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<document id="BB36907C148603B5728A4F2B75DDA98C" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850361EF7E1C5E7D81FC1D6F87" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Scotophaeus anahita Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="B5C9F25A-2966-43AF-91AF-994A6B783F96" docUuidSource="ZooBank" docVersion="8" lastPageNumber="99" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="98" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DA9CDC309E0ABFA8BF5E698ABD047816">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="D0A68497379C52B20C13850E8165A13D">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="50EA63863231CFDC9A454E848B8C5020">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
<mods:nameIdentifier id="53770FBF1BAE96CB95EA784957A11F67" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0B171C772305C17D29B948D502773E1A">Chatzaki, Maria</mods:namePart>
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<mods:nameIdentifier id="A1924113825018BC74C565B2F31E54EB" type="email">maria.chatzaki@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FD5A8668C71D6864F606CFE1E7667AB0">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="DFE363CBBA21AC5C63DBF219EE1DB0F8">Sergei.</mods:affiliation>
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<mods:namePart id="F84A44A648FCBC3A7E8A1B1699A94A6E">Marusik, Yuri M.</mods:namePart>
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<mods:date id="B67FF49F433DB5CD5EB6C7E6AF08F1A9">2021</mods:date>
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<treatment id="03F987850361EF7E1C5E7D81FC1D6F87" ID-DOI="http://doi.org/10.5281/zenodo.4906490" ID-GBIF-Taxon="182597852" ID-Zenodo-Dep="4906490" LSID="urn:lsid:zoobank.org:act:B5C9F25A-2966-43AF-91AF-994A6B783F96" httpUri="http://treatment.plazi.org/id/03F987850361EF7E1C5E7D81FC1D6F87" lastPageId="31" lastPageNumber="99" pageId="30" pageNumber="98">
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<paragraph id="8BEF36930361EF7F1C5E7D81FC0B6E4C" blockId="30.[369,1218,754,852]" box="[618,970,754,781]" pageId="30" pageNumber="98">
<heading id="D0A781FF0361EF7F1C5E7D81FC0B6E4C" box="[618,970,754,781]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="98" reason="2">
<taxonomicName id="4C504D100361EF7F1C5E7D81FCA96E4C" ID-CoL="98YVM" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[618,872,754,781]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="98" phylum="Arthropoda" rank="species" species="anahita" status="sp. nov.">
<emphasis id="B924EA810361EF7F1C5E7D81FCA96E4C" bold="true" box="[618,872,754,781]" italics="true" pageId="30" pageNumber="98">Scotophaeus anahita</emphasis>
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<taxonomicNameLabel id="A21757FA0361EF7F1D5B7D80FC0B6E4C" box="[879,970,755,781]" pageId="30" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BEF36930361EF7F1C857C49FC426E15" blockId="30.[369,1218,754,852]" box="[689,899,826,852]" pageId="30" pageNumber="98">
<figureCitation id="136B2A160361EF7F1C857C49FC426E15" box="[689,899,826,852]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="30" pageNumber="98">Fig. 19A–C, G–H</figureCitation>
</paragraph>
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<heading id="D0A781FF0361EF7F1E897C0FFEF76ED7" bold="true" box="[189,310,892,918]" fontSize="11" level="3" pageId="30" pageNumber="98" reason="3">
<emphasis id="B924EA810361EF7F1E897C0FFEF76ED7" bold="true" box="[189,310,892,918]" pageId="30" pageNumber="98">Diagnosis</emphasis>
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The two new species of
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<emphasis id="B924EA810361EF7F1FD37CDFFDBF6E87" box="[487,638,940,966]" italics="true" pageId="30" pageNumber="98">Scotophaeus</emphasis>
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described here,
<taxonomicName id="4C504D100361EF7F1D727CDFFC7E6E87" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[838,959,940,966]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="98" phylum="Arthropoda" rank="species" species="anahita" status="sp. nov.">
<emphasis id="B924EA810361EF7F1D727CDFFC926E87" box="[838,851,940,966]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1D507CDFFC7E6E87" box="[868,959,940,966]" italics="true" pageId="30" pageNumber="98">anahita</emphasis>
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and
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<emphasis id="B924EA810361EF7F1A507CDFFBB06E87" box="[1124,1137,940,966]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1AB67CDFFAC96E87" box="[1154,1288,940,966]" italics="true" pageId="30" pageNumber="98">elburzensis</emphasis>
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<taxonomicNameLabel id="A21757FA0361EF7F1B267CDFFAB36E87" box="[1298,1394,940,966]" pageId="30" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, are very similar in the structure of their epigynal sclerotization and somatic characters but are clearly different in the structure of the endogyne. In
<taxonomicName id="4C504D100361EF7F1C8E7C87FCEE694F" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[698,815,1012,1038]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="98" phylum="Arthropoda" rank="species" species="anahita" status="sp. nov.">
<emphasis id="B924EA810361EF7F1C8E7C87FD08694F" box="[698,713,1012,1038]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1CE27C87FCEE694F" box="[726,815,1012,1038]" italics="true" pageId="30" pageNumber="98">anahita</emphasis>
</taxonomicName>
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, the lateral ducts are spiraled (vs relatively straight, slightly curved outward in
<taxonomicName id="4C504D100361EF7F1C527B6BFCC66970" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[614,775,1047,1074]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="98" phylum="Arthropoda" rank="species" species="elburzensis" status="sp. nov.">
<emphasis id="B924EA810361EF7F1C527B6BFDB56973" box="[614,628,1048,1074]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1CB77B64FCC66970" box="[643,775,1047,1073]" italics="true" pageId="30" pageNumber="98">elburzensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0361EF7F1D247B6BFCAC6973" box="[784,877,1048,1074]" pageId="30" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
) and the terminal lobes are smaller than the basal lobes and located anteriorly (vs larger than the basal lobes and located posteriorly in
<taxonomicName id="4C504D100361EF7F1AE17B4FFAB76914" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[1237,1398,1083,1110]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="98" phylum="Arthropoda" rank="species" species="elburzensis" status="sp. nov.">
<emphasis id="B924EA810361EF7F1AE17B4FFB256917" box="[1237,1252,1084,1110]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1AC67B48FAB76914" box="[1266,1398,1083,1109]" italics="true" pageId="30" pageNumber="98">elburzensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0361EF7F1E897B13FED6693B" box="[189,279,1120,1146]" pageId="30" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
) (cf.
<figureCitation id="136B2A160361EF7F1F667B2CFE19693B" box="[338,472,1119,1146]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="30" pageNumber="98">Fig. 19B, E</figureCitation>
). Similarly, they resemble
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<emphasis id="B924EA810361EF7F1D3D7B13FCD9693B" box="[777,792,1120,1146]" italics="true" pageId="30" pageNumber="98">S</emphasis>
.
<emphasis id="B924EA810361EF7F1D127B13FCAD6938" box="[806,876,1120,1145]" italics="true" pageId="30" pageNumber="98">nanus</emphasis>
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Wunderlich,
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in the form of the epigynal atrium, but the configuration of the spermathecal cluster is very different from both of the new species (see
<bibRefCitation id="EFC14B620361EF7F1EC77BD4FE026983" author="Wunderlich J." box="[243,451,1191,1218]" pageId="30" pageNumber="98" pagination="743 - 747" refId="ref18382" refString="Wunderlich J. 1995. Beschreibung einer bisher unbekannten Art der Gattung Scotophaeus Simon aus Osterreich (Arachnida: Araneae: Gnaphosidae). Beitrage zur Araneologie 4 (1994): 743 - 747. [publ. in Dec. 1995]" type="book chapter" year="1995">Wunderlich 1995</bibRefCitation>
: figs 6–8).
</paragraph>
</subSubSection>
<subSubSection id="C34A65180361EF7F1E897B9AFD7A6817" pageId="30" pageNumber="98" type="etymology">
<paragraph id="8BEF36930361EF7F1E897B9AFE856842" blockId="30.[189,324,1257,1283]" box="[189,324,1257,1283]" pageId="30" pageNumber="98">
<heading id="D0A781FF0361EF7F1E897B9AFE856842" bold="true" box="[189,324,1257,1283]" fontSize="11" level="3" pageId="30" pageNumber="98" reason="3">
<emphasis id="B924EA810361EF7F1E897B9AFE856842" bold="true" box="[189,324,1257,1283]" pageId="30" pageNumber="98">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930361EF7F1E897A6BFD7A6817" blockId="30.[189,1399,1304,1367]" pageId="30" pageNumber="98">The specific epithet is a noun in apposition and refers to the divinity of the Waters (associated with fertility, healing and wisdom) in Persian mythology.</paragraph>
</subSubSection>
<subSubSection id="C34A65180361EF7F1E897A0CFE9A6B75" pageId="30" pageNumber="98" type="materials_examined">
<paragraph id="8BEF36930361EF7F1E897A0CFEAD68D9" blockId="30.[189,364,1406,1433]" box="[189,364,1406,1433]" pageId="30" pageNumber="98">
<heading id="D0A781FF0361EF7F1E897A0CFEAD68D9" bold="true" box="[189,364,1406,1433]" fontSize="11" level="3" pageId="30" pageNumber="98" reason="3">
<emphasis id="B924EA810361EF7F1E897A0CFEAD68D9" bold="true" box="[189,364,1406,1433]" pageId="30" pageNumber="98">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930361EF7F1EDE7ADDFE9A6B75" blockId="30.[189,1399,1454,1588]" pageId="30" pageNumber="98">
<materialsCitation id="3B383CCE0361EF7F1EDE7ADDFE9A6B75" ID-GBIF-Occurrence="3127434301" collectingDate="2016-05-19" collectionCode="ZMMU" collectorName="P. Ponel" country="Iran" county="Qamsar" elevation="2710" latitude="33.616665" location="Barzok Protected Area" longLatPrecision="1204" longitude="51.316666" municipality="Barzok Protected Area" pageId="30" pageNumber="98" specimenCount="1" specimenCount-female="1" stateProvince="Isfahan Province" typeStatus="holotype">
<emphasis id="B924EA810361EF7F1EDE7ADDFE9D6889" bold="true" box="[234,348,1454,1480]" pageId="30" pageNumber="98">
<typeStatus id="54EB88310361EF7F1EDE7ADDFE9D6889" box="[234,348,1454,1480]" pageId="30" pageNumber="98" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34776030361EF7F1E897AA1FEC968AD" box="[189,264,1490,1516]" name="Iran" pageId="30" pageNumber="98">IRAN</collectingCountry>
•
<specimenCount id="9D56FD1A0361EF7F1F117AA1FEFD68AD" box="[293,316,1490,1516]" pageId="30" pageNumber="98" type="female">♀</specimenCount>
;
<collectingRegion id="4994F8710361EF7F1F787AA1FDD768AD" box="[332,534,1490,1516]" country="Iran" name="Esfahan" pageId="30" pageNumber="98">Isfahan Province</collectingRegion>
,
<collectingCounty id="628E4E1F0361EF7F1C107AA1FDBE68AD" box="[548,639,1490,1516]" pageId="30" pageNumber="98">Qamsar</collectingCounty>
&amp;
<collectingMunicipality id="6B8BACE90361EF7F1C9C7AA1FC7668AD" box="[680,951,1490,1516]" pageId="30" pageNumber="98">Barzok Protected Area</collectingMunicipality>
,
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<quantity id="4CA89B760361EF7F1DF17AA1FBD068AD" box="[965,1041,1490,1517]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="30" pageNumber="98" unit="km" value="55.0">55 km</quantity>
SW of Qamsar
</locationDeviation>
,
<locationDeviation id="9A5D2FB10361EF7F1AED7AA1FED76B51" location="Kamoo" pageId="30" pageNumber="98">
<quantity id="4CA89B760361EF7F1AED7AA1FAE468AD" box="[1241,1317,1490,1516]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="30" pageNumber="98" unit="km" value="14.0">14 km</quantity>
NE of Kamoo
</locationDeviation>
,
<locationDeviation id="9A5D2FB10361EF7F1F157A85FD146B51" box="[289,725,1526,1552]" pageId="30" pageNumber="98">near the road of Gargash observatory</locationDeviation>
;
<geoCoordinate id="EE6450540361EF7F1CD47A87FC8A6B51" box="[736,843,1524,1552]" degrees="33" direction="north" minutes="37" orientation="latitude" pageId="30" pageNumber="98" precision="925" value="33.616665">33°37′ N</geoCoordinate>
,
<geoCoordinate id="EE6450540361EF7F1D627A87FC7E6B51" box="[854,959,1524,1552]" degrees="51" direction="east" minutes="19" orientation="longitude" pageId="30" pageNumber="98" precision="925" value="51.316666">51°19′ E</geoCoordinate>
;
<elevation id="007DD1A00361EF7F1DFF7A85FBA36B51" box="[971,1122,1526,1552]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="30" pageNumber="98" unit="m" value="2710.0">
<quantity id="4CA89B760361EF7F1DFF7A85FBE26B51" box="[971,1059,1526,1552]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="30" pageNumber="98" unit="m" value="2710.0">2710 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEE10530361EF7F1A5B7A85FACC6B51" box="[1135,1293,1526,1552]" pageId="30" pageNumber="98" value="2016-05-19">
<collectingDate id="EFAAE9BB0361EF7F1A5B7A85FACC6B51" box="[1135,1293,1526,1552]" pageId="30" pageNumber="98" value="2016-05-19">19 May 2016</collectingDate>
</date>
;
<collectorName id="26A553450361EF7F1B2D7A85FAB66B51" box="[1305,1399,1526,1552]" pageId="30" pageNumber="98">P. Ponel</collectorName>
leg.;
<collectionCode id="ED41AE560361EF7F1ECC7969FE966B75" box="[248,343,1562,1588]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="30" pageNumber="98" type="Museum">ZMMU</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A65180361EF7E1E89792FFEF86F17" lastPageId="31" lastPageNumber="99" pageId="30" pageNumber="98" type="description">
<paragraph id="8BEF36930361EF7F1E89792FFE8F6B37" blockId="30.[189,334,1628,1654]" box="[189,334,1628,1654]" pageId="30" pageNumber="98">
<heading id="D0A781FF0361EF7F1E89792FFE8F6B37" bold="true" box="[189,334,1628,1654]" fontSize="11" level="3" pageId="30" pageNumber="98" reason="3">
<emphasis id="B924EA810361EF7F1E89792FFE8F6B37" bold="true" box="[189,334,1628,1654]" pageId="30" pageNumber="98">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930361EF7F1EDE79F8FE846BE4" blockId="30.[187,1400,1675,2025]" box="[234,325,1675,1701]" pageId="30" pageNumber="98">
<emphasis id="B924EA810361EF7F1EDE79F8FE846BE4" bold="true" box="[234,325,1675,1701]" pageId="30" pageNumber="98">Female</emphasis>
</paragraph>
<paragraph id="8BEF36930361EF7E1E8979DCFCAC6C07" blockId="30.[187,1400,1675,2025]" lastBlockId="31.[189,1399,265,327]" lastPageId="31" lastPageNumber="99" pageId="30" pageNumber="98">
Total length 11.7. Carapace 5.90 long, 3.80 wide. Eye sizes and interdistance of PMEs: AME = 0.18, ALE = 0.18, PME = 0.20, PLE = 0.20, PME–PME 0.15. Carapace yellowish brown, covered with dark, short setae. Chelicera (
<figureCitation id="136B2A160361EF7F1FF67984FDEB6A50" box="[450,554,1783,1809]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="30" pageNumber="98">Fig. 19G</figureCitation>
) dark brown (almost black), with 3 promarginal teeth. Sternum yellowish brown with dark border, covered with black setae. Maxillae and labium dark brown with white tip. Maxillae longer than wide, with medial indentation on the external edge. Legs yellowish brown with lighter femora and darker brownish metatarsi and tarsi. Abdomen gray, covered with white plumose hairs and dark setae dorsally, and yellow with dark setae ventrally. Spinnerets yellowish brown and uniform in color. Measurements of legs: I: 14.90 (4.18, 2.43, 3.48, 2.83, 2.00), II: 13.95 (3.93, 2.28, 3.18, 2.75, 1.83), III: 13.45 (3.75, 1.90, 2.95, 3.00, 1.85), IV: 18.23 (4.85, 2.20, 4.08, 5.05, 2.05). Spination: I: Fe: 2d, 1pl; Ti: 4v; Mt: 2v. II: Fe: 2d, 2pl; Ti: 4v; Mt: 2v. III: Fe: 2d, 4pl, 2rl; Ti: 1d, 3pl, 3rl, 6v; Mt: 5pl, 6rl, 9v. IV: Fe: 3d, 2pl, 2rl; Ti: 2d, 3pl, 3rl, 6v; Mt: 6pl, 6rl, 6v.
</paragraph>
<paragraph id="8BEF36930360EF7E1E897E1DFDBC6CB0" blockId="31.[189,1399,366,497]" pageId="31" pageNumber="99">
Epigyne as in
<figureCitation id="136B2A160360EF7E1F567E1DFDD26CC9" box="[354,531,366,392]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[189,1393,771,1817]" captionTargetId="figure-136@31.[189,1398,771,1819]" captionTargetPageId="31" captionText="Fig. 19. A–C, G–H. Scotophaeus anahita sp. nov., holotype, ♀ (ZMMU). A, H. Epigyne, ventral view. B–C. Epigyne, dorsal view. G. Chelicera, posterior view. – D–F, I. Scotophaeus elburzensis sp. nov., ♀ (D–E = specimen from Tehran Province, SMF; F, I = holotype, ZMMU). D, I. Epigyne, ventral view. E–F. Epigyne, dorsal view. Abbreviations: Bl = basal lobe; Ld = lateral duct; Lm = lateral margin; rLd = ridge of lateral duct; Tl = terminal lobe. Scale bars: A, C, F, H–I = 0.1 mm; D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4906061" httpUri="https://zenodo.org/record/4906061/files/figure.png" pageId="31" pageNumber="99">Fig. 19A–C, H</figureCitation>
. Epigynal plate almost as long as wide, with transverse atrium posteriorly; atrium with chitinized ʻridgeʼ of the lateral ducts (
<emphasis id="B924EA810360EF7E1D367EE1FCF26CEA" box="[770,819,402,427]" italics="true" pageId="31" pageNumber="99">rLD</emphasis>
) posteriorly and bordered by lateral margin (
<emphasis id="B924EA810360EF7E1B0B7EE1FAA56CEA" box="[1343,1380,402,427]" italics="true" pageId="31" pageNumber="99">Lm</emphasis>
); lateral ducts (
<emphasis id="B924EA810360EF7E1F6F7EC7FEBB6C8F" box="[347,378,436,462]" italics="true" pageId="31" pageNumber="99">Ld</emphasis>
) wide and spiraled; each receptacle with two pairs of concentric sacs (
<emphasis id="B924EA810360EF7E1A997EC7FB066C8F" box="[1197,1223,436,462]" italics="true" pageId="31" pageNumber="99">Bl</emphasis>
and
<emphasis id="B924EA810360EF7E1ACA7EC7FAD76C8F" box="[1278,1302,436,462]" italics="true" pageId="31" pageNumber="99">Tl</emphasis>
); dorsal blind tubes arise from the second pair.
</paragraph>
<paragraph id="8BEF36930360EF7E1EDE7D6AFEEB6F72" blockId="31.[189,313,537,598]" box="[234,298,537,563]" pageId="31" pageNumber="99">
<emphasis id="B924EA810360EF7E1EDE7D6AFEEB6F72" bold="true" box="[234,298,537,563]" pageId="31" pageNumber="99">Male</emphasis>
</paragraph>
<paragraph id="8BEF36930360EF7E1E897D4FFEF86F17" blockId="31.[189,313,537,598]" box="[189,313,572,598]" pageId="31" pageNumber="99">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A65180360EF7E1E897D0EFC1D6F87" box="[189,988,637,710]" pageId="31" pageNumber="99" type="distribution">
<paragraph id="8BEF36930360EF7E1E897D0EFE976FD6" blockId="31.[189,342,637,663]" box="[189,342,637,663]" pageId="31" pageNumber="99">
<heading id="D0A781FF0360EF7E1E897D0EFE976FD6" bold="true" box="[189,342,637,663]" fontSize="11" level="3" pageId="31" pageNumber="99" reason="3">
<emphasis id="B924EA810360EF7E1E897D0EFE976FD6" bold="true" box="[189,342,637,663]" pageId="31" pageNumber="99">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930360EF7E1E897DDFFC1D6F87" blockId="31.[189,988,684,710]" box="[189,988,684,710]" pageId="31" pageNumber="99">
Known only from the
<typeStatus id="54EB88310360EF7E1FF67DDFFE346F87" box="[450,501,684,710]" pageId="31" pageNumber="99">type</typeStatus>
locality in
<collectingRegion id="4994F8710360EF7E1C4D7DDFFC836F87" box="[633,834,684,710]" country="Iran" name="Esfahan" pageId="31" pageNumber="99">Isfahan Province</collectingRegion>
, central
<collectingCountry id="F34776030360EF7E1D917DDFFC196F87" box="[933,984,684,710]" name="Iran" pageId="31" pageNumber="99">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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