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<document id="C8A1FFC29FB29F58710E4A478DC15B26" ID-DOI="10.11646/zootaxa.3640.3.9" ID-GBIF-Dataset="d7cead4e-7c14-426f-bb28-e452e3a2a9e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="224176" ID-ZooBank="0BD5E6B0-0F43-4719-8A41-EFA23A2C04E3" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460594132249" checkinUser="plazi" docAuthor="Wang, Yang, Wei, Cong &amp; Zhang, Yalin" docDate="2013" docId="03C4878FFFF8FF9512A0E3A5149FE17B" docLanguage="en" docName="zt03640p478.pdf" docOrigin="Zootaxa 3640 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diramus nigromaculatus Wang &amp; Zhang, sp. nov." docType="treatment" docVersion="8" lastPageNumber="476" masterDocId="FFFDFFF7FFFBFF961237E1741749E42B" masterDocTitle="Diramus, a new genus of the leafhopper subfamily Evacanthinae (Hemiptera: Cicadellidae), with description of three new species from Thailand" masterLastPageNumber="478" masterPageNumber="473" pageNumber="476" updateTime="1698326454364" updateUser="plazi">
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<mods:title id="3838722E8C422C1EF7A2B68F4D3D155B">Diramus, a new genus of the leafhopper subfamily Evacanthinae (Hemiptera: Cicadellidae), with description of three new species from Thailand</mods:title>
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<mods:namePart id="E0076FBF503C71A99AA9ADFE4E8825B5">Wang, Yang</mods:namePart>
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<mods:namePart id="F8D0290B9BACC47B31D702D44EBD8FFF">Wei, Cong</mods:namePart>
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<mods:namePart id="D3256DCB7953A1D2720AF6ABFD6FF0AA">Zhang, Yalin</mods:namePart>
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<mods:title id="707E38EE1E57D705BE0835A5DCF0784E">Zootaxa</mods:title>
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<mods:date id="2A1DDDC6C9E36E99519550570D724C16">2013</mods:date>
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<mods:identifier id="2235EF9E7D9933DC203F63EE62E7AB32" type="GBIF-Dataset">d7cead4e-7c14-426f-bb28-e452e3a2a9e1</mods:identifier>
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<treatment id="03C4878FFFF8FF9512A0E3A5149FE17B" ID-DOI="http://doi.org/10.5281/zenodo.6152615" ID-GBIF-Taxon="119566339" ID-Zenodo-Dep="6152615" LSID="urn:lsid:plazi:treatment:03C4878FFFF8FF9512A0E3A5149FE17B" httpUri="http://treatment.plazi.org/id/03C4878FFFF8FF9512A0E3A5149FE17B" lastPageNumber="476" pageId="3" pageNumber="476">
<subSubSection id="C3776512FFF8FF9512A0E3A516CDE727" pageId="3" pageNumber="476" type="nomenclature">
<paragraph id="8BD23699FFF8FF9512A0E3A515B5E6C0" blockId="3.[151,764,721,780]" box="[151,764,721,747]" pageId="3" pageNumber="476">
<heading id="D09A81F5FFF8FF9512A0E3A515B5E6C0" bold="true" box="[151,764,721,747]" fontSize="11" level="1" pageId="3" pageNumber="476" reason="1">
<emphasis id="B919EA8BFFF8FF9512A0E3A515B5E6C0" bold="true" box="[151,764,721,747]" pageId="3" pageNumber="476">
<taxonomicName id="4C6D4D1AFFF8FF9512A0E3A515D9E6C0" ID-CoL="8JRR7" authority="Wang &amp; Zhang" authorityName="Wang &amp; Zhang" box="[151,656,721,747]" class="Insecta" family="Cicadellidae" genus="Diramus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="nigromaculatus" status="sp. nov.">
<emphasis id="B919EA8BFFF8FF9512A0E3A5168FE6C0" bold="true" box="[151,454,721,747]" italics="true" pageId="3" pageNumber="476">Diramus nigromaculatus</emphasis>
Wang &amp; Zhang
</taxonomicName>
,
<taxonomicNameLabel id="A22A57F0FFF8FF9510A9E3A515B5E6C0" box="[670,764,721,747]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD23699FFF8FF9512A0E38016CDE727" blockId="3.[151,764,721,780]" box="[151,388,756,780]" pageId="3" pageNumber="476">
<figureCitation id="13562A1CFFF8FF9512A0E3801641E727" box="[151,264,756,780]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[185,1380,421,1974]" captionTargetId="figure@2.[151,1436,385,1981]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Male, habitus, face, head and forewing. 1 – 5: D. khaokus sp. nov.; 6 – 10: D. nigromaculatus sp. nov.; 11 – 15: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224177/files/figure.png" pageId="3" pageNumber="476">Figs 6–10</figureCitation>
,
<figureCitation id="13562A1CFFF8FF951323E3801614E727" box="[276,349,756,780]" captionStart="FIGURES 16 – 30" captionStartId="4.[151,264,1242,1265]" captionTargetBox="[175,1409,197,1205]" captionTargetId="figure@4.[151,1436,175,1218]" captionTargetPageId="4" captionText="FIGURES 16 – 30. Male genitalia. 16 – 20: D. khaokus sp. nov.; 21 – 25: D. nigromaculatus sp. nov.; 26 – 30: D. uncatus sp. nov.; 16, 21, 26: male pygofer, lateral view; 17, 22, 27: aedeagus, lateral view; 18, 23, 28: aedeagus, ventral view; 19, 24, 29: connective and style, dorsal view; 20, 25, 30: subgenital plate, ventral view." httpUri="https://zenodo.org/record/224178/files/figure.png" pageId="3" pageNumber="476">21–25</figureCitation>
,
<figureCitation id="13562A1CFFF8FF95135FE38016CDE727" box="[360,388,756,780]" captionStart="FIGURES 31 – 33" captionStartId="4.[151,264,1729,1752]" captionTargetBox="[184,1339,1372,1696]" captionTargetId="figure@4.[151,1436,1359,1724]" captionTargetPageId="4" captionText="FIGURES 31 – 33. Male pygofer side. 31: D. khaokus sp. nov.; 32: D. nigromaculatus sp. nov.; 33: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224179/files/figure.png" pageId="3" pageNumber="476">32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776512FFF8FF9512A0E24914D5E005" pageId="3" pageNumber="476" type="description">
<paragraph id="8BD23699FFF8FF9512A0E249169AE7EA" blockId="3.[151,1437,829,1361]" pageId="3" pageNumber="476">
<emphasis id="B919EA8BFFF8FF9512A0E2491660E77D" bold="true" box="[151,297,829,854]" pageId="3" pageNumber="476">Description.</emphasis>
Body length (including forewing): ɗ, 5.4–5.6mm. Vertex (
<figureCitation id="13562A1CFFF8FF9511F0E2491342E77E" box="[967,1035,829,853]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[185,1380,421,1974]" captionTargetId="figure@2.[151,1436,385,1981]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Male, habitus, face, head and forewing. 1 – 5: D. khaokus sp. nov.; 6 – 10: D. nigromaculatus sp. nov.; 11 – 15: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224177/files/figure.png" pageId="3" pageNumber="476">Fig. 9</figureCitation>
) mostly white, with irregular black patch on middle part and posterior margin; clypellus (
<figureCitation id="13562A1CFFF8FF951120E2151417E752" box="[791,862,865,889]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[185,1380,421,1974]" captionTargetId="figure@2.[151,1436,385,1981]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Male, habitus, face, head and forewing. 1 – 5: D. khaokus sp. nov.; 6 – 10: D. nigromaculatus sp. nov.; 11 – 15: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224177/files/figure.png" pageId="3" pageNumber="476">Fig. 8</figureCitation>
) dark brown, with off-white near apical margin. Forewing (
<figureCitation id="13562A1CFFF8FF951324E2F21621E7B5" box="[275,360,902,926]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[185,1380,421,1974]" captionTargetId="figure@2.[151,1436,385,1981]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Male, habitus, face, head and forewing. 1 – 5: D. khaokus sp. nov.; 6 – 10: D. nigromaculatus sp. nov.; 11 – 15: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224177/files/figure.png" pageId="3" pageNumber="476">Fig. 10</figureCitation>
), mostly nearly dark ochreous, with large white patch near costal margin; clavus marked with large yellowish green patch.
</paragraph>
<paragraph id="8BD23699FFF8FF9512F0E2BA14D5E005" blockId="3.[151,1437,829,1361]" pageId="3" pageNumber="476">
Male pygofer (
<figureCitation id="13562A1CFFF8FF951341E2BA1691E7CD" box="[374,472,974,998]" captionStart="FIGURES 16 – 30" captionStartId="4.[151,264,1242,1265]" captionTargetBox="[175,1409,197,1205]" captionTargetId="figure@4.[151,1436,175,1218]" captionTargetPageId="4" captionText="FIGURES 16 – 30. Male genitalia. 16 – 20: D. khaokus sp. nov.; 21 – 25: D. nigromaculatus sp. nov.; 26 – 30: D. uncatus sp. nov.; 16, 21, 26: male pygofer, lateral view; 17, 22, 27: aedeagus, lateral view; 18, 23, 28: aedeagus, ventral view; 19, 24, 29: connective and style, dorsal view; 20, 25, 30: subgenital plate, ventral view." httpUri="https://zenodo.org/record/224178/files/figure.png" pageId="3" pageNumber="476">Figs. 21</figureCitation>
,
<figureCitation id="13562A1CFFF8FF9513DFE2BA154CE7CD" box="[488,517,974,998]" captionStart="FIGURES 31 – 33" captionStartId="4.[151,264,1729,1752]" captionTargetBox="[184,1339,1372,1696]" captionTargetId="figure@4.[151,1436,1359,1724]" captionTargetPageId="4" captionText="FIGURES 31 – 33. Male pygofer side. 31: D. khaokus sp. nov.; 32: D. nigromaculatus sp. nov.; 33: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224179/files/figure.png" pageId="3" pageNumber="476">32</figureCitation>
) with short, slender ventral process arising near base, just reaching middle of ventral margin. Aedeagus (
<figureCitation id="13562A1CFFF8FF9513FCE2851507E022" box="[459,590,1009,1033]" captionStart="FIGURES 16 – 30" captionStartId="4.[151,264,1242,1265]" captionTargetBox="[175,1409,197,1205]" captionTargetId="figure@4.[151,1436,175,1218]" captionTargetPageId="4" captionText="FIGURES 16 – 30. Male genitalia. 16 – 20: D. khaokus sp. nov.; 21 – 25: D. nigromaculatus sp. nov.; 26 – 30: D. uncatus sp. nov.; 16, 21, 26: male pygofer, lateral view; 17, 22, 27: aedeagus, lateral view; 18, 23, 28: aedeagus, ventral view; 19, 24, 29: connective and style, dorsal view; 20, 25, 30: subgenital plate, ventral view." httpUri="https://zenodo.org/record/224178/files/figure.png" pageId="3" pageNumber="476">Figs 23, 24</figureCitation>
) with paired apical processes extending laterally but somewhat straight in ventral view. Other characteristics as shown in
<figureCitation id="13562A1CFFF8FF951090E5621450E005" box="[679,793,1046,1070]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1975,1998]" captionTargetBox="[185,1380,421,1974]" captionTargetId="figure@2.[151,1436,385,1981]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Male, habitus, face, head and forewing. 1 – 5: D. khaokus sp. nov.; 6 – 10: D. nigromaculatus sp. nov.; 11 – 15: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224177/files/figure.png" pageId="3" pageNumber="476">Figs 6–10</figureCitation>
,
<figureCitation id="13562A1CFFF8FF951113E5621424E005" box="[804,877,1046,1070]" captionStart="FIGURES 16 – 30" captionStartId="4.[151,264,1242,1265]" captionTargetBox="[175,1409,197,1205]" captionTargetId="figure@4.[151,1436,175,1218]" captionTargetPageId="4" captionText="FIGURES 16 – 30. Male genitalia. 16 – 20: D. khaokus sp. nov.; 21 – 25: D. nigromaculatus sp. nov.; 26 – 30: D. uncatus sp. nov.; 16, 21, 26: male pygofer, lateral view; 17, 22, 27: aedeagus, lateral view; 18, 23, 28: aedeagus, ventral view; 19, 24, 29: connective and style, dorsal view; 20, 25, 30: subgenital plate, ventral view." httpUri="https://zenodo.org/record/224178/files/figure.png" pageId="3" pageNumber="476">21–25</figureCitation>
,
<figureCitation id="13562A1CFFF8FF95114FE56214DEE005" box="[888,919,1046,1070]" captionStart="FIGURES 31 – 33" captionStartId="4.[151,264,1729,1752]" captionTargetBox="[184,1339,1372,1696]" captionTargetId="figure@4.[151,1436,1359,1724]" captionTargetPageId="4" captionText="FIGURES 31 – 33. Male pygofer side. 31: D. khaokus sp. nov.; 32: D. nigromaculatus sp. nov.; 33: D. uncatus sp. nov." httpUri="https://zenodo.org/record/224179/files/figure.png" pageId="3" pageNumber="476">32</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3776512FFF8FF9512F0E54F13DEE0B7" pageId="3" pageNumber="476" type="materials_examined">
<paragraph id="8BD23699FFF8FF9512F0E54F13DEE0B7" blockId="3.[151,1437,829,1361]" pageId="3" pageNumber="476">
<emphasis id="B919EA8BFFF8FF9512F0E54F1633E078" bold="true" box="[199,378,1082,1107]" pageId="3" pageNumber="476">
<typeStatus id="54D6883BFFF8FF9512F0E54F1649E078" box="[199,256,1083,1107]" pageId="3" pageNumber="476">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F37A7609FFF8FF9513BDE54F1556E078" box="[394,543,1083,1107]" name="Thailand" pageId="3" pageNumber="476">THAILAND</collectingCountry>
:
<typeStatus id="54D6883BFFF8FF951006E54E15D0E079" box="[561,665,1082,1106]" pageId="3" pageNumber="476" type="holotype">Holotype</typeStatus>
ɗ (QSBG), Loei, Phu Kradueng,
<quantity id="4C959B7CFFF8FF951676E54E13CDE079" box="[1089,1156,1082,1106]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="476" unit="m" value="420.0">420m</quantity>
,
<date id="FFD31059FFF8FF9516A1E54E1211E079" box="[1174,1368,1082,1106]" pageId="3" pageNumber="476" value="2008-06-26" valueMax="2008-07-01" valueMin="2008-06-26">26.vi.–1.vii.2008</date>
, coll. Thonghuay.
<typeStatus id="54D6883BFFF8FF951315E51516DAE052" box="[290,403,1121,1145]" pageId="3" pageNumber="476" type="paratype">Paratypes</typeStatus>
: 1ɗ (UIUC), Phetchabun, Nam Nao,
<quantity id="4C959B7CFFF8FF951102E514143EE053" box="[821,887,1120,1144]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="3" pageNumber="476" unit="m" value="868.0">868m</quantity>
,
<date id="FFD31059FFF8FF9511B5E5141358E053" box="[898,1041,1120,1144]" pageId="3" pageNumber="476" value="2006-10-09" valueMax="2006-10-16" valueMin="2006-10-09">9–16.x.2006</date>
, coll. Leng Janteab; 1ɗ (NWAFU), Phetchabun, Khao Kho,
<quantity id="4C959B7CFFF8FF95139EE5F016A5E0B7" box="[425,492,1156,1180]" metricMagnitude="2" metricUnit="m" metricValue="2.42" pageId="3" pageNumber="476" unit="m" value="242.0">242m</quantity>
,
<date id="FFD31059FFF8FF9513C0E5F015C9E0B7" box="[503,640,1156,1180]" pageId="3" pageNumber="476" value="2006-11-05" valueMax="2006-11-06" valueMin="2006-11-05">5–6.xi.2006</date>
, coll. Somchai Chachumnan &amp; Saink Singtong.
</paragraph>
</subSubSection>
<subSubSection id="C3776512FFF8FF9512F0E5DD155CE107" pageId="3" pageNumber="476" type="discussion">
<paragraph id="8BD23699FFF8FF9512F0E5DD155CE107" blockId="3.[151,1437,829,1361]" pageId="3" pageNumber="476">
<emphasis id="B919EA8BFFF8FF9512F0E5DD1672E0EA" bold="true" box="[199,315,1193,1217]" pageId="3" pageNumber="476">Remarks.</emphasis>
This species can be easily recognized by the following characteristics: 1) vertex predominantly white, marked with a large black patch on disc and at base, respectively; 2) forewing with a large white marking near costal margin; 3) apical processes of aedeagus in ventral view somewhat straight, which is only extended laterally but not curved anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C3776512FFF8FF9512F0E44D149FE17B" box="[199,982,1336,1361]" pageId="3" pageNumber="476" type="etymology">
<paragraph id="8BD23699FFF8FF9512F0E44D149FE17B" blockId="3.[151,1437,829,1361]" box="[199,982,1336,1361]" pageId="3" pageNumber="476">
<emphasis id="B919EA8BFFF8FF9512F0E44D1607E17A" bold="true" box="[199,334,1337,1361]" pageId="3" pageNumber="476">Etymology.</emphasis>
The specific name refers to the black patch on the vertex.
</paragraph>
</subSubSection>
</treatment>
</document>
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