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<document id="D5F8B3C499C1A21FE0DE72747BA8ECAC" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank &amp; Ngiam, Robin W. J." docDate="2017" docId="03C3A90C985DFF9BFF6A88B4FE73E8C3" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus pasia van Tol 1990" docType="treatment" docVersion="8" lastPageNumber="218" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="216" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="77ECD1B450FD9A1A5A034221CB94E863">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="9D90991B036F8D2F20D288EB0709C2CA">Stokvis, Frank</mods:namePart>
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<treatment id="03C3A90C985DFF9BFF6A88B4FE73E8C3" ID-DOI="http://doi.org/10.5281/zenodo.5631250" ID-GBIF-Taxon="137580773" ID-Zenodo-Dep="5631250" LSID="urn:lsid:plazi:treatment:03C3A90C985DFF9BFF6A88B4FE73E8C3" httpUri="http://treatment.plazi.org/id/03C3A90C985DFF9BFF6A88B4FE73E8C3" lastPageId="17" lastPageNumber="218" pageId="15" pageNumber="216">
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<heading id="D09DAF76985DFF85FF6A88B4FDA3EFC8" bold="true" box="[151,574,151,178]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
<taxonomicName id="4C6A6399985DFF85FF6A88B4FDA3EFC8" ID-CoL="3TQ4K" authority="van Tol, 1990" authorityName="van Tol" authorityYear="1990" box="[151,574,151,178]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC408985DFF85FF6A88B4FDA3EFC8" bold="true" box="[151,574,151,178]" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF6A88B4FE0DEFCB" bold="true" box="[151,400,151,177]" italics="true" pageId="15" pageNumber="216">Leptogomphus pasia</emphasis>
van
<bibRefCitation id="EFFB65EB985DFF85FE3088BBFDA3EFC8" author="Tol" box="[461,574,152,178]" pageId="15" pageNumber="216" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol, 1990</bibRefCitation>
</emphasis>
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(
<figureCitation id="1351049F985DFF85FF628899FF78EFA9" box="[159,229,186,211]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="15" pageNumber="216">Figs 1</figureCitation>
,
<figureCitation id="1351049F985DFF85FF0E8898FE9EEFA9" box="[243,259,187,211]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="15" pageNumber="216">2</figureCitation>
,
<figureCitation id="1351049F985DFF85FEF28898FE82EFA9" box="[271,287,187,211]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="15" pageNumber="216">3</figureCitation>
,
<figureCitation id="1351049F985DFF85FED68899FEA6EFA9" box="[299,315,186,211]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="15" pageNumber="216">6</figureCitation>
,
<figureCitation id="1351049F985DFF85FEBA8898FEF8EFA9" box="[327,357,187,211]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="15" pageNumber="216">14</figureCitation>
,
<figureCitation id="1351049F985DFF85FE8C8898FE12EFA9" box="[369,399,187,211]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="15" pageNumber="216">21</figureCitation>
,
<figureCitation id="1351049F985DFF85FE668898FE27EFA9" box="[411,442,187,211]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="15" pageNumber="216">39</figureCitation>
,
<figureCitation id="1351049F985DFF85FE388899FE7EEFA9" box="[453,483,186,211]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="15" pageNumber="216">56</figureCitation>
,
<figureCitation id="1351049F985DFF85FE128899FD90EFA9" box="[495,525,186,211]" captionStart="FIGURES 64–67" captionStartId="35.[151,264,1766,1789]" captionTargetBox="[149,1433,669,1739]" captionTargetId="figure@35.[149,1434,668,1739]" captionTargetPageId="35" captionText="FIGURES 64–67. Male synthorax lateral: (64) L. sii holotype; (65) L. sii teneral male Gunung Pueh; (66) L. pasia SAB12_GOM1; (67) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068030/files/figure.png" pageId="15" pageNumber="216">66</figureCitation>
,
<figureCitation id="1351049F985DFF85FDE48898FDAAEFA9" box="[537,567,187,211]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="15" pageNumber="216">71</figureCitation>
,
<figureCitation id="1351049F985DFF85FDBE8898FDFCEFA9" box="[579,609,187,211]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="15" pageNumber="216">81</figureCitation>
,
<figureCitation id="1351049F985DFF85FD938898FD11EFA9" box="[622,652,187,211]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="15" pageNumber="216">89</figureCitation>
,
<figureCitation id="1351049F985DFF85FD6A8899FD28EFA9" box="[663,693,186,211]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="15" pageNumber="216">95</figureCitation>
,
<figureCitation id="1351049F985DFF85FD3C8898FD70EFA9" box="[705,749,187,211]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="15" pageNumber="216">101</figureCitation>
,
<figureCitation id="1351049F985DFF85FD048898FCB8EFA9" box="[761,805,187,211]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="15" pageNumber="216">107</figureCitation>
,
<figureCitation id="1351049F985DFF85FCCC8898FCC1EFA9" box="[817,860,187,211]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="15" pageNumber="216">113</figureCitation>
,
<figureCitation id="1351049F985DFF85FC958898FC09EFA9" box="[872,916,187,211]" captionStart="FIGURES 118–119" captionStartId="45.[151,264,604,627]" captionTargetBox="[156,1432,193,583]" captionTargetId="figure@45.[156,1432,193,583]" captionTargetPageId="45" captionText="FIGURES 118–119. Ventral view female abdominal S8 (all scale bars 0.5mm): (118) L. coomansi SAR11_12_GOM48; (119) L. pasia SAB12_GOM2." httpUri="https://zenodo.org/record/1068050/files/figure.png" pageId="15" pageNumber="216">119</figureCitation>
,
<figureCitation id="1351049F985DFF85FC628898FC56EFA9" box="[927,971,187,211]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="15" pageNumber="216">121</figureCitation>
,
<figureCitation id="1351049F985DFF85FC2A8898FB9EEFA9" box="[983,1027,187,211]" captionStart="FIGURE 127" captionStartId="49.[151,250,1596,1618]" captionTargetBox="[151,1436,193,1574]" captionTargetId="figure@49.[151,1436,193,1574]" captionTargetPageId="49" captionText="FIGURE 127. Distribution of L. pasia (white circle) and L. sii (blacksquare)." httpUri="https://zenodo.org/record/1068056/files/figure.png" pageId="15" pageNumber="216">127</figureCitation>
,
<figureCitation id="1351049F985DFF85FBF28898FBA7EFA9" box="[1039,1082,187,211]" captionStart="FIGURES 130–132" captionStartId="52.[151,264,1763,1786]" captionTargetBox="[453,1133,193,1742]" captionTargetId="figure@52.[453,1133,193,1742]" captionTargetPageId="52" captionText="FIGURES 130–132. Habitus of some Bornean Leptogomphus species:(130) L. pasia holotype (left hand side, flipped to right for consistency with other images); (131) L.sii holotype male; (132) L. williamsoni male SAR13_14_GOM56." httpUri="https://zenodo.org/record/1068062/files/figure.png" pageId="15" pageNumber="216">130</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB7A3F34985DFF85FF6A8921FD25EE4D" pageId="15" pageNumber="216">
<taxonomicName id="4C6A6399985DFF85FF6A8921FDBDEE62" ID-CoL="3TQ4K" authority="van Tol 1990: 97" authorityName="van Tol" authorityPageNumber="97" authorityYear="1990" box="[151,544,258,280]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC408985DFF85FF6A8921FEF7EE62" box="[151,362,258,280]" italics="true" pageId="15" pageNumber="216">Leptogomphus pasia</emphasis>
van
<treatmentCitation id="0ACB3E0B985DFF85FE618921FDBDEE62" author="Tol" box="[412,544,258,280]" page="97" pageId="15" pageNumber="216" year="1990">
<bibRefCitation id="EFFB65EB985DFF85FE618921FE66EE62" author="Tol" box="[412,507,258,280]" pageId="15" pageNumber="216" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
: 97
</treatmentCitation>
</taxonomicName>
–99, 101, 102,
<figureCitation id="1351049F985DFF85FD478921FC8FEE62" box="[698,786,258,280]" captionStart-0="FIGURE1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURES 4–9" captionStartId-0="4.[151,250,1913,1935]" captionStartId-1="5.[151,250,1294,1316]" captionStartId-2="6.[151,250,1871,1893]" captionStartId-3="9.[151,264,1713,1736]" captionTargetBox-0="[168,1423,1134,1886]" captionTargetBox-1="[243,1384,200,1237]" captionTargetBox-2="[209,1373,710,1811]" captionTargetBox-3="[151,1436,193,1692]" captionTargetId-0="figure@4.[151,1436,1127,1891]" captionTargetId-1="figure@5.[151,1436,193,1273]" captionTargetId-2="figure@6.[151,1436,700,1850]" captionTargetId-3="figure@9.[151,1436,193,1692]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="9" captionText-0="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." captionText-1="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." captionText-2="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." captionText-3="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri-0="https://zenodo.org/record/1067998/files/figure.png" httpUri-1="https://zenodo.org/record/1068000/files/figure.png" httpUri-2="https://zenodo.org/record/1068002/files/figure.png" httpUri-3="https://zenodo.org/record/1068004/files/figure.png" pageId="15" pageNumber="216">Figs 1–5</figureCitation>
(part, original description
<typeStatus id="54D1A6B8985DFF85FBD98921FBE0EE62" box="[1060,1149,258,280]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
female Long Pasia,
<collectingRegion id="49AED6F8985DFF85FAB28921FA0DEE62" box="[1359,1424,258,280]" country="Malaysia" name="Sabah" pageId="15" pageNumber="216">Sabah</collectingRegion>
),
<tableCitation id="C6E82DA1985DFF85FF3A8902FE8CEE4D" box="[199,273,289,311]" captionStart="TABLE 2" captionStartId="3.[151,239,256,279]" captionTargetBox="[159,1375,443,1403]" captionText="TABLE 2. Leptogomphus from Borneo used in the molecular analyses; collectors and more details of location given in material lists under each species. Collection codes and BOLD Process IDs are given along with basic location details. The collection codes can be used to locate the sequences on the BOLD website, and also appear as BOLD Sample IDs there." pageId="15" pageNumber="216">Table 2</tableCitation>
(part, female characters);—
<treatmentCitation id="0ACB3E0B985DFF85FDD38902FD29EE4D" author="Orr" box="[558,692,289,311]" page="96" pageId="15" pageNumber="216" year="2003">
<bibRefCitation id="EFFB65EB985DFF85FDD38902FD13EE4D" author="Orr" box="[558,654,289,311]" pageId="15" pageNumber="216" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr 2003</bibRefCitation>
: 96
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3704B91985DFF85FF6A8947FB79EDE6" pageId="15" pageNumber="216" type="materials_examined">
<paragraph id="8BD5181A985DFF85FF6A8947FE06EEBE" blockId="15.[151,1437,356,2037]" pageId="15" pageNumber="216">
<materialsCitation id="3B021247985DFF85FF6A8947FE06EEBE" ID-GBIF-Occurrence="1702634987" collectingDate="1987-04-03" collectionCode="RMNH" collectorName="C. van Actherberg" country="Malaysia" county="Beaufort" elevation="1250" latitude="4.4333" location="Material" longLatPrecision="7" longitude="115.6667" municipality="Long Pasia area" pageId="15" pageNumber="216" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B91EC408985DFF85FF6A8947FD8BEE07" bold="true" box="[151,534,356,381]" pageId="15" pageNumber="216">
<location id="8EB54EC1985DFF85FF6A8947FF62EE07" LSID="urn:lsid:plazi:treatment:03C3A90C985DFF9BFF6A88B4FE73E8C3:8EB54EC1985DFF85FF6A8947FF62EE07" box="[151,255,356,381]" country="Malaysia" county="Beaufort" latitude="4.4333" longLatPrecision="7" longitude="115.6667" municipality="Long Pasia area" name="Material" pageId="15" pageNumber="216" stateProvince="Sabah">Material</location>
studied.
<typeStatus id="54D1A6B8985DFF85FE938947FE34EE07" box="[366,425,356,381]" pageId="15" pageNumber="216">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B91EC408985DFF85FDD88947FD04EE07" bold="true" box="[549,665,356,381]" pageId="15" pageNumber="216">
<typeStatus id="54D1A6B8985DFF85FDD88947FD09EE07" box="[549,660,356,381]" pageId="15" pageNumber="216" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D6CD393985DFF85FD5F8947FD50EE06" box="[674,717,356,380]" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(JvT 16524),
<location id="8EB54EC1985DFF85FC8D8947FB9CEE07" LSID="urn:lsid:plazi:treatment:03C3A90C985DFF9BFF6A88B4FE73E8C3:8EB54EC1985DFF85FC8D8947FB9CEE07" box="[880,1025,356,381]" country="Malaysia" county="Beaufort" latitude="4.4333" longLatPrecision="7" longitude="115.6667" municipality="Long Pasia area" name="Sungai Pasia" pageId="15" pageNumber="216" stateProvince="Sabah">Sungai Pasia</location>
near confluence with
<location id="8EB54EC1985DFF85FB028947FA05EE07" LSID="urn:lsid:plazi:treatment:03C3A90C985DFF9BFF6A88B4FE73E8C3:8EB54EC1985DFF85FB028947FA05EE07" box="[1279,1432,356,381]" country="Malaysia" county="Beaufort" latitude="4.4333" longLatPrecision="7" longitude="115.6667" municipality="Long Pasia area" name="Sungai Maga" pageId="15" pageNumber="216" stateProvince="Sabah">Sungai Maga</location>
,
<collectingMunicipality id="6BB18260985DFF85FF6A89ABFED3EEE5" box="[151,334,391,416]" pageId="15" pageNumber="216">Long Pasia area</collectingMunicipality>
,
<quantity id="4C92B5FF985DFF85FEA789A4FE32EEDA" box="[346,431,391,416]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="216" unit="km" value="105.0">105 km</quantity>
south of
<collectingCounty id="62B46096985DFF85FDE789A4FDE2EEDA" box="[538,639,391,416]" pageId="15" pageNumber="216">Beaufort</collectingCounty>
,
<collectingRegion id="49AED6F8985DFF85FD7689A4FD4FEEDA" box="[651,722,391,416]" country="Malaysia" name="Sabah" pageId="15" pageNumber="216">Sabah</collectingRegion>
,
<collectingCountry id="F37D588A985DFF85FD2389A4FCDAEEDA" box="[734,839,391,416]" name="Malaysia" pageId="15" pageNumber="216">Malaysia</collectingCountry>
,
<geoCoordinate id="EE5E7EDD985DFF85FCAE89ABFC25EEDA" box="[851,952,392,416]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="5" value="4.4333">4.4333N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985DFF85FC3E89A4FBA2EEDA" box="[963,1087,391,416]" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="5" value="115.6667">115.6667E</geoCoordinate>
,
<elevation id="0047FF29985DFF85FBB689A4FB06EEDA" box="[1099,1179,391,416]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="216" unit="m" value="1250.0">
<quantity id="4C92B5FF985DFF85FBB689A4FB06EEDA" box="[1099,1179,391,416]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="216" unit="m" value="1250.0">1250m</quantity>
</elevation>
,
<collectingDate id="EF90C732985DFF85FB5589ABFA8BEEDA" box="[1192,1302,391,416]" pageId="15" pageNumber="216" value="1987-04-03">
<date id="FFD43EDA985DFF85FB5589ABFA8BEEDA" box="[1192,1302,391,416]" pageId="15" pageNumber="216" value="1987-04-03">3 iv 1987</date>
</collectingDate>
, leg.
<collectorName id="269F7DCC985DFF85FAAE89ABFE85EEBF" pageId="15" pageNumber="216">C. van Actherberg</collectorName>
, in
<collectionCode id="ED7B80DF985DFF85FEC2898EFE0AEEBE" box="[319,407,429,452]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="216">RMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD5181A985DFF85FF3A89ECFC6AED76" blockId="15.[151,1437,356,2037]" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF3A89ECFE1CEE92" bold="true" box="[199,385,463,488]" pageId="15" pageNumber="216">Other material</emphasis>
.
<emphasis id="B91EC408985DFF85FE6B89ECFD16EE92" bold="true" box="[406,651,463,488]" pageId="15" pageNumber="216">
<collectingRegion id="49AED6F8985DFF85FE6B89ECFD9CEE92" box="[406,513,463,488]" country="Malaysia" name="Sarawak" pageId="15" pageNumber="216">
<collectingCountry id="F37D588A985DFF85FE6B89ECFD9CEE92" box="[406,513,463,488]" name="Malaysia" pageId="15" pageNumber="216">Sarawak</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F37D588A985DFF85FDE889ECFD1BEE92" box="[533,646,463,488]" name="Malaysia" pageId="15" pageNumber="216">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="9D6CD393985DFF85FD6489F3FD54EE9D" box="[665,713,464,488]" pageId="15" pageNumber="216" type="male">1 ♂</specimenCount>
(teneral), Pa’Gelawat,
<geoCoordinate id="EE5E7EDD985DFF85FC1F89F3FBC8EE92" box="[994,1109,464,488]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="1" value="3.79817">3.79817N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985DFF85FB9A89ECFB6DEE92" box="[1127,1264,463,488]" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="1" value="115.46537">115.46537E</geoCoordinate>
,
<quantity id="4C92B5FF985DFF85FAFF89F3FA05EE92" box="[1282,1432,464,488]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="15" pageNumber="216" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">1100–1300m</quantity>
, Pulong Tau NP, Miri Division,
<date id="FFD43EDA985DFF85FE0989D7FDF2ED76" box="[500,623,500,525]" pageId="15" pageNumber="216" value="2012-04-18">18 iv 2012</date>
, leg. R.A. Dow, in collection Dow.
</paragraph>
<paragraph id="8BD5181A985DFF85FF3A8A34FB79EDE6" blockId="15.[151,1437,356,2037]" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF3A8A34FE0DED4A" bold="true" box="[199,400,535,560]" pageId="15" pageNumber="216">
<collectingRegion id="49AED6F8985DFF85FF3A8A34FE89ED4A" box="[199,276,535,560]" country="Malaysia" name="Sabah" pageId="15" pageNumber="216">Sabah</collectingRegion>
,
<collectingCountry id="F37D588A985DFF85FEDE8A34FE0DED4A" box="[291,400,535,560]" name="Malaysia" pageId="15" pageNumber="216">Malaysia</collectingCountry>
</emphasis>
:
<specimenCount id="9D6CD393985DFF85FE5C8A3BFE53ED55" box="[417,462,536,560]" pageId="15" pageNumber="216" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED7B80DF985DFF85FE1D8A3BFD84ED4A" box="[480,537,536,560]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14076" name="Society of Amateur Botanists" pageId="15" pageNumber="216">SAB</collectionCode>
12_GOM1,
<collectionCode id="ED7B80DF985DFF85FD5E8A34FD61ED4A" box="[675,764,535,560]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="216">RMNH</collectionCode>
.INS.507699, used for description below and illustrations), Kipungit streams, Poring Hot Springs, Mount Kinabalu NP,
<geoCoordinate id="EE5E7EDD985DFF85FCBF8A1FFC3AED2F" box="[834,935,572,597]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="5" value="6.0886">6.0886N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985DFF85FC4F8A1FFBB1ED2F" box="[946,1068,572,597]" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="5" value="116.7675">116.7675E</geoCoordinate>
(Poring),
<date id="FFD43EDA985DFF85FB5D8A1FFA81ED2E" box="[1184,1308,572,597]" pageId="15" pageNumber="216" value="2012-09-11">11 ix 2012</date>
, leg. J. van Tol, in
<collectionCode id="ED7B80DF985DFF85FF148A43FEDCED0D" box="[233,321,608,631]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="216">RMNH</collectionCode>
;
<specimenCount id="9D6CD393985DFF85FEB08A43FEE5ED0D" box="[333,376,608,632]" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED7B80DF985DFF85FE758A7CFE7EED02" box="[392,483,607,632]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="216">RMNH</collectionCode>
.INS.507715), Silau-Silau stream, Park Headquarters area, same national park,
<geoCoordinate id="EE5E7EDD985DFF85FAA38A7CFA05ED02" box="[1374,1432,607,632]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="5555" value="6.0">6.0N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985DFF85FF6A8AA7FE9FEDE7" box="[151,258,644,669]" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="55" value="116.533">116.533E</geoCoordinate>
(approximate coordinates for Silau-Silau stream),
<date id="FFD43EDA985DFF85FCC78AA7FC28EDE6" box="[826,949,644,669]" pageId="15" pageNumber="216" value="2012-09-13">13 ix 2012</date>
, leg. J. van Tol, in
<collectionCode id="ED7B80DF985DFF85FB748AA6FB7CEDE6" box="[1161,1249,645,668]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="216">RMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3704B91985DFF9BFF3A8A84FAA5EBDA" lastPageId="17" lastPageNumber="218" pageId="15" pageNumber="216" type="description">
<paragraph id="8BD5181A985DFF85FF3A8A84FDB3EB7E" blockId="15.[151,1437,356,2037]" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF3A8A84FC1CEDBA" bold="true" box="[199,897,679,704]" pageId="15" pageNumber="216">Description of male (SAB12_GOM1, RMNH.INS.507699).</emphasis>
<emphasis id="B91EC408985DFF85FC748A84FC5BEDBA" box="[905,966,679,704]" italics="true" pageId="15" pageNumber="216">Head</emphasis>
(
<figureCitation id="1351049F985DFF85FC2B8A84FBBDEDBA" box="[982,1056,679,704]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="15" pageNumber="216">Figs 6</figureCitation>
,
<figureCitation id="1351049F985DFF85FBD08A8BFBD1EDBA" box="[1069,1100,680,704]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="15" pageNumber="216">14</figureCitation>
). Median lobe labium brown, lateral lobes mostly pale, hooks dark brown. Labrum black and very dark brown with pair of small, widely separated yellow basal marks. Mandible bases pale, black area above, more extensive on right side than on left. Genae mostly black except adjacent to mandible bases, where paler. Clypeus black with indistinct dark brown area below antefrons and very small paler patch centrally on anteclypeus. Ante- and postfrons not very sharply divided, antefrons mostly black, postfrons largely occupied by transverse yellow stripe, not quite divided centrally, (
<figureCitation id="1351049F985DFF85FA928B7FFF2BECE2" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="15" pageNumber="216">Fig. 14</figureCitation>
). Vertex and occiput black. Pair of rounded transverse tubercles behind lateral ocelli; ocelli whitish, vertex smooth behind tubercles. Occipital plate deepest at eye margins, becoming shallower gradually over most of length, more abruptly centrally where forms “v” shape; raised centrally at rear and slightly along much of rest of rear margin and at posterior corners.
</paragraph>
<paragraph id="8BD5181A985DFF85FF3A8C2CFC18E9AF" blockId="15.[151,1437,356,2037]" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF3A8C2CFE85EB52" box="[199,280,1039,1064]" italics="true" pageId="15" pageNumber="216">Thorax</emphasis>
. Prothorax mostly very dark brown, yellow markings laterally on anterior and middle pronotal lobes, small, very narrowly divided central posterior marking on middle pronotal lobe, posterior pronotal lobe with narrow, faint central longitudinal pale stripe. Middle pronotal lobe with pair of small raised areas bearing tiny tubercles and short setae, placed centrally on dorsum of each half. Synthorax dark brown with pale yellow markings as follows (
<figureCitation id="1351049F985DFF85FE7B8CBCFE41EBC2" box="[390,476,1183,1208]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="15" pageNumber="216">Figs 56</figureCitation>
,
<figureCitation id="1351049F985DFF85FE158CBCFD95EBC2" box="[488,520,1183,1208]" captionStart="FIGURES 64–67" captionStartId="35.[151,264,1766,1789]" captionTargetBox="[149,1433,669,1739]" captionTargetId="figure@35.[149,1434,668,1739]" captionTargetPageId="35" captionText="FIGURES 64–67. Male synthorax lateral: (64) L. sii holotype; (65) L. sii teneral male Gunung Pueh; (66) L. pasia SAB12_GOM1; (67) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068030/files/figure.png" pageId="15" pageNumber="216">66</figureCitation>
): short, narrow mesothoracic collar, broadly divided at middorsal carina, joined to narrow ca linear dorsal thoracic stripes that extend beyond level of apex of antealar crest, slightly convergent towards antealar carina, terminating just short of antealar carina. Narrow, irregular antehumeral stripe, separated from spot below dorsal thoracic stripe on mesepisternum, near antealar carina. Irregular, narrow, stripe running most of length of mesepimeron. Metepisternum with irregular stripe running from metakatepisternum, below spiracle, on left broken near antealar carina, on right broken into four patches (
<figureCitation id="1351049F985DFF85FBE48D77FBF6EA17" box="[1049,1131,1364,1389]" captionStart="FIGURES 64–67" captionStartId="35.[151,264,1766,1789]" captionTargetBox="[149,1433,669,1739]" captionTargetId="figure@35.[149,1434,668,1739]" captionTargetPageId="35" captionText="FIGURES 64–67. Male synthorax lateral: (64) L. sii holotype; (65) L. sii teneral male Gunung Pueh; (66) L. pasia SAB12_GOM1; (67) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068030/files/figure.png" pageId="15" pageNumber="216">Fig. 66</figureCitation>
shows the right hand side, flipped to the left for consistency). Broad stripe on metepimeron, widening toward antealar carina but deeply excised by black on upper margin shortly before antealar carina. Mesokatepisternum yellow to front, metakatepisternum mostly pale, except to rear so narrowly separated from marking on metepisternum. Venter mostly pale with indistinct central brown area near abdomen. Legs (right anterior and middle detached below coxa, right posterior completely removed for DNA extraction) robust and relatively short. Coxae mostly pale with brown areas, remainder mostly brown, becoming paler distally on left anterior tibia and tarsus where claws almost white except brown tips; claws similar on left middle tarsus. Wings: sectors of arculus separated at origin with 5 cross veins up to and at first bifurcation of superior sector in Fw,
<quantity id="4C92B5FF985DFF85FCAE8E57FC1DE9F7" box="[851,896,1652,1677]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="216" unit="in" value="3.0">3 in</quantity>
Hw. Discoidal field with 2 rows of cells from origin, transitioning to three rows before level of nodus in both wings. 16 Ax in Fw, 11 Hw, 11 (left) or 12 (right) Px in Fw, 13 Px in Hw. Pt brown, covering ca 4-5 underlying cells.
</paragraph>
<paragraph id="8BD5181A985DFF9AFF3A8EFCFCA1EEAA" blockId="15.[151,1437,356,2037]" lastBlockId="16.[151,1437,151,789]" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216">
<emphasis id="B91EC408985DFF85FF3A8EFCFEACE982" box="[199,305,1759,1784]" italics="true" pageId="15" pageNumber="216">Abdomen</emphasis>
. Slender after base of S3, expanding moderately from base of S7, maximum width and height reached apical part of S8, then almost constant. Dorsum of S10 with pair of prominent ca conical protuberances basally (
<figureCitation id="1351049F985DFF85FF058F04FEC1E83A" box="[248,348,1831,1856]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="15" pageNumber="216">Fig. 107</figureCitation>
), bearing many small tubercles on posterior faces. Brown with yellow markings as follow: S1 mostly yellow lower laterally, yellow dorsal mark. S2 lateral mark including much of auricle, another lateral mark placed posteriorly, irregular stripe along ventral margin of tergite in posterior ca one-half, narrow yellowish middorsal stripe. S3 with narrow basal lateral marking, S3–7 with very narrow, irregular, middorsal stripe, this also indistinctly present on basal ca one half of S8. S9–10 entirely brown. Cerci (
<figureCitation id="1351049F985DFF85FC0F8F9BFBCAE8AA" box="[1010,1111,1976,2001]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="15" pageNumber="216">Figs 100</figureCitation>
,
<figureCitation id="1351049F985DFF85FB9E8F9BFB13E8AA" box="[1123,1166,1976,2000]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="15" pageNumber="216">107</figureCitation>
) broad at base in lateral view (
<figureCitation id="1351049F985DFF85FF1C8FFFFEDBE88E" box="[225,326,2012,2037]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="15" pageNumber="216">Fig. 107</figureCitation>
), running slightly downwards to tip along upper margin, contracting abruptly on lower margin shortly after base (so basal part appears as ventral spur), then running gently upward to sharp tip; an interior ventral ridge bearing small teeth clearly visible from basal spur to shortly before apex. In dorsal view (
<figureCitation id="1351049F9842FF9AFB48889FFA89EFAE" box="[1205,1300,188,213]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="16" pageNumber="217">Fig. 101</figureCitation>
) broadest at base, outer margin curved in quite sharply in basal ca one-third, then running gently inwards to sharp tip, where turned slightly out; short blunt basal spur on inner margin, then running straight for much of length, turned out towards tips; the inner ventral ridge expands inwards near tips, visible as ca semicircluar expansion in dorsal view. Pale greyish except basally, extreme tips and teeth of ventral ridge, where dark. Epiproct (
<figureCitation id="1351049F9842FF9AFB4B896FFA83EE1E" box="[1206,1310,332,357]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="16" pageNumber="217">Figs 101</figureCitation>
,
<figureCitation id="1351049F9842FF9AFAD3896FFAC5EE1E" box="[1326,1368,332,356]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="16" pageNumber="217">113</figureCitation>
) dark brown, black at apex, only just shorter than cerci, somewhat compressed so shape of deep divide not clear, in lateral view (
<figureCitation id="1351049F9842FF9AFF2289B7FEDCEED6" box="[223,321,404,429]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="16" pageNumber="217">Fig. 107</figureCitation>
) tapering from moderately broad base, ca straight over much of length, upturned at apex, where sharp. In ventral view arms broad, evenly tapering to apices.
</paragraph>
<paragraph id="8BD5181A9842FF9AFF3A89FFFC4EED8A" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="217">
Accessory genitalia as shown in
<figureCitation id="1351049F9842FF9AFDCA89FFFD2CEE8E" box="[567,689,476,501]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="16" pageNumber="217">Figures 89</figureCitation>
,
<figureCitation id="1351049F9842FF9AFD4389FFFD41EE8F" box="[702,732,476,501]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="16" pageNumber="217">95</figureCitation>
, anterior hamule small, tapering to sharp tip, hooked to rear in apical ca one-half, ca two-fifths height of posterior hamule. Posterior hamule very large, directed postero-ventrally, anterior margin curving from base to ca three-quarters length, where running approximately parallel to lower margin of tergite of S2, after this abruptly narrowed and directed ventrally to sharp apex, posterior margin more gently and evenly curved from base to apex, apex turned slightly out. Vesical not fully inflated, low, seemingly simple. Penis (
<figureCitation id="1351049F9842FF9AFEC18AACFE13EDD2" box="[316,398,655,680]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="16" pageNumber="217">Fig. 95</figureCitation>
) with penultimate segment extended to rear of join with terminal segment, bifurcated in this part. Terminal segment not extending far beyond penultimate segment, dorsal margin almost straight in lateral view, apically with short cornua situated at end of ventrally directed stalk.
</paragraph>
<paragraph id="8BD5181A9842FF9AFF3A8ADEFAB1EC6F" blockId="16.[151,1437,151,789]" box="[199,1324,764,789]" pageId="16" pageNumber="217">Measurements (mm). Hw (possibly not fully inflated) 30, abdomen excluding anal appendages 35.5.</paragraph>
<caption id="DF1548929842FF9AFF6A8FAEFE76E8B8" httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="16" pageNumber="217" targetBox="[151,1436,841,1912]" targetPageId="16">
<paragraph id="8BD5181A9842FF9AFF6A8FAEFE76E8B8" blockId="16.[151,1437,1933,1986]" pageId="16" pageNumber="217">
<emphasis id="B91EC4089842FF9AFF6A8FAEFECAE8D9" bold="true" box="[151,343,1933,1956]" pageId="16" pageNumber="217">FIGURES 14–17.</emphasis>
Male head frontal: (14)
<taxonomicName id="4C6A63999842FF9AFD9D8FACFD28E8DE" authorityName="van Tol" authorityYear="1990" box="[608,693,1935,1956]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089842FF9AFD9D8FACFD28E8DE" box="[608,693,1935,1956]" italics="true" pageId="16" pageNumber="217">L. pasia</emphasis>
</taxonomicName>
SAB12_GOM1; (15)
<taxonomicName id="4C6A63999842FF9AFC5E8FACFC44E8DE" authorityName="Dow &amp; Stokvis &amp; Ngiam" authorityYear="2017" box="[931,985,1935,1956]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="sii">
<emphasis id="B91EC4089842FF9AFC5E8FACFC44E8DE" box="[931,985,1935,1956]" italics="true" pageId="16" pageNumber="217">L. sii</emphasis>
</taxonomicName>
holotype; (16)
<taxonomicName id="4C6A63999842FF9AFB7D8FACFB7DE8DE" authorityName="Dow &amp; Stokvis &amp; Ngiam" authorityYear="2017" box="[1152,1248,1934,1956]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="schieli">
<emphasis id="B91EC4089842FF9AFB7D8FACFB7DE8DE" box="[1152,1248,1934,1956]" italics="true" pageId="16" pageNumber="217">L. schieli</emphasis>
</taxonomicName>
holotype; (17)
<taxonomicName id="4C6A63999842FF9AFA7A8FACFE90E8B8" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="williamsoni">
<emphasis id="B91EC4089842FF9AFA7A8FACFE90E8B8" italics="true" pageId="16" pageNumber="217">L. williamsoni</emphasis>
</taxonomicName>
SAR11_12_GOM56.
</paragraph>
</caption>
<paragraph id="8BD5181A9843FF9BFF3A88B4FC7DEE67" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
<emphasis id="B91EC4089843FF9BFF3A88B4FE31EFCA" bold="true" box="[199,428,151,176]" pageId="17" pageNumber="218">Variation in males.</emphasis>
The teneral male from Pa’Gelawat agrees with the male described in most features that can be usefully compared, in particular the cerci, although distorted, have the distinctive basal ventral spur and the interior ventral ridge bearing small teeth; however the semicircular expansion near the tips of the cerci is less evident in this male, although that may be due to the condition of the cerci.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A890BFD92EE1F" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
Measurements (mm). 13–16 Ax in Fw,
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Hw, 13 Px in all wings. Hw (not fully inflated) 29.5, abdomen excluding anal appendages ca 35.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A894CFB4AEED7" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
<emphasis id="B91EC4089843FF9BFF3A894CFDDFEEF2" bold="true" box="[199,578,367,392]" pageId="17" pageNumber="218">Descriptive notes on the female.</emphasis>
A full description of the female
<typeStatus id="54D1A6B89843FF9BFC4F894CFB89EEF2" box="[946,1044,367,392]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
was given by van
<bibRefCitation id="EFFB65EB9843FF9BFB1B894CFAFDEEF2" author="Tol" box="[1254,1376,367,392]" pageId="17" pageNumber="218" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol (1990)</bibRefCitation>
; here we note a few additional points and discuss variation in the female collected on Mount Kinabalu.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8994FB22ED1A" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
The female from Mount Kinabalu has a small mark centrally on the anteclypeus and a pair of small, widely spaced transverse streaks on the postclypeus; van
<bibRefCitation id="EFFB65EB9843FF9BFD2889FFFCD0EE8E" author="Tol" box="[725,845,476,501]" pageId="17" pageNumber="218" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol (1990)</bibRefCitation>
states that the clypeus is black in the
<typeStatus id="54D1A6B89843FF9BFAFB89FFFAF6EE8F" box="[1286,1387,476,501]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
, but there is a small, faint central pale mark on the anteclypeus. Only a single pair of transverse tubercles are present in the female from Mount Kinabalu, the “smaller pair latero-posteriorly”, present in the
<typeStatus id="54D1A6B89843FF9BFBAF8A07FB29ED47" box="[1106,1204,548,573]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
are absent, but there is peculiar shallow groove in the same position; the smaller pair are very small in the
<typeStatus id="54D1A6B89843FF9BFBA88A64FB26ED1A" box="[1109,1211,583,608]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8A4EFC15EDB6" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">In the female from Mount Kinabalu, the antehumeral stripes are broken as they approach the wing bases, leaving a separate pale mark by the antealar carina. It has 18 (right) to 19 (left) Ax in the Fw, 12 (right) to 13 (left) in Hw; 14 (left) to 15 (right) Px in Fw, 14 (right) to 16 (left) in Hw.</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8AF4FB3FEB07" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
The dorsal marking of S1, described as a middorsal stripe in the
<typeStatus id="54D1A6B89843FF9BFC1A8AF4FBD1ED8A" box="[999,1100,727,752]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
, is a broad yellow marking occupying most of the dorsum of the segment in the Mount Kinabalu specimen; the middorsal abdominal markings otherwise agrees with that described for the
<typeStatus id="54D1A6B89843FF9BFD728B3CFD69EC42" box="[655,756,799,824]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
, except that it is only present on the basal ca one quarter of S8. The lateral markings of S2 and S
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the
<typeStatus id="54D1A6B89843FF9BFD728B67FD6CEC27" box="[655,753,836,861]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
are described as “lateral side of segment 2 and antero-lateral side of segment 3 also dark yellow” (van
<bibRefCitation id="EFFB65EB9843FF9BFD808B44FD75ECFA" author="Tol" box="[637,744,871,896]" pageId="17" pageNumber="218" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
), giving the impression that the entire side of S2 and of the anterior part of S3 are yellow; in the Mount Kinabalu specimen these markings are centrally placed lateral longitudinal stripes, very similar to those seen in other species. However examination of the
<typeStatus id="54D1A6B89843FF9BFB458B8CFA87ECB2" box="[1208,1306,943,968]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
reveals the same condition as seen in the Mount Kinabalu specimen, and the expression in the original description is misleading. Additionally there is a small yellow mark at the base of S4 and a tiny one at the base of S5 on the left in the Mount Kinabalu female. The vulvar scale in the Mount Kinabalu female is the same as that of the
<typeStatus id="54D1A6B89843FF9BFACE8C3FFA05EB4F" box="[1331,1432,1052,1077]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
; the sternum of S
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this specimen has a slight swelling in the same position as the distinct tubercle seen in
<taxonomicName id="4C6A63999843FF9BFA788C62FE9EEB06" authorityName="Laidlaw" authorityYear="1936" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC4089843FF9BFA788C62FE9EEB06" italics="true" pageId="17" pageNumber="218">L. coomansi</emphasis>
</taxonomicName>
females. The cerci of the Mount Kinabalu specimen are rather pale in dorsal view.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8CAAFAA5EBDA" blockId="17.[151,1437,151,1977]" box="[199,1336,1159,1184]" pageId="17" pageNumber="218">Measurements of Mount Kinabalu female (mm). Hw 33.5, abdomen excluding anal appendages 35.5.</paragraph>
</subSubSection>
<subSubSection id="C3704B919843FF9BFF3A8C8FFE73E8C3" pageId="17" pageNumber="218" type="discussion">
<paragraph id="8BD5181A9843FF9BFF3A8C8FFD21E957" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
<emphasis id="B91EC4089843FF9BFF3A8C8FFEA6EBBF" bold="true" box="[199,315,1196,1221]" pageId="17" pageNumber="218">Remarks.</emphasis>
Both males examined have very pale cerci, although both are teneral this may reflect the mature condition since the anal appendages of teneral individuals of other Bornean
<taxonomicName id="4C6A63999843FF9BFBEF8CECFB2AEB92" authorityName="Selys in Borneo" authorityYear="1878" box="[1042,1207,1231,1256]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91EC4089843FF9BFBEF8CECFB2AEB92" box="[1042,1207,1231,1256]" italics="true" pageId="17" pageNumber="218">Leptogomphus</emphasis>
</taxonomicName>
males are typically somewhat darker. The ventral spur on the male cercus is unusual in Bornean
<taxonomicName id="4C6A63999843FF9BFC098CD7FB04EA77" authorityName="Selys in Borneo" authorityYear="1878" box="[1012,1177,1268,1293]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91EC4089843FF9BFC098CD7FB04EA77" box="[1012,1177,1268,1293]" italics="true" pageId="17" pageNumber="218">Leptogomphus</emphasis>
</taxonomicName>
, but a similar structure is evident in, for instance,
<taxonomicName id="4C6A63999843FF9BFE388D34FCD0EA4A" authority="Selys, 1891" authorityName="Selys" authorityYear="1891" box="[453,845,1303,1328]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B91EC4089843FF9BFE388D34FD23EA4A" box="[453,702,1303,1328]" italics="true" pageId="17" pageNumber="218">Leptogomphus gestroi</emphasis>
Selys, 1891
</taxonomicName>
, known from
<collectingCountry id="F37D588A9843FF9BFC108D3AFBA1EA4A" box="[1005,1084,1305,1328]" name="Myanmar" pageId="17" pageNumber="218">Burma</collectingCountry>
,
<collectingCountry id="F37D588A9843FF9BFBB58D34FB33EA4A" box="[1096,1198,1303,1328]" name="Thailand" pageId="17" pageNumber="218">Thailand</collectingCountry>
,
<collectingCountry id="F37D588A9843FF9BFB468D34FB63EA4A" box="[1211,1278,1303,1328]" name="China" pageId="17" pageNumber="218">China</collectingCountry>
and
<collectingCountry id="F37D588A9843FF9BFACB8D34FA04EA4A" box="[1334,1433,1303,1328]" name="Vietnam" pageId="17" pageNumber="218">Vietnam</collectingCountry>
. More significantly,
<taxonomicName id="4C6A63999843FF9BFE898D1EFD9AEA2F" authorityName="Selys" authorityYear="1878" box="[372,519,1340,1365]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B91EC4089843FF9BFE898D1EFD9AEA2F" box="[372,519,1340,1365]" italics="true" pageId="17" pageNumber="218">L. lansbergei</emphasis>
</taxonomicName>
from
<collectingRegion id="49AED6F89843FF9BFDB18D1EFDE3EA2E" box="[588,638,1341,1364]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="218">Java</collectingRegion>
and
<collectingRegion id="49AED6F89843FF9BFD488D1FFC8FEA2E" box="[693,786,1340,1364]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="218">Sumatra</collectingRegion>
also has this structure, and in fact the anal appendages of
<taxonomicName id="4C6A63999843FF9BFF6A8D42FED5EA02" authorityName="lansbergei Selys" authorityYear="1878" box="[151,328,1375,1400]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="lansbergei" subSpecies="lansbergei">
<emphasis id="B91EC4089843FF9BFF6A8D42FED5EA02" box="[151,328,1375,1400]" italics="true" pageId="17" pageNumber="218">L. l. lansbergei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A63999843FF9BFE7A8D42FE78EA02" authorityName="van Tol" authorityYear="1990" box="[391,485,1377,1400]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089843FF9BFE7A8D42FE78EA02" box="[391,485,1377,1400]" italics="true" pageId="17" pageNumber="218">L. pasia</emphasis>
</taxonomicName>
are extremely similar (see
<figureCitation id="1351049F9843FF9BFCD78D7CFCE3EA02" box="[810,894,1375,1400]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="17" pageNumber="218">
Fig.
<quantity id="4C92B5FF9843FF9BFC9F8D7CFCE3EA02" box="[866,894,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="17" pageNumber="218" unit="in" value="15.0">15</quantity>
</figureCitation>
in
<bibRefCitation id="EFFB65EB9843FF9BFC578D7CFB8BEA02" author="Ris" box="[938,1046,1375,1400]" pageId="17" pageNumber="218" refString="Ris, F. (1927) Odonaten von Sumatra, gesammelt von Edward Jacobson. Zoologische Mededelingen, 10, 1 - 49." type="journal article" year="1927">Ris 1927</bibRefCitation>
), although the two differ in their markings. Note that the similarity is less marked when compared with van Tol’s illustrations of
<taxonomicName id="4C6A63999843FF9BFB138DA6FA01EAE7" authorityName="lansbergei Selys" authorityYear="1878" box="[1262,1436,1412,1437]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="lansbergei" subSpecies="lansbergei">
<emphasis id="B91EC4089843FF9BFB138DA6FA01EAE7" box="[1262,1436,1412,1437]" italics="true" pageId="17" pageNumber="218">L. l. lansbergei</emphasis>
</taxonomicName>
(
<figureCitation id="1351049F9843FF9BFF628D84FE80EABA" box="[159,285,1447,1472]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="17" pageNumber="218">Figs 30, 31</figureCitation>
on page 103). It is also interesting to note that the cerci of
<taxonomicName id="4C6A63999843FF9BFC518D8AFBC9EABA" authorityName="lansbergei Selys" authorityYear="1878" box="[940,1108,1447,1472]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="lansbergei" subSpecies="lansbergei">
<emphasis id="B91EC4089843FF9BFC518D8AFBC9EABA" box="[940,1108,1447,1472]" italics="true" pageId="17" pageNumber="218">L. l. lansbergei</emphasis>
</taxonomicName>
are whitish in colour, as they appear to be in
<taxonomicName id="4C6A63999843FF9BFEAF8DEEFE32EA9E" authorityName="van Tol" authorityYear="1990" box="[338,431,1485,1508]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089843FF9BFEAF8DEEFE32EA9E" box="[338,431,1485,1508]" italics="true" pageId="17" pageNumber="218">L. pasia</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFB65EB9843FF9BFE3C8DEFFDA3EA9E" author="Ris" box="[449,574,1484,1509]" pageId="17" pageNumber="218" refString="Ris, F. (1927) Odonaten von Sumatra, gesammelt von Edward Jacobson. Zoologische Mededelingen, 10, 1 - 49." type="journal article" year="1927">Ris (1927)</bibRefCitation>
, as noted by van
<bibRefCitation id="EFFB65EB9843FF9BFCEF8DEFFC12EA9E" author="Tol" box="[786,911,1484,1509]" pageId="17" pageNumber="218" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol (1990)</bibRefCitation>
, considered that the anal appendages of
<taxonomicName id="4C6A63999843FF9BFA918DEEFE91E972" authorityName="lansbergei Selys" authorityYear="1878" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="subSpecies" species="lansbergei" subSpecies="lansbergei">
<emphasis id="B91EC4089843FF9BFA918DEEFE91E972" italics="true" pageId="17" pageNumber="218">L. l. lansbergei</emphasis>
</taxonomicName>
and
<emphasis id="B91EC4089843FF9BFEB58DD2FE7FE972" box="[328,482,1519,1544]" italics="true" pageId="17" pageNumber="218">L. l. assimilis</emphasis>
are different, in which case in our view they should be treated as distinct species. However this is a matter in need of further study.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8E14FD98E9E2" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
<bibRefCitation id="EFFB65EB9843FF9BFF3A8E14FEE9E92A" author="Laidlaw" box="[199,372,1591,1616]" pageId="17" pageNumber="218" refString="Laidlaw, F. F. (1931) Revised list of the dragonflies of Borneo. Journal of the Federated Malay States Museums, 16 (3 / 4), 234 - 250." type="journal article" year="1931">Laidlaw (1931)</bibRefCitation>
recorded a male of
<emphasis id="B91EC4089843FF9BFDAA8E1AFD73E92A" box="[599,750,1591,1616]" italics="true" pageId="17" pageNumber="218">L. l. assimilis</emphasis>
from Bettotan in
<collectingRegion id="49AED6F89843FF9BFC4A8E14FC62E92A" box="[951,1023,1591,1616]" country="Malaysia" name="Sabah" pageId="17" pageNumber="218">Sabah</collectingRegion>
; as already noted by van
<bibRefCitation id="EFFB65EB9843FF9BFAE38E14FA0AE92A" author="Tol" box="[1310,1431,1591,1616]" pageId="17" pageNumber="218" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol (1990)</bibRefCitation>
, this specimen cannot be located now. It is very unlikely that the Bettotan specimen was actually
<emphasis id="B91EC4089843FF9BFB2D8E7EFAF8E90F" box="[1232,1381,1628,1653]" italics="true" pageId="17" pageNumber="218">L. l. assimilis</emphasis>
, and possibly it was actually
<taxonomicName id="4C6A63999843FF9BFE598EA2FE63E9E2" authorityName="van Tol" authorityYear="1990" box="[420,510,1665,1688]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089843FF9BFE598EA2FE63E9E2" box="[420,510,1665,1688]" italics="true" pageId="17" pageNumber="218">L. pasia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8E87FD87E837" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
The male of
<taxonomicName id="4C6A63999843FF9BFEAA8E86FE2CE9C6" authorityName="van Tol" authorityYear="1990" box="[343,433,1701,1724]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089843FF9BFEAA8E86FE2CE9C6" box="[343,433,1701,1724]" italics="true" pageId="17" pageNumber="218">L. pasia</emphasis>
</taxonomicName>
was associated to the female based on similarity of markings and the match is supported by the molecular results; the two samples, from separate populations, are identical in COI and have 99.6% identical sites in ITS. In all of our analyses (
<figureCitation id="1351049F9843FF9BFDDD8ECFFDFAE87E" box="[544,615,1772,1797]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="17" pageNumber="218">Figs 1</figureCitation>
,
<figureCitation id="1351049F9843FF9BFD888ECFFD18E87E" box="[629,645,1772,1796]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="17" pageNumber="218">2</figureCitation>
,
<figureCitation id="1351049F9843FF9BFD6C8ECFFD02E87E" box="[657,671,1772,1796]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="17" pageNumber="218">3</figureCitation>
) this species is the sister of
<taxonomicName id="4C6A63999843FF9BFC218ECEFB89E87E" authorityName="Dow &amp; Stokvis &amp; Ngiam" authorityYear="2017" box="[988,1044,1773,1796]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="sii">
<emphasis id="B91EC4089843FF9BFC218ECEFB89E87E" box="[988,1044,1773,1796]" italics="true" pageId="17" pageNumber="218">L. sii</emphasis>
</taxonomicName>
, however given the morphological similarity with
<taxonomicName id="4C6A63999843FF9BFEB78F32FE7CE852" authorityName="Selys" authorityYear="1878" box="[330,481,1807,1832]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B91EC4089843FF9BFEB78F32FE7CE852" box="[330,481,1807,1832]" italics="true" pageId="17" pageNumber="218">L. lansbergei</emphasis>
</taxonomicName>
, an expanded analysis including
<taxonomicName id="4C6A63999843FF9BFC948F32FB9DE852" authorityName="Selys" authorityYear="1878" box="[873,1024,1807,1832]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B91EC4089843FF9BFC948F32FB9DE852" box="[873,1024,1807,1832]" italics="true" pageId="17" pageNumber="218">L. lansbergei</emphasis>
</taxonomicName>
is highly desirable to establish the relationships among these species.
</paragraph>
<paragraph id="8BD5181A9843FF9BFF3A8F74FE73E8C3" blockId="17.[151,1437,151,1977]" pageId="17" pageNumber="218">
Confirmed specimens of
<taxonomicName id="4C6A63999843FF9BFE1B8F7AFDDEE80A" authorityName="van Tol" authorityYear="1990" box="[486,579,1881,1904]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC4089843FF9BFE1B8F7AFDDEE80A" box="[486,579,1881,1904]" italics="true" pageId="17" pageNumber="218">L. pasia</emphasis>
</taxonomicName>
are from the Mount Kinabalu, the Long Pasia area of
<collectingRegion id="49AED6F89843FF9BFB488F74FB64E80A" box="[1205,1273,1879,1904]" country="Malaysia" name="Sabah" pageId="17" pageNumber="218">Sabah</collectingRegion>
and the Tama Abu range in
<collectingRegion id="49AED6F89843FF9BFEC48F5FFE04E8EF" box="[313,409,1916,1941]" country="Malaysia" name="Sarawak" pageId="17" pageNumber="218">
<collectingCountry id="F37D588A9843FF9BFEC48F5FFE04E8EF" box="[313,409,1916,1941]" name="Malaysia" pageId="17" pageNumber="218">Sarawak</collectingCountry>
</collectingRegion>
(almost contiguous with the Long Pasia area of
<collectingRegion id="49AED6F89843FF9BFC2E8F5FFB84E8EF" box="[979,1049,1916,1941]" country="Malaysia" name="Sabah" pageId="17" pageNumber="218">Sabah</collectingRegion>
) (
<figureCitation id="1351049F9843FF9BFBCE8F5FFB04E8EE" box="[1075,1177,1916,1941]" captionStart="FIGURE 127" captionStartId="49.[151,250,1596,1618]" captionTargetBox="[151,1436,193,1574]" captionTargetId="figure@49.[151,1436,193,1574]" captionTargetPageId="49" captionText="FIGURE 127. Distribution of L. pasia (white circle) and L. sii (blacksquare)." httpUri="https://zenodo.org/record/1068056/files/figure.png" pageId="17" pageNumber="218">Fig. 127</figureCitation>
). All records are from upland and montane locations.
</paragraph>
</subSubSection>
</treatment>
</document>
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