8" pageNumber="76" reason="3">
<emphasis id="B924EA810377EF691E897E7AFE856C62" bold="true" box="[189,324,265,291]" pageId="8" pageNumber="76">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930377EF691E897E4AFDF96C39" blockId="8.[189,1399,313,376]" pageId="8" pageNumber="76">The specific epithet is a noun in apposition and refers to Ardashir I (equivalent to Greek Artaxérxēs), the founder of the Sasanian Empire.</paragraph>
</subSubSection>
<subSubSection id="C34A65180377EF691E897ED2FE746F74" pageId="8" pageNumber="76" type="materials_examined">
<paragraph id="8BEF36930377EF691E897ED2FEAD6CFA" blockId="8.[189,364,417,443]" box="[189,364,417,443]" pageId="8" pageNumber="76">
<heading id="D0A781FF0377EF691E897ED2FEAD6CFA" bold="true" box="[189,364,417,443]" fontSize="11" level="3" pageId="8" pageNumber="76" reason="3">
<emphasis id="B924EA810377EF691E897ED2FEAD6CFA" bold="true" box="[189,364,417,443]" pageId="8" pageNumber="76">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930377EF691EDE7EA2FE746F74" blockId="8.[189,1398,465,566]" pageId="8" pageNumber="76">
<materialsCitation id="3B383CCE0377EF691EDE7EA2FE746F74" ID-GBIF-Occurrence="3127434327" collectingDate="2004-04-15" collectionCode="SMF" collectorName="V. Vignoli &amp; P. Crucitti" country="Iran" elevation="1481" latitude="31.216667" location="Abarqu" longLatPrecision="1217" longitude="53.416668" pageId="8" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Yazd Province" typeStatus="holotype">
<emphasis id="B924EA810377EF691EDE7EA2FE9D6CAA" bold="true" box="[234,348,465,491]" pageId="8" pageNumber="76">
<typeStatus id="54EB88310377EF691EDE7EA2FE9D6CAA" box="[234,348,465,491]" pageId="8" pageNumber="76" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34776030377EF691E897E85FEC86F51" box="[189,265,502,528]" name="Iran" pageId="8" pageNumber="76">IRAN</collectingCountry>
•
<specimenCount id="9D56FD1A0377EF691F1C7E84FE806F50" box="[296,321,503,529]" pageId="8" pageNumber="76" type="male">♂</specimenCount>
;
<collectingRegion id="4994F8710377EF691F7B7E85FDC26F51" box="[335,515,502,528]" country="Iran" name="Yazd" pageId="8" pageNumber="76">Yazd Province</collectingRegion>
,
<location id="8E8F60480377EF691C247E85FDAF6F51" LSID="urn:lsid:plazi:treatment:03F987850379EF691C5579BDFC7E68C2:8E8F60480377EF691C247E85FDAF6F51" box="[528,622,502,528]" country="Iran" latitude="31.216667" longLatPrecision="1217" longitude="53.416668" name="Abarqu" pageId="8" pageNumber="76" stateProvince="Yazd Province">Abarqu</location>
;
<geoCoordinate id="EE6450540377EF691C487E87FD2D6F50" box="[636,748,500,529]" degrees="31" direction="north" minutes="13" orientation="latitude" pageId="8" pageNumber="76" precision="925" value="31.216667">31°13′ N</geoCoordinate>
,
<geoCoordinate id="EE6450540377EF691CCF7E87FCA96F50" box="[763,872,500,529]" degrees="53" direction="east" minutes="25" orientation="longitude" pageId="8" pageNumber="76" precision="925" value="53.416668">53°25′ E</geoCoordinate>
;
<elevation id="007DD1A00377EF691D437E85FBD96F51" box="[887,1048,502,529]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="8" pageNumber="76" unit="m" value="1481.0">
<quantity id="4CA89B760377EF691D437E85FC156F50" box="[887,980,502,529]" metricMagnitude="3" metricUnit="m" metricValue="1.4809999999999999" pageId="8" pageNumber="76" unit="m" value="1481.0">1481 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEE10530377EF691A1C7E85FB0D6F51" box="[1064,1228,502,529]" pageId="8" pageNumber="76" value="2004-04-15">
<collectingDate id="EFAAE9BB0377EF691A1C7E85FB0D6F51" box="[1064,1228,502,529]" pageId="8" pageNumber="76" value="2004-04-15">15 Apr. 2004</collectingDate>
</date>
;
<collectorName id="26A553450377EF691AED7E84FA976F51" box="[1241,1366,502,529]" pageId="8" pageNumber="76">V. Vignoli</collectorName>
&amp;
<collectorName id="26A553450377EF691E897D6FFEF26F74" box="[189,307,539,566]" pageId="8" pageNumber="76">P. Crucitti</collectorName>
leg.;
<collectionCode id="ED41AE560377EF691F417D68FE706F74" box="[373,433,539,565]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="76">SMF</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A65180377EF691E897D2DFEF8684E" pageId="8" pageNumber="76" type="description">
<paragraph id="8BEF36930377EF691E897D2DFE8F6F39" blockId="8.[189,334,606,632]" box="[189,334,606,632]" pageId="8" pageNumber="76">
<heading id="D0A781FF0377EF691E897D2DFE8F6F39" bold="true" box="[189,334,606,632]" fontSize="11" level="3" pageId="8" pageNumber="76" reason="3">
<emphasis id="B924EA810377EF691E897D2DFE8F6F39" bold="true" box="[189,334,606,632]" pageId="8" pageNumber="76">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930377EF691EDE7DFCFEEB6FE8" blockId="8.[187,1399,655,977]" box="[234,298,655,681]" pageId="8" pageNumber="76">
<emphasis id="B924EA810377EF691EDE7DFCFEEB6FE8" bold="true" box="[234,298,655,681]" pageId="8" pageNumber="76">Male</emphasis>
</paragraph>
<paragraph id="8BEF36930377EF691E897DC0FDD36E90" blockId="8.[187,1399,655,977]" pageId="8" pageNumber="76">
Habitus as in
<figureCitation id="136B2A160377EF691F6B7DC0FE7C6F8C" box="[351,445,691,717]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[221,1385,525,1854]" captionTargetId="figure-110@7.[202,1385,525,1854]" captionTargetPageId="7" captionText="Fig. 3. A–D. Berlandina artaxerxes sp. nov., holotype, ♂ (SMF). A. Habitus, dorsal view. B–D. Palp, ventral, proventral and retrolateral views. –E–G. Berlandina mesopotamica Al-Khazali,2020, ♀ (MHNG). E. Habitus, dorsal view. F–G. Epigyne, ventral and dorsal views. Abbreviations: Ao = apical outgrowth; Eb = embolar base; Po = pointed process. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906013" httpUri="https://zenodo.org/record/4906013/files/figure.png" pageId="8" pageNumber="76">Fig. 3A</figureCitation>
. Total length 5.55. Carapace 2.95 long, 2.15 wide. Eye sizes and interdistance of PMEs:AME = 0.13, ALE = 0.13, PME = 0.13, PLE = 0.15, PME–PME = 0.07. Carapace, sternum, labium, chelicerae and maxillae yellowish brown; carapace with small dark submarginal patches. Chelicera with a small tooth on promargin and a serrated, bifurcate keel on retromargin. Legs the same color as carapace, without annulations. Abdomen gray, without any clear pattern, although some brown markings are visible. Spinnerets light brown and uniform in color. Measurements of legs: I: 7.00 (1.95, 1.12, 1.54, 1.31, 1.08), II: 6.29 (1.68, 1.08, 1.24, 1.26, 1.03), III: 5.04 + missing Ta (1.62, 0.99, 0.94, 1.49, Ta missing), IV: 8.58 (2.25, 1.20, 1.49, 2.39, 1.25).
</paragraph>
<paragraph id="8BEF36930377EF691E897C8AFAF669E6" blockId="8.[188,1399,1017,1191]" pageId="8" pageNumber="76">
Palp as in
<figureCitation id="136B2A160377EF691F067C8AFE716952" box="[306,432,1017,1043]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[221,1385,525,1854]" captionTargetId="figure-110@7.[202,1385,525,1854]" captionTargetPageId="7" captionText="Fig. 3. A–D. Berlandina artaxerxes sp. nov., holotype, ♂ (SMF). A. Habitus, dorsal view. B–D. Palp, ventral, proventral and retrolateral views. –E–G. Berlandina mesopotamica Al-Khazali,2020, ♀ (MHNG). E. Habitus, dorsal view. F–G. Epigyne, ventral and dorsal views. Abbreviations: Ao = apical outgrowth; Eb = embolar base; Po = pointed process. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906013" httpUri="https://zenodo.org/record/4906013/files/figure.png" pageId="8" pageNumber="76">Fig. 3B–D</figureCitation>
. Tibia with massive apophysis, as long as tibia, pointed tip directed dorso-anteriorly with angle about 45° to the axis of the palp; cymbium ovoid, 1.6 times as long as wide; median apophysis long, oblique, about as long as tibial apophysis; embolar base (
<emphasis id="B924EA810377EF691DA97B30FC7F691C" box="[925,958,1091,1117]" italics="true" pageId="8" pageNumber="76">Eb</emphasis>
) complex, with 2 outgrowths: a basal pointed process (
<emphasis id="B924EA810377EF691FBD7B1AFE6B69C3" box="[393,426,1129,1154]" italics="true" pageId="8" pageNumber="76">Po</emphasis>
) weakly sclerotized, and a sclerotized apical outgrowth (
<emphasis id="B924EA810377EF691A607B1BFBB469C3" box="[1108,1141,1128,1154]" italics="true" pageId="8" pageNumber="76">Ao</emphasis>
) [tip appears broken] parallel to a bent, long embolus; embolus proper gently curved and making a course of about 90°.
</paragraph>
<paragraph id="8BEF36930377EF691EDE7BA3FE8469AB" blockId="8.[189,325,1232,1295]" box="[234,325,1232,1258]" pageId="8" pageNumber="76">
<emphasis id="B924EA810377EF691EDE7BA3FE8469AB" bold="true" box="[234,325,1232,1258]" pageId="8" pageNumber="76">Female</emphasis>
</paragraph>
<paragraph id="8BEF36930377EF691E897B86FEF8684E" blockId="8.[189,325,1232,1295]" box="[189,313,1269,1295]" pageId="8" pageNumber="76">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A65180377EF691E897A4BFC7E68C2" box="[189,959,1336,1411]" pageId="8" pageNumber="76" type="distribution">
<paragraph id="8BEF36930377EF691E897A4BFE976813" blockId="8.[189,342,1336,1362]" box="[189,342,1336,1362]" pageId="8" pageNumber="76">
<heading id="D0A781FF0377EF691E897A4BFE976813" bold="true" box="[189,342,1336,1362]" fontSize="11" level="3" pageId="8" pageNumber="76" reason="3">
<emphasis id="B924EA810377EF691E897A4BFE976813" bold="true" box="[189,342,1336,1362]" pageId="8" pageNumber="76">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930377EF691E897A1AFC7E68C2" blockId="8.[189,959,1384,1411]" box="[189,959,1384,1411]" pageId="8" pageNumber="76">
Known only from the
<typeStatus id="54EB88310377EF691FF67A1AFE3468C2" box="[450,501,1385,1411]" pageId="8" pageNumber="76">type</typeStatus>
locality in
<collectingRegion id="4994F8710377EF691C4C7A1BFCE468C3" box="[632,805,1384,1410]" country="Iran" name="Yazd" pageId="8" pageNumber="76">Yazd Province</collectingRegion>
, central
<collectingCountry id="F34776030377EF691DBC7A1AFC7A68C2" box="[904,955,1385,1411]" name="Iran" pageId="8" pageNumber="76">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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