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<document id="50AD45A4A9B9FD6454595BD0E8941582" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850370EF711C4E7D2AFE0E698A" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echemus caspicus Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="CFBF122D-F516-4D05-B4F9-85AD591F4511" docUuidSource="ZooBank" docVersion="8" lastPageNumber="84" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="83" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3DC298A20136BF40FCF736EDCA82887F">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="F49CBE7520ADD3B3AA04DDAF27506873">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="84E4E91F9CFED8A70B511B002000C93C">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
<mods:nameIdentifier id="2FD176B2CC12EEDAEE68585358D18DA9" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="53D1CFE662242DAD2D2BBBFD4571732F">Chatzaki, Maria</mods:namePart>
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<mods:nameIdentifier id="AFE707725EAA90B5B7A919ADF8FE9518" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="C3853CD86C47F933945BF974C4EA6AA9">Author</mods:roleTerm>
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<mods:namePart id="2F768C9631EC8EA9B4A78609A49D4066">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="B0B15A2B422D87D8724D5B1E509ACE08">Sergei.</mods:affiliation>
<mods:nameIdentifier id="C523C4B4E942938C8008D54B3ED257AA" type="email">Sergei.Esyunin@psu.ru</mods:nameIdentifier>
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<mods:namePart id="A6C6AA018C10EA52DFBC9A73C983358E">Marusik, Yuri M.</mods:namePart>
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<mods:title id="DBECC71EDC26E226A81669D98251A95C">European Journal of Taxonomy</mods:title>
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<mods:date id="743DFED30974C470E80C5F00299687FD">2021</mods:date>
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<mods:number id="88F242CC8D7DE34AE266834CC1494A33">2021-06-04</mods:number>
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<treatment id="03F987850370EF711C4E7D2AFE0E698A" ID-DOI="http://doi.org/10.5281/zenodo.4906471" ID-GBIF-Taxon="182597838" ID-Zenodo-Dep="4906471" LSID="urn:lsid:zoobank.org:act:CFBF122D-F516-4D05-B4F9-85AD591F4511" httpUri="http://treatment.plazi.org/id/03F987850370EF711C4E7D2AFE0E698A" lastPageId="16" lastPageNumber="84" pageId="15" pageNumber="83">
<subSubSection id="C34A65180370EF6E1C4E7D2AFC7B6F35" box="[634,954,601,628]" pageId="15" pageNumber="83" type="nomenclature">
<paragraph id="8BEF36930370EF6E1C4E7D2AFC7B6F35" blockId="15.[369,1218,601,699]" box="[634,954,601,628]" pageId="15" pageNumber="83">
<heading id="D0A781FF0370EF6E1C4E7D2AFC7B6F35" box="[634,954,601,628]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="83" reason="2">
<taxonomicName id="4C504D100370EF6E1C4E7D2AFC986F35" ID-CoL="98YVT" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[634,857,601,628]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="caspicus" status="sp. nov.">
<emphasis id="B924EA810370EF6E1C4E7D2AFC986F35" bold="true" box="[634,857,601,628]" italics="true" pageId="15" pageNumber="83">Echemus caspicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0370EF6E1D547D29FC7B6F35" box="[864,954,602,628]" pageId="15" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C34A65180370EF6E1F457D0EFCFD6FFA" pageId="15" pageNumber="83" type="description">
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<uri id="FFC13A910370EF6E1F457D0EFB036FD9" box="[369,1218,637,664]" pageId="15" pageNumber="83">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BEF36930370EF6E1CC37DD2FCFD6FFA" blockId="15.[369,1218,601,699]" box="[759,828,673,699]" pageId="15" pageNumber="83">
<figureCitation id="136B2A160370EF6E1CC37DD2FCFD6FFA" box="[759,828,673,699]" captionStart="Fig" captionStartId="15.[189,233,1964,1990]" captionTargetBox="[189,1398,1219,1921]" captionTargetId="figure-193@15.[189,1398,1219,1926]" captionTargetPageId="15" captionText="Fig. 9. Echemus caspicus sp. nov., holotype, ♀ (MHNG). A–B. Epigyne, dorsal and ventral views. C. Chelicerae, posterior view. D. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="15" pageNumber="83">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A65180370EF6E1E897D91FE5F6941" pageId="15" pageNumber="83" type="diagnosis">
<paragraph id="8BEF36930370EF6E1E897D91FEF76FBD" blockId="15.[189,310,738,764]" box="[189,310,738,764]" pageId="15" pageNumber="83">
<heading id="D0A781FF0370EF6E1E897D91FEF76FBD" bold="true" box="[189,310,738,764]" fontSize="11" level="3" pageId="15" pageNumber="83" reason="3">
<emphasis id="B924EA810370EF6E1E897D91FEF76FBD" bold="true" box="[189,310,738,764]" pageId="15" pageNumber="83">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930370EF6E1E897C62FE5F6941" blockId="15.[187,1399,785,1024]" pageId="15" pageNumber="83">
The new species is most similar to the generotype
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<emphasis id="B924EA810370EF6E1CC27C61FCC96E6A" box="[758,776,786,811]" italics="true" pageId="15" pageNumber="83">E</emphasis>
.
<emphasis id="B924EA810370EF6E1D237C62FC666E6A" box="[791,935,785,811]" italics="true" pageId="15" pageNumber="83">angustifrons</emphasis>
(Westring, 1861)
</taxonomicName>
by having an S-shaped septum (rather than rectangular as in, e.g.,
<taxonomicName id="4C504D100370EF6E1C817C46FC2B6E0E" authority="Chatzaki, 2019" authorityName="Chatzaki" authorityYear="2019" box="[693,1002,821,847]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="kaltsasi">
<emphasis id="B924EA810370EF6E1C817C46FD066E0F" box="[693,711,821,846]" italics="true" pageId="15" pageNumber="83">E</emphasis>
.
<emphasis id="B924EA810370EF6E1CE27C46FCF16E0E" box="[726,816,821,847]" italics="true" pageId="15" pageNumber="83">kaltsasi</emphasis>
<bibRefCitation id="EFC14B620370EF6E1D037C46FC2B6E0E" author="Chatzaki M. &amp; Van Keer J." box="[823,1002,821,847]" pageId="15" pageNumber="83" pagination="434 - 460" refId="ref17144" refString="Chatzaki M. &amp; Van Keer J. 2019. Ground spiders (Araneae: Gnaphosidae, Liocranidae, Prodidomidae) from the Greek islands Rodos, Symi and Karpathos, with the description of new species. Zootaxa 4646 (3): 434 - 460. https: // doi. org / 10.11646 / zootaxa. 4646.3.2" type="journal article" year="2019">Chatzaki, 2019</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C504D100370EF6E1A177C46FF386E33" authority="Kovblyuk &amp; Seyyar, 2009" authorityName="Kovblyuk &amp; Seyyar" authorityYear="2009" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="B924EA810370EF6E1A177C46FBF46E0F" box="[1059,1077,821,846]" italics="true" pageId="15" pageNumber="83">E</emphasis>
.
<emphasis id="B924EA810370EF6E1A707C46FBBA6E0E" box="[1092,1147,821,847]" italics="true" pageId="15" pageNumber="83">levyi</emphasis>
Kovblyuk &amp; Seyyar, 2009
</taxonomicName>
) but differs by the much narrower anterior part (or stalk), giving a bell-shaped form to the septum (widened part of septum/stalk of stem ratio&gt;2, vs &lt;
<quantity id="4CA89B760370EF6E1D187C08FCAE6ED4" box="[812,879,891,918]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="15" pageNumber="83" unit="in" value="1.5">1.5 in</quantity>
<taxonomicName id="4C504D100370EF6E1D437C0FFBEB6ED7" baseAuthorityName="Westring" baseAuthorityYear="1861" box="[887,1066,892,918]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B924EA810370EF6E1D437C0FFC486ED4" box="[887,905,892,917]" italics="true" pageId="15" pageNumber="83">E</emphasis>
.
<emphasis id="B924EA810370EF6E1DAC7C0FFBEB6ED7" box="[920,1066,892,918]" italics="true" pageId="15" pageNumber="83">angustifrons</emphasis>
</taxonomicName>
, cf.
<figureCitation id="136B2A160370EF6E1A6A7C08FB766ED7" box="[1118,1207,891,918]" captionStart="Fig" captionStartId="15.[189,233,1964,1990]" captionTargetBox="[189,1398,1219,1921]" captionTargetId="figure-193@15.[189,1398,1219,1926]" captionTargetPageId="15" captionText="Fig. 9. Echemus caspicus sp. nov., holotype, ♀ (MHNG). A–B. Epigyne, dorsal and ventral views. C. Chelicerae, posterior view. D. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="15" pageNumber="83">Fig. 9B</figureCitation>
and
<bibRefCitation id="EFC14B620370EF6E1AC47C08FEB36EF8" author="Chatzaki M. &amp; Van Keer J." pageId="15" pageNumber="83" pagination="434 - 460" refId="ref17144" refString="Chatzaki M. &amp; Van Keer J. 2019. Ground spiders (Araneae: Gnaphosidae, Liocranidae, Prodidomidae) from the Greek islands Rodos, Symi and Karpathos, with the description of new species. Zootaxa 4646 (3): 434 - 460. https: // doi. org / 10.11646 / zootaxa. 4646.3.2" type="journal article" year="2019">
Chatzaki &amp;
<collectingRegion id="4994F8710370EF6E1E897CD3FF2A6EFB" box="[189,235,928,954]" country="Turkey" name="Van" pageId="15" pageNumber="83">Van</collectingRegion>
Keer 2019
</bibRefCitation>
: fig. 9). Also, the anterior epigynal margin is equally sclerotized, round and reaches the lateral sides of the plate in the new species (vs extending anteriorly and forming a triangular sclerotization in
<taxonomicName id="4C504D100370EF6E1EEF7C94FE4F6941" baseAuthorityName="Westring" baseAuthorityYear="1861" box="[219,398,998,1024]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B924EA810370EF6E1EEF7C94FF2C6941" box="[219,237,999,1024]" italics="true" pageId="15" pageNumber="83">E</emphasis>
.
<emphasis id="B924EA810370EF6E1EC87C95FE4F6941" box="[252,398,998,1024]" italics="true" pageId="15" pageNumber="83">angustifrons</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A65180370EF6E1E897B5BFE5569D5" pageId="15" pageNumber="83" type="etymology">
<paragraph id="8BEF36930370EF6E1E897B5BFE856903" blockId="15.[189,324,1064,1090]" box="[189,324,1064,1090]" pageId="15" pageNumber="83">
<heading id="D0A781FF0370EF6E1E897B5BFE856903" bold="true" box="[189,324,1064,1090]" fontSize="11" level="3" pageId="15" pageNumber="83" reason="3">
<emphasis id="B924EA810370EF6E1E897B5BFE856903" bold="true" box="[189,324,1064,1090]" pageId="15" pageNumber="83">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930370EF6E1E897B25FE5569D5" blockId="15.[189,1399,1110,1172]" pageId="15" pageNumber="83">
The specific epithet is an adjective and refers to the Caspian region, in which the
<typeStatus id="54EB88310370EF6E1A527B24FB586930" box="[1126,1177,1111,1137]" pageId="15" pageNumber="83">type</typeStatus>
locality of the new species in located.
</paragraph>
</subSubSection>
<caption id="DF2F661B0370EF6E1E8978DFFA8A6AAB" ID-DOI="http://doi.org/10.5281/zenodo.4906031" ID-Zenodo-Dep="4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="15" pageNumber="83" startId="15.[189,233,1964,1990]" targetBox="[189,1398,1219,1921]" targetPageId="15">
<paragraph id="8BEF36930370EF6E1E8978DFFA8A6AAB" blockId="15.[189,1399,1964,2026]" pageId="15" pageNumber="83">
<emphasis id="B924EA810370EF6E1E8978DFFECF6A87" bold="true" box="[189,270,1964,1990]" pageId="15" pageNumber="83">Fig. 9.</emphasis>
<taxonomicName id="4C504D100370EF6E1F2D78DFFE376A87" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[281,502,1964,1990]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="83" phylum="Arthropoda" rank="species" species="caspicus" status="sp. nov.">
<emphasis id="B924EA810370EF6E1F2D78DFFE376A87" box="[281,502,1964,1990]" italics="true" pageId="15" pageNumber="83">Echemus caspicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0370EF6E1C3478DEFD9E6A86" box="[512,607,1965,1991]" pageId="15" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (MHNG).
<emphasis id="B924EA810370EF6E1DB378DFFC7E6A87" bold="true" box="[903,959,1964,1990]" pageId="15" pageNumber="83">A–B</emphasis>
. Epigyne, dorsal and ventral views.
<emphasis id="B924EA810370EF6E1E8B78BCFF136AA8" bold="true" box="[191,210,1999,2025]" pageId="15" pageNumber="83">C</emphasis>
. Chelicerae, posterior view.
<emphasis id="B924EA810370EF6E1C1678A3FDF66AAB" bold="true" box="[546,567,2000,2026]" pageId="15" pageNumber="83">D</emphasis>
. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<subSubSection id="C34A6518036FEF711E897E7AFE986CDC" pageId="16" pageNumber="84" type="materials_examined">
<paragraph id="8BEF3693036FEF711E897E7AFEAD6C63" blockId="16.[189,364,264,291]" box="[189,364,264,291]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="16" pageNumber="84" reason="3">
<emphasis id="B924EA81036FEF711E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" pageId="16" pageNumber="84">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036FEF711EDE7E4AFE986CDC" blockId="16.[189,1398,313,413]" pageId="16" pageNumber="84">
<materialsCitation id="3B383CCE036FEF711EDE7E4AFE986CDC" ID-GBIF-Occurrence="3127434311" collectingDate="2017-07" collectionCode="MHNG" collectorName="R. Rafiei-Jahed" country="Iran" elevation="523" latitude="36.766666" location="Shast Kalateh" longLatPrecision="1185" longitude="54.183334" pageId="16" pageNumber="84" specimenCount="1" specimenCount-female="1" stateProvince="Golestan Province" typeStatus="holotype">
<emphasis id="B924EA81036FEF711EDE7E4AFE9D6C12" bold="true" box="[234,348,313,339]" pageId="16" pageNumber="84">
<typeStatus id="54EB8831036FEF711EDE7E4AFE9D6C12" box="[234,348,313,339]" pageId="16" pageNumber="84" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3477603036FEF711E897E2DFEC96C39" box="[189,264,350,376]" name="Iran" pageId="16" pageNumber="84">IRAN</collectingCountry>
•
<specimenCount id="9D56FD1A036FEF711F2B7E2CFEF76C38" box="[287,310,351,377]" pageId="16" pageNumber="84" type="female">♀</specimenCount>
;
<collectingRegion id="4994F871036FEF711F767E2DFDDA6C39" box="[322,539,350,376]" country="Iran" name="Golestan" pageId="16" pageNumber="84">Golestan Province</collectingRegion>
,
<location id="8E8F6048036FEF711C117E2DFD086C39" LSID="urn:lsid:plazi:treatment:03F987850370EF711C4E7D2AFE0E698A:8E8F6048036FEF711C117E2DFD086C39" box="[549,713,350,376]" country="Iran" latitude="36.766666" longLatPrecision="1185" longitude="54.183334" name="Shast Kalateh" pageId="16" pageNumber="84" stateProvince="Golestan Province">Shast Kalateh</location>
;
<geoCoordinate id="EE645054036FEF711CE77E2EFCFF6C38" box="[723,830,349,377]" degrees="36" direction="north" minutes="46" orientation="latitude" pageId="16" pageNumber="84" precision="925" value="36.766666">36°46′ N</geoCoordinate>
,
<geoCoordinate id="EE645054036FEF711D7C7E2EFC6F6C38" box="[840,942,349,377]" degrees="54" direction="east" minutes="11" orientation="longitude" pageId="16" pageNumber="84" precision="925" value="54.183334">54°11′ E</geoCoordinate>
;
<elevation id="007DD1A0036FEF711D8D7E2DFB816C39" box="[953,1088,350,377]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="16" pageNumber="84" unit="m" value="523.0">
<quantity id="4CA89B76036FEF711D8D7E2DFBC36C38" box="[953,1026,350,377]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" pageId="16" pageNumber="84" unit="m" value="523.0">523 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEE1053036FEF711A787E2DFB786C39" box="[1100,1209,350,376]" pageId="16" pageNumber="84" value="2017-07">
<collectingDate id="EFAAE9BB036FEF711A787E2DFB786C39" box="[1100,1209,350,376]" pageId="16" pageNumber="84" value="2017-07">Jul. 2017</collectingDate>
</date>
;
<collectorName id="26A55345036FEF711AF77E2CFAB76C39" box="[1219,1398,350,377]" pageId="16" pageNumber="84">R. Rafiei-Jahed</collectorName>
leg.;
<collectionCode id="ED41AE56036FEF711ECC7EF0FE976CDC" box="[248,342,387,413]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="16" pageNumber="84" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036FEF711E897EB5FEF86970" pageId="16" pageNumber="84" type="description">
<paragraph id="8BEF3693036FEF711E897EB5FE8F6CA1" blockId="16.[189,334,454,480]" box="[189,334,454,480]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711E897EB5FE8F6CA1" bold="true" box="[189,334,454,480]" fontSize="11" level="3" pageId="16" pageNumber="84" reason="3">
<emphasis id="B924EA81036FEF711E897EB5FE8F6CA1" bold="true" box="[189,334,454,480]" pageId="16" pageNumber="84">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036FEF711EDE7E84FE846F50" blockId="16.[187,1400,503,790]" box="[234,325,503,529]" pageId="16" pageNumber="84">
<emphasis id="B924EA81036FEF711EDE7E84FE846F50" bold="true" box="[234,325,503,529]" pageId="16" pageNumber="84">Female</emphasis>
</paragraph>
<paragraph id="8BEF3693036FEF711E897D6FFC886E57" blockId="16.[187,1400,503,790]" pageId="16" pageNumber="84">
Habitus as in
<figureCitation id="136B2A16036FEF711F517D6FFE056F76" box="[357,452,540,567]" captionStart="Fig" captionStartId="15.[189,233,1964,1990]" captionTargetBox="[189,1398,1219,1921]" captionTargetId="figure-193@15.[189,1398,1219,1926]" captionTargetPageId="15" captionText="Fig. 9. Echemus caspicus sp. nov., holotype, ♀ (MHNG). A–B. Epigyne, dorsal and ventral views. C. Chelicerae, posterior view. D. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="16" pageNumber="84">Fig. 9D</figureCitation>
. Total length 5.29. Carapace 1.87 long, 1.26 wide. Eye sizes and interdistance of PMEs: AME = 0.14, ALE = 0.13, PME = 0.12, PLE = 0.11, PME–PME = 0.07. Carapace, sternum, labium, chelicerae and maxillae dark brown, without any pattern. Chelicerae (
<figureCitation id="136B2A16036FEF711A7C7D14FB606FC0" box="[1096,1185,615,641]" captionStart="Fig" captionStartId="15.[189,233,1964,1990]" captionTargetBox="[189,1398,1219,1921]" captionTargetId="figure-193@15.[189,1398,1219,1926]" captionTargetPageId="15" captionText="Fig. 9. Echemus caspicus sp. nov., holotype, ♀ (MHNG). A–B. Epigyne, dorsal and ventral views. C. Chelicerae, posterior view. D. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="16" pageNumber="84">Fig. 9C</figureCitation>
) with 3 pro- and 1 retromarginal teeth. Legs the same color as carapace, without annulations. Abdomen and spinnerets gray, without any pattern. Spinnerets uniform in color and slightly lighter in the apical segment. Measurements of legs: I: 4.59 (1.34, 0.91, 0.98, 0.76, 0.60), II: 4.26 (1.31, 0.83, 0.93, 0.66, 0.53), III: 4.04 (1.12, 0.73, 0.76, 0.78, 0.65), IV: 5.48 (1.47, 0.86, 1.11, 1.35, 0.69).
</paragraph>
<paragraph id="8BEF3693036FEF711E897C4CFBE76E89" blockId="16.[189,1402,831,969]" pageId="16" pageNumber="84">
Epigyne as in
<figureCitation id="136B2A16036FEF711F5B7C4CFE356E18" box="[367,500,831,857]" captionStart="Fig" captionStartId="15.[189,233,1964,1990]" captionTargetBox="[189,1398,1219,1921]" captionTargetId="figure-193@15.[189,1398,1219,1926]" captionTargetPageId="15" captionText="Fig. 9. Echemus caspicus sp. nov., holotype, ♀ (MHNG). A–B. Epigyne, dorsal and ventral views. C. Chelicerae, posterior view. D. Habitus, dorsal view. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906031" httpUri="https://zenodo.org/record/4906031/files/figure.png" pageId="16" pageNumber="84">Fig. 9A–B</figureCitation>
. Atrium large (ca 1.8 times as wide as long), wider than epigynal plate’s length, lateral margins distinct; septum thin, stalk 4.6 times thinner than atrium’s width; accessorial gland transverse and as long as receptacles, parallel to copulatory ducts; copulatory ducts transverse, moderately coiled, connected to oval receptacles posteriorly and to accessorial glands.
</paragraph>
<paragraph id="8BEF3693036FEF711EDE7C81FEEB694D" blockId="16.[189,313,1010,1073]" box="[234,298,1010,1036]" pageId="16" pageNumber="84">
<emphasis id="B924EA81036FEF711EDE7C81FEEB694D" bold="true" box="[234,298,1010,1036]" pageId="16" pageNumber="84">Male</emphasis>
</paragraph>
<paragraph id="8BEF3693036FEF711E897B64FEF86970" blockId="16.[189,313,1010,1073]" box="[189,313,1047,1073]" pageId="16" pageNumber="84">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A6518036FEF711E897B29FE0E698A" pageId="16" pageNumber="84" type="distribution">
<paragraph id="8BEF3693036FEF711E897B29FE976935" blockId="16.[189,342,1114,1140]" box="[189,342,1114,1140]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711E897B29FE976935" bold="true" box="[189,342,1114,1140]" fontSize="11" level="3" pageId="16" pageNumber="84" reason="3">
<emphasis id="B924EA81036FEF711E897B29FE976935" bold="true" box="[189,342,1114,1140]" pageId="16" pageNumber="84">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036FEF711E897BF8FE0E698A" blockId="16.[189,1399,1163,1227]" pageId="16" pageNumber="84">
Known only from the
<typeStatus id="54EB8831036FEF711FFE7BF8FE3C69E4" box="[458,509,1163,1189]" pageId="16" pageNumber="84">type</typeStatus>
locality in
<collectingRegion id="4994F871036FEF711CB37BF8FCA469E4" box="[647,869,1163,1189]" country="Iran" name="Golestan" pageId="16" pageNumber="84">Golestan Province</collectingRegion>
, northern
<collectingCountry id="F3477603036FEF711DD47BF8FBD269E4" box="[992,1043,1163,1189]" name="Iran" pageId="16" pageNumber="84">Iran</collectingCountry>
. This is the first record of the genus
<taxonomicName id="4C504D10036FEF711F3D7BC3FEB2698B" authorityName="Simon" authorityYear="1878" box="[265,371,1200,1226]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B924EA81036FEF711F3D7BC3FEB2698B" box="[265,371,1200,1226]" italics="true" pageId="16" pageNumber="84">Echemus</emphasis>
</taxonomicName>
in
<collectingCountry id="F3477603036FEF711FAC7BC2FE0A698A" box="[408,459,1201,1227]" name="Iran" pageId="16" pageNumber="84">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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