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<document id="DD9267707F245BB876BCA38EE474DF82" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850374EF6C1C697A61FBE46E21" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptodrassus iranicus Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="3FD0F576-4945-4079-B0BB-503412D640D5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="81" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="79" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0318741427E5AB65653C91D7BC020FA9">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="82E0016A3C9F7D33EDC4D5DD1CD352B7">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="3C980823D9CCCA2B2BEE7751290088B2">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="DC2FC86281C859E38E80292876C39B6E">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="8E7E708C662508B0F733B067A3A051FD">Esyunin, Sergei L.</mods:namePart>
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<mods:namePart id="223454F9936254B295AFDAD184FAC404">Marusik, Yuri M.</mods:namePart>
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<mods:date id="1148CDD629F28AED3C83D1A2D6F8DE14">2021</mods:date>
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<heading id="D0A781FF0374EF6A1C697A61FC16686C" box="[605,983,1298,1325]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="79" reason="2">
<taxonomicName id="4C504D100374EF6A1C697A61FCB5686D" ID-CoL="98YVV" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[605,884,1298,1325]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="79" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B924EA810374EF6A1C697A61FCB5686D" bold="true" box="[605,884,1298,1325]" italics="true" pageId="11" pageNumber="79">Cryptodrassus iranicus</emphasis>
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<taxonomicNameLabel id="A21757FA0374EF6A1D487A60FC16686C" box="[892,983,1299,1325]" pageId="11" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BEF36930374EF6A1CC37A2AFCFD6835" blockId="11.[374,1214,1298,1396]" box="[759,828,1369,1396]" pageId="11" pageNumber="79">
<figureCitation id="136B2A160374EF6A1CC37A2AFCFD6835" box="[759,828,1369,1396]" captionStart="Fig" captionStartId="12.[189,232,1786,1812]" captionTargetBox="[191,1398,265,1747]" captionTargetId="figure-11@12.[189,1398,265,1749]" captionTargetPageId="12" captionText="Fig. 7. Cryptodrassus iranicus sp. nov., holotype, ♂ (MHNG). A, C, H. Palp, ventral, proventral and retroventral views. B, D. Palp, retrolateral view. EG. Bulb, ventral, proventral and retroventral views. I. Habitus, dorsal view. J. Chelicerae, posterior view. Abbreviations: Eo = embolar outgrowth; Ma = median apophysis; Tb = tegular bulge. Scale bars: AB, E, J = 0.2 mm; D, H = 0.1 mm; I = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906027" httpUri="https://zenodo.org/record/4906027/files/figure.png" pageId="11" pageNumber="79">Fig. 7</figureCitation>
</paragraph>
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<paragraph id="8BEF36930374EF6A1E897AE8FEF768F4" blockId="11.[189,310,1435,1461]" box="[189,310,1435,1461]" pageId="11" pageNumber="79">
<heading id="D0A781FF0374EF6A1E897AE8FEF768F4" bold="true" box="[189,310,1435,1461]" fontSize="11" level="3" pageId="11" pageNumber="79" reason="3">
<emphasis id="B924EA810374EF6A1E897AE8FEF768F4" bold="true" box="[189,310,1435,1461]" pageId="11" pageNumber="79">Diagnosis</emphasis>
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The new species differs from the congeners by having a distinct tegular bulge (
<emphasis id="B924EA810374EF6A1AB77AB9FB6368A5" box="[1155,1186,1482,1508]" italics="true" pageId="11" pageNumber="79">Tb</emphasis>
) near the base of embolus.
</paragraph>
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<subSubSection id="C34A65180374EF6A1E897943FB196B39" box="[189,1240,1584,1657]" pageId="11" pageNumber="79" type="etymology">
<paragraph id="8BEF36930374EF6A1E897943FE856B0B" blockId="11.[189,324,1584,1610]" box="[189,324,1584,1610]" pageId="11" pageNumber="79">
<heading id="D0A781FF0374EF6A1E897943FE856B0B" bold="true" box="[189,324,1584,1610]" fontSize="11" level="3" pageId="11" pageNumber="79" reason="3">
<emphasis id="B924EA810374EF6A1E897943FE856B0B" bold="true" box="[189,324,1584,1610]" pageId="11" pageNumber="79">Etymology</emphasis>
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<paragraph id="8BEF36930374EF6A1E89792DFB196B39" blockId="11.[189,1240,1630,1657]" box="[189,1240,1630,1657]" pageId="11" pageNumber="79">The specific epithet is an adjective and refers to the collection locality of the new species.</paragraph>
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<subSubSection id="C34A65180374EF6A1E8979D3FB766A4C" pageId="11" pageNumber="79" type="materials_examined">
<paragraph id="8BEF36930374EF6A1E8979D3FEAD6BFB" blockId="11.[189,364,1696,1722]" box="[189,364,1696,1722]" pageId="11" pageNumber="79">
<heading id="D0A781FF0374EF6A1E8979D3FEAD6BFB" bold="true" box="[189,364,1696,1722]" fontSize="11" level="3" pageId="11" pageNumber="79" reason="3">
<emphasis id="B924EA810374EF6A1E8979D3FEAD6BFB" bold="true" box="[189,364,1696,1722]" pageId="11" pageNumber="79">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B383CCE0374EF6A1EDE79BCFB766A4C" ID-GBIF-Occurrence="3127434312" collectingDate="2017-04" collectionCode="MHNG" collectorName="M. Amiri" country="Iran" location="Zarrin Joob Vil" pageId="11" pageNumber="79" specimenCount="1" specimenCount-male="1" stateProvince="Kermanshah Province" typeStatus="holotype">
<paragraph id="8BEF36930374EF6A1EDE79BCFE9D6BA8" blockId="11.[189,1207,1743,1805]" box="[234,348,1743,1769]" pageId="11" pageNumber="79">
<emphasis id="B924EA810374EF6A1EDE79BCFE9D6BA8" bold="true" box="[234,348,1743,1769]" pageId="11" pageNumber="79">
<typeStatus id="54EB88310374EF6A1EDE79BCFE9D6BA8" box="[234,348,1743,1769]" pageId="11" pageNumber="79" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BEF36930374EF6A1E897980FB766A4C" blockId="11.[189,1207,1743,1805]" box="[189,1207,1779,1805]" pageId="11" pageNumber="79">
<collectingCountry id="F34776030374EF6A1E897980FEC96A4C" box="[189,264,1779,1805]" name="Iran" pageId="11" pageNumber="79">IRAN</collectingCountry>
<specimenCount id="9D56FD1A0374EF6A1F167980FEFA6A4C" box="[290,315,1779,1805]" pageId="11" pageNumber="79" type="male"></specimenCount>
;
<collectingRegion id="4994F8710374EF6A1F737980FD8F6A4C" box="[327,590,1779,1805]" country="Iran" name="Kermanshah" pageId="11" pageNumber="79">Kermanshah Province</collectingRegion>
,
<location id="8E8F60480374EF6A1C6E7980FCD26A4C" LSID="urn:lsid:plazi:treatment:03F987850374EF6C1C697A61FBE46E21:8E8F60480374EF6A1C6E7980FCD26A4C" box="[602,787,1779,1805]" country="Iran" name="Zarrin Joob Vil" pageId="11" pageNumber="79" stateProvince="Kermanshah Province">Zarrin Joob Vil</location>
.;
<date id="FFEE10530374EF6A1D167980FC5B6A4C" box="[802,922,1779,1805]" pageId="11" pageNumber="79" value="2017-04">
<collectingDate id="EFAAE9BB0374EF6A1D167980FC5B6A4C" box="[802,922,1779,1805]" pageId="11" pageNumber="79" value="2017-04">Apr. 2017</collectingDate>
</date>
;
<collectorName id="26A553450374EF6A1D927980FBD56A4C" box="[934,1044,1779,1805]" pageId="11" pageNumber="79">M. Amiri</collectorName>
leg.;
<collectionCode id="ED41AE560374EF6A1A627980FB756A4C" box="[1110,1204,1779,1805]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="79" type="Museum">MHNG</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C34A65180374EF6C1E897847FEF86FAF" lastPageId="13" lastPageNumber="81" pageId="11" pageNumber="79" type="description">
<paragraph id="8BEF36930374EF6A1E897847FE8F6A0F" blockId="11.[189,334,1844,1870]" box="[189,334,1844,1870]" pageId="11" pageNumber="79">
<heading id="D0A781FF0374EF6A1E897847FE8F6A0F" bold="true" box="[189,334,1844,1870]" fontSize="11" level="3" pageId="11" pageNumber="79" reason="3">
<emphasis id="B924EA810374EF6A1E897847FE8F6A0F" bold="true" box="[189,334,1844,1870]" pageId="11" pageNumber="79">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930374EF6A1EDE7817FEEB6A3F" blockId="11.[189,1400,1892,2025]" box="[234,298,1892,1918]" pageId="11" pageNumber="79">
<emphasis id="B924EA810374EF6A1EDE7817FEEB6A3F" bold="true" box="[234,298,1892,1918]" pageId="11" pageNumber="79">Male</emphasis>
</paragraph>
<paragraph id="8BEF36930374EF6A1E8978F4FAB96AA9" blockId="11.[189,1400,1892,2025]" pageId="11" pageNumber="79">
Habitus as in
<figureCitation id="136B2A160374EF6A1F5578F4FE756AE3" box="[353,436,1927,1954]" captionStart="Fig" captionStartId="12.[189,232,1786,1812]" captionTargetBox="[191,1398,265,1747]" captionTargetId="figure-11@12.[189,1398,265,1749]" captionTargetPageId="12" captionText="Fig. 7. Cryptodrassus iranicus sp. nov., holotype, ♂ (MHNG). A, C, H. Palp, ventral, proventral and retroventral views. B, D. Palp, retrolateral view. EG. Bulb, ventral, proventral and retroventral views. I. Habitus, dorsal view. J. Chelicerae, posterior view. Abbreviations: Eo = embolar outgrowth; Ma = median apophysis; Tb = tegular bulge. Scale bars: AB, E, J = 0.2 mm; D, H = 0.1 mm; I = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906027" httpUri="https://zenodo.org/record/4906027/files/figure.png" pageId="11" pageNumber="79">Fig. 7I</figureCitation>
. Total length 3.59. Carapace 1.57 long, 1.26 wide. Eye sizes and interdistance of PMEs: AME = 0.08, ALE = 0.09, PME = 0.11, PLE = 0.07, PMEPME = 0.03. Carapace, chelicerae, sternum, maxillae and labium light reddish brown. Chelicerae (
<figureCitation id="136B2A160374EF6A1D9178BDFC376AA8" box="[933,1014,1998,2025]" captionStart="Fig" captionStartId="12.[189,232,1786,1812]" captionTargetBox="[191,1398,265,1747]" captionTargetId="figure-11@12.[189,1398,265,1749]" captionTargetPageId="12" captionText="Fig. 7. Cryptodrassus iranicus sp. nov., holotype, ♂ (MHNG). A, C, H. Palp, ventral, proventral and retroventral views. B, D. Palp, retrolateral view. EG. Bulb, ventral, proventral and retroventral views. I. Habitus, dorsal view. J. Chelicerae, posterior view. Abbreviations: Eo = embolar outgrowth; Ma = median apophysis; Tb = tegular bulge. Scale bars: AB, E, J = 0.2 mm; D, H = 0.1 mm; I = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906027" httpUri="https://zenodo.org/record/4906027/files/figure.png" pageId="11" pageNumber="79">Fig. 7J</figureCitation>
) with a small promarginal tooth.
</paragraph>
<caption id="DF2F661B0373EF6D1E897989FAFD6A3E" ID-DOI="http://doi.org/10.5281/zenodo.4906027" ID-Zenodo-Dep="4906027" httpUri="https://zenodo.org/record/4906027/files/figure.png" pageId="12" pageNumber="80" startId="12.[189,232,1786,1812]" targetBox="[191,1398,265,1747]" targetPageId="12">
<paragraph id="8BEF36930373EF6D1E897989FAFD6A3E" blockId="12.[189,1399,1786,1919]" pageId="12" pageNumber="80">
<emphasis id="B924EA810373EF6D1E897989FECD6A54" bold="true" box="[189,268,1786,1813]" pageId="12" pageNumber="80">Fig. 7.</emphasis>
<taxonomicName id="4C504D100373EF6D1F227989FDEA6A55" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[278,555,1786,1812]" class="Arachnida" family="Gnaphosidae" genus="Cryptodrassus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="80" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov.">
<emphasis id="B924EA810373EF6D1F227989FDEA6A55" box="[278,555,1786,1812]" italics="true" pageId="12" pageNumber="80">Cryptodrassus iranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0373EF6D1C007988FD506A54" box="[564,657,1787,1813]" pageId="12" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (MHNG).
<emphasis id="B924EA810373EF6D1D867989FBD56A54" bold="true" box="[946,1044,1786,1813]" pageId="12" pageNumber="80">A, C, H</emphasis>
. Palp, ventral, proventral and retroventral views.
<emphasis id="B924EA810373EF6D1F95786DFE196A79" bold="true" box="[417,472,1822,1848]" pageId="12" pageNumber="80">B, D</emphasis>
. Palp, retrolateral view.
<emphasis id="B924EA810373EF6D1CCC786EFCF06A76" bold="true" box="[760,817,1821,1847]" pageId="12" pageNumber="80">EG</emphasis>
. Bulb, ventral, proventral and retroventral views.
<emphasis id="B924EA810373EF6D1E897832FF096A1A" bold="true" box="[189,200,1857,1883]" pageId="12" pageNumber="80">I</emphasis>
. Habitus, dorsal view.
<emphasis id="B924EA810373EF6D1FD47832FE2C6A1A" bold="true" box="[480,493,1857,1883]" pageId="12" pageNumber="80">J</emphasis>
. Chelicerae, posterior view. Abbreviations:
<emphasis id="B924EA810373EF6D1DCB7831FBE16A1A" box="[1023,1056,1858,1883]" italics="true" pageId="12" pageNumber="80">Eo</emphasis>
= embolar outgrowth;
<emphasis id="B924EA810373EF6D1B027831FA9C6A1A" box="[1334,1373,1858,1883]" italics="true" pageId="12" pageNumber="80">Ma</emphasis>
= median apophysis;
<emphasis id="B924EA810373EF6D1FAB7817FE7F6A3F" box="[415,446,1892,1918]" italics="true" pageId="12" pageNumber="80">Tb</emphasis>
= tegular bulge. Scale bars: AB, E, J = 0.2 mm; D, H = 0.1 mm; I = 1 mm.
</paragraph>
</caption>
<paragraph id="8BEF36930372EF6C1E897E7AFE856CF4" blockId="13.[187,1400,265,437]" pageId="13" pageNumber="81">Maxillae with strong lateral protrusion. Legs the same color as carapace, with few spines and without annulations.Abdomen light brown, without any markings, with a distinct reddish brown scutum anteriorly. Spinnerets light brown and uniform in color. Measurements of legs: I: 4.78 (1.35, 0.79, 1.18, 0.85, 0.61), II: 3.94 (1.10, 0.72, 0.93, 0.70, 0.49), III: 3.16 (0.86, 0.52, 0.62, 0.69, 0.47), IV: 4.52 (1.19, 0.69, 0.98, 1.04, 0.62).</paragraph>
<paragraph id="8BEF36930372EF6C1E897EAFFA836FC9" blockId="13.[189,1400,476,648]" pageId="13" pageNumber="81">
Palp as in
<figureCitation id="136B2A160372EF6C1F057EAFFE706CB6" box="[305,433,476,503]" captionStart="Fig" captionStartId="12.[189,232,1786,1812]" captionTargetBox="[191,1398,265,1747]" captionTargetId="figure-11@12.[189,1398,265,1749]" captionTargetPageId="12" captionText="Fig. 7. Cryptodrassus iranicus sp. nov., holotype, ♂ (MHNG). A, C, H. Palp, ventral, proventral and retroventral views. B, D. Palp, retrolateral view. EG. Bulb, ventral, proventral and retroventral views. I. Habitus, dorsal view. J. Chelicerae, posterior view. Abbreviations: Eo = embolar outgrowth; Ma = median apophysis; Tb = tegular bulge. Scale bars: AB, E, J = 0.2 mm; D, H = 0.1 mm; I = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906027" httpUri="https://zenodo.org/record/4906027/files/figure.png" pageId="13" pageNumber="81">Fig. 7AH</figureCitation>
. Femur and patella unmodified; tibial apophysis small, sharply pointed, shorter than tibia; cymbium with rounded and short tip; bulb elongate oval; tegulum with antero-prolateral bulge (
<emphasis id="B924EA810372EF6C1B7A7D73FAAC6F5B" box="[1358,1389,512,538]" italics="true" pageId="13" pageNumber="81">Tb</emphasis>
) near base of embolus; median apophysis (
<emphasis id="B924EA810372EF6C1C877D55FD1B6F7E" box="[691,730,550,575]" italics="true" pageId="13" pageNumber="81">Ma</emphasis>
) (or conductor sensu
<bibRefCitation id="EFC14B620372EF6C1DD57D56FB136F7E" author="Chatzaki M. &amp; Thaler K. &amp; Mylonas M." box="[993,1234,549,575]" pageId="13" pageNumber="81" pagination="559 - 601" refId="ref17204" refString="Chatzaki M., Thaler K. &amp; Mylonas M. 2002. Ground spiders (Gnaphosidae; Araneae) of Crete (Greece). Taxonomy and distribution. I. Revue suisse de Zoologie 109 (3): 559 - 601. https: // doi. org / 10.5962 / bhl. part. 79611" type="journal article" year="2002">
Chatzaki
<emphasis id="B924EA810372EF6C1A667D55FB4F6F7E" box="[1106,1166,549,575]" italics="true" pageId="13" pageNumber="81">et al.</emphasis>
2002
</bibRefCitation>
and
<bibRefCitation id="EFC14B620372EF6C1B3A7D56FF3B6F22" author="Chatzaki M." pageId="13" pageNumber="81" pagination="521 - 545" refId="ref17086" refString="Chatzaki M. 2018. On the ground spider genera Marjanus gen. nov., Lasophorus gen. nov. and Turkozelotes Kovblyuk &amp; Seyyar, 2009 (Araneae: Gnaphosidae) from Greece. Zootaxa 4392 (3): 521 - 545. https: // doi. org / 10.11646 / zootaxa. 4392.3.5" type="journal article" year="2018">Chatzaki 2018</bibRefCitation>
) subtriangular in ventral view, lacking hook; embolus relatively short, filiform, originating at ca 8 oclock position and terminating at about 12, terminal part with short, spine-like outgrowth (
<emphasis id="B924EA810372EF6C1B257D1DFAF36FC6" box="[1297,1330,622,647]" italics="true" pageId="13" pageNumber="81">Eo</emphasis>
).
</paragraph>
<paragraph id="8BEF36930372EF6C1EDE7DC3FE846F8B" blockId="13.[189,325,688,750]" box="[234,325,688,714]" pageId="13" pageNumber="81">
<emphasis id="B924EA810372EF6C1EDE7DC3FE846F8B" bold="true" box="[234,325,688,714]" pageId="13" pageNumber="81">Female</emphasis>
</paragraph>
<paragraph id="8BEF36930372EF6C1E897DA7FEF86FAF" blockId="13.[189,325,688,750]" box="[189,313,724,750]" pageId="13" pageNumber="81">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A65180372EF6C1E897C65FBE46E21" box="[189,1061,790,864]" pageId="13" pageNumber="81" type="distribution">
<paragraph id="8BEF36930372EF6C1E897C65FE976E71" blockId="13.[189,342,790,816]" box="[189,342,790,816]" pageId="13" pageNumber="81">
<heading id="D0A781FF0372EF6C1E897C65FE976E71" bold="true" box="[189,342,790,816]" fontSize="11" level="3" pageId="13" pageNumber="81" reason="3">
<emphasis id="B924EA810372EF6C1E897C65FE976E71" bold="true" box="[189,342,790,816]" pageId="13" pageNumber="81">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930372EF6C1E897C35FBE46E21" blockId="13.[189,1061,838,864]" box="[189,1061,838,864]" pageId="13" pageNumber="81">
Known only from the
<typeStatus id="54EB88310372EF6C1FF67C35FE346E21" box="[450,501,838,864]" pageId="13" pageNumber="81">type</typeStatus>
locality in
<collectingRegion id="4994F8710372EF6C1C4D7C35FC416E21" box="[633,896,838,864]" country="Iran" name="Kermanshah" pageId="13" pageNumber="81">Kermanshah Province</collectingRegion>
, western
<collectingCountry id="F34776030372EF6C1DDA7C35FBE06E21" box="[1006,1057,838,864]" name="Iran" pageId="13" pageNumber="81">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>