394 lines
61 KiB
XML
394 lines
61 KiB
XML
<document id="3790EF142AEDD9FA67F8198A6AC01DEB" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank & Ngiam, Robin W. J." docDate="2017" docId="03C3A90C9844FF90FF6A8DD1FDD6ED1A" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus schieli Dow & Stokvis & Ngiam 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="227" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="223" updateTime="1698505057438" updateUser="ExternalLinkService">
|
||
<mods:mods id="F76A2347976DF1371BA1FE92E32D62EA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A79D0D041C22448D2FBCECE9C3E0B620">
|
||
<mods:title id="9AB31822103846855A0DAFC0864E2E8B">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="8CAD56E5FDD03D5E28B2909104EC1462" type="personal">
|
||
<mods:role id="B316E94E23BEB27331F3ED68409E7361">
|
||
<mods:roleTerm id="CD82C89A1B333CE8D30A936743EBC603">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8F610EA03D34ACFC3B628D59457B6444">Dow, Rory A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2F4162F5293819BFE9238C4004C060A7" type="personal">
|
||
<mods:role id="8300D1DEFB48171CCB080E391CE646AA">
|
||
<mods:roleTerm id="D37FF39555706660F2D9EE1DB363BDE7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7DDCF4685D79C7C322FD14B478C26D83">Stokvis, Frank</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8B3EC09E35132F8D89099A806B7A9BB1" type="personal">
|
||
<mods:role id="A3B85147400A85D9A63DC7452DFBED61">
|
||
<mods:roleTerm id="23DDD956C11C442ABE4702F60A3B5A6F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A7F55C9EF385B83523D7F9AAEE6DE2DE">Ngiam, Robin W. J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="12FD0CACCC746FA061C4EAB1C7AAD1F4">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F2D7A17EFCD454A58ED558FAA0B2644B" type="host">
|
||
<mods:titleInfo id="265EC69DA13A4FC211731958D4247627">
|
||
<mods:title id="D5389F1F4AE7D6F6B7D3096272B2779F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1CA75581B211AF3450588FEFB7464476">
|
||
<mods:date id="4FAAA67D6EFC48FDDD48667C0418A5AA">2017</mods:date>
|
||
<mods:detail id="4C2C58358C908D5C160C10E76D243310" type="pubDate">
|
||
<mods:number id="321A995BA46EA4A9278DD759458E0392">2017-11-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BC9F563579BC3CD28C7D0C537ECF1ADC" type="volume">
|
||
<mods:number id="98D9ABA4B51D1915658AE723E9D0CB8C">4358</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="58BFC28D799B4157F0D5414293385539" type="issue">
|
||
<mods:number id="0C23883B076BFB9F9F241365859DDDDE">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1CF0EAB886DBD64BCCA5A53635628681" unit="page">
|
||
<mods:start id="C7157A1E62CC6E07525500FEBB69C322">201</mods:start>
|
||
<mods:end id="67CC925D4F8A46AEFC3462C8CCE74E4D">257</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="EB187B0237E9028A6FFF08B00FF808D0">journal article</mods:classification>
|
||
<mods:identifier id="D1AF3CF5CFE67FA1370C006E05D5D682" type="CLB-Dataset">31287</mods:identifier>
|
||
<mods:identifier id="A4BF48407956E7086D724C5744BA110C" type="DOI">10.11646/zootaxa.4358.2.1</mods:identifier>
|
||
<mods:identifier id="3028664D7C3C866A0FB9005D24DC7A22" type="GBIF-Dataset">d226f633-d454-4e32-b12f-6d00c30bddfc</mods:identifier>
|
||
<mods:identifier id="8721DF3151E3602EFF9FC5C3550AAEB2" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="AA6D2AF4E6DFBAFF9FBA8A9904D1FCD8" type="Zenodo-Dep">1067996</mods:identifier>
|
||
<mods:identifier id="A29501EC45779D9A55080033733A26B0" type="ZooBank">8861BCC0-022F-4803-98E8-D28B90F666E4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C3A90C9844FF90FF6A8DD1FDD6ED1A" ID-DOI="http://doi.org/10.5281/zenodo.5631254" ID-GBIF-Taxon="137580778" ID-Zenodo-Dep="5631254" LSID="urn:lsid:plazi:treatment:03C3A90C9844FF90FF6A8DD1FDD6ED1A" httpUri="http://treatment.plazi.org/id/03C3A90C9844FF90FF6A8DD1FDD6ED1A" lastPageId="26" lastPageNumber="227" pageId="22" pageNumber="223">
|
||
<subSubSection id="C3704B919844FF9CFF6A8DD1FC4EE957" pageId="22" pageNumber="223" type="nomenclature">
|
||
<paragraph id="8BD5181A9844FF9CFF6A8DD1FD99E976" blockId="22.[151,979,1522,1581]" box="[151,516,1522,1548]" pageId="22" pageNumber="223">
|
||
<heading id="D09DAF769844FF9CFF6A8DD1FD99E976" bold="true" box="[151,516,1522,1548]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
|
||
<emphasis id="B91EC4089844FF9CFF6A8DD1FD99E976" bold="true" box="[151,516,1522,1548]" pageId="22" pageNumber="223">
|
||
<taxonomicName id="4C6A63999844FF9CFF6A8DD1FE02E976" ID-CoL="9X355" authority="Dow & Stokvis & Ngiam, 2017" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,415,1522,1548]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="schieli" status="sp. nov.">
|
||
<emphasis id="B91EC4089844FF9CFF6A8DD1FE02E976" bold="true" box="[151,415,1522,1548]" italics="true" pageId="22" pageNumber="223">Leptogomphus schieli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D79739844FF9CFE5A8DD1FD99E976" box="[423,516,1522,1548]" pageId="22" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9844FF9CFF6A8E37FC4EE957" blockId="22.[151,979,1522,1581]" box="[151,979,1556,1581]" pageId="22" pageNumber="223">
|
||
(
|
||
<figureCitation id="1351049F9844FF9CFF628E37FF7AE957" box="[159,231,1556,1581]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="22" pageNumber="223">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFF0E8E37FE8CE957" box="[243,273,1556,1581]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="22" pageNumber="223">16</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFEE08E37FEA6E957" box="[285,315,1556,1581]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="22" pageNumber="223">25</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFEBA8E36FEF8E957" box="[327,357,1557,1581]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="22" pageNumber="223">33</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFE8C8E36FE12E957" box="[369,399,1557,1581]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="22" pageNumber="223">43</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFE668E36FE27E957" box="[411,442,1557,1581]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="22" pageNumber="223">48</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFE3B8E37FE79E957" box="[454,484,1556,1581]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="22" pageNumber="223">58</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFE128E37FD90E957" box="[495,525,1556,1581]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="22" pageNumber="223">62</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFDE48E37FDAAE957" box="[537,567,1556,1581]" captionStart="FIGURES 74–77" captionStartId="37.[151,264,1356,1379]" captionTargetBox="[151,1435,194,1335]" captionTargetId="figure@37.[151,1436,193,1335]" captionTargetPageId="37" captionText="FIGURES 74–77. Female synthorax dorsal: (74) L pendleburyi SAR16_GOM8; (75) L. schieli paratype SAR15_GOM6; (76) L. williamsoni SAR13_14_GOM51; (77) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068034/files/figure.png" pageId="22" pageNumber="223">75</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFDBE8E37FDFCE957" box="[579,609,1556,1581]" captionStart="FIGURES 84–87" captionStartId="39.[151,264,1427,1450]" captionTargetBox="[151,1435,194,1406]" captionTargetId="figure@39.[151,1436,193,1406]" captionTargetPageId="39" captionText="FIGURES 84–87. Female synthorax lateral: (84) L pendleburyi SAR16_GOM8; (85) L. schieli paratype SAR15_GOM6; (86) L. williamsoni SAR13_14_GOM51, right hand side, flipped for consistency;(87) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068038/files/figure.png" pageId="22" pageNumber="223">85</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFD938E36FD11E957" box="[622,652,1557,1581]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="22" pageNumber="223">91</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFD6A8E36FD28E957" box="[663,693,1557,1581]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="22" pageNumber="223">97</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFD3C8E36FD70E957" box="[705,749,1557,1581]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="22" pageNumber="223">103</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFD048E36FCB8E957" box="[761,805,1557,1581]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="22" pageNumber="223">109</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFCCC8E37FCC1E957" box="[817,860,1556,1581]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="22" pageNumber="223">115</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFC958E36FC09E957" box="[872,916,1557,1581]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="22" pageNumber="223">123</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9844FF9CFC5D8E36FC56E957" box="[928,971,1557,1581]" captionStart="FIGURE128" captionStartId="50.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1589]" captionTargetId="figure@50.[151,1436,193,1589]" captionTargetPageId="50" captionText="FIGURE128. Distributionof L. pendleburyi (yellowcircle), L. schieli (black square) and L. species from Pulong Tau National Park, Sarawak (white square)." httpUri="https://zenodo.org/record/1068058/files/figure.png" pageId="22" pageNumber="223">128</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919844FF9CFF6A8E7FFB99E908" box="[151,1028,1628,1651]" pageId="22" pageNumber="223" type="description">
|
||
<paragraph id="8BD5181A9844FF9CFF6A8E7FFB99E908" blockId="22.[151,1028,1628,1651]" box="[151,1028,1628,1651]" pageId="22" pageNumber="223">
|
||
<taxonomicName id="4C6A63999844FF9CFF6A8E7FFEB1E908" authority="Dow & Stokvis & Ngiam, 2017" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,300,1628,1650]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
<emphasis id="B91EC4089844FF9CFF6A8E7FFEB1E908" box="[151,300,1628,1650]" italics="true" pageId="22" pageNumber="223">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D79739844FF9CFECF8E7EFE30E908" box="[306,429,1628,1650]" pageId="22" pageNumber="223" rank="species">new species</taxonomicNameLabel>
|
||
;—Dow
|
||
<emphasis id="B91EC4089844FF9CFDF98E7DFDAAE908" box="[516,567,1628,1651]" italics="true" pageId="22" pageNumber="223">et al.</emphasis>
|
||
2016: 10, 13 (both sexes Gunung Penrissen).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919844FF9CFF6A8EBCFBBEE83D" pageId="22" pageNumber="223" type="materials_examined">
|
||
<paragraph id="8BD5181A9844FF9CFF6A8EBCFE1FE985" blockId="22.[151,1437,1695,2008]" pageId="22" pageNumber="223">
|
||
<typeStatus id="54D1A6B89844FF9CFF6A8EBCFE99E9C2" box="[151,260,1695,1720]" pageId="22" pageNumber="223" type="holotype">
|
||
<emphasis id="B91EC4089844FF9CFF6A8EBCFE99E9C2" bold="true" box="[151,260,1695,1720]" pageId="22" pageNumber="223">Holotype</emphasis>
|
||
</typeStatus>
|
||
<specimenCount id="9D6CD3939844FF9CFEF68EBCFEABE9CD" box="[267,310,1695,1719]" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF9844FF9CFEB88EBCFEE2E9C2" box="[325,383,1695,1720]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="22" pageNumber="223">SAR</collectionCode>
|
||
15_GOM8), by very small, steep stream below peak of Gunung Penrissen, Borneo Highlands resort area on Gunung Penrissen, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9844FF9CFD068EE0FCF1E9A6" box="[763,876,1731,1756]" direction="north" orientation="latitude" pageId="22" pageNumber="223" precision="1" value="1.11905">1.11905N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9844FF9CFC878EE7FB9EE9A6" box="[890,1027,1732,1756]" direction="east" orientation="longitude" pageId="22" pageNumber="223" precision="1" value="110.22793">110.22793E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9844FF9CFBEF8EE7FB07E9A6" box="[1042,1178,1731,1756]" pageId="22" pageNumber="223" value="2015-07-14">14 vii 2015</date>
|
||
, leg. G.T. Reels, to be deposited in
|
||
<collectionCode id="ED7B80DF9844FF9CFEDB8ECBFEE3E985" box="[294,382,1768,1791]" pageId="22" pageNumber="223">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9844FF9CFF3A8F28FBBEE83D" blockId="22.[151,1437,1695,2008]" pageId="22" pageNumber="223">
|
||
<emphasis id="B91EC4089844FF9CFF3A8F28FEDBE85E" bold="true" box="[199,326,1803,1828]" pageId="22" pageNumber="223">
|
||
<typeStatus id="54D1A6B89844FF9CFF3A8F28FEDCE85E" box="[199,321,1803,1828]" pageId="22" pageNumber="223" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939844FF9CFEB28F2FFE0DE859" box="[335,400,1804,1828]" pageId="22" pageNumber="223" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF9844FF9CFE5C8F28FE47E85E" box="[417,474,1803,1828]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="22" pageNumber="223">SAR</collectionCode>
|
||
15_GOM6, 7; supposition), seepage below peak of Gunung Penrissen, same area as
|
||
<typeStatus id="54D1A6B89844FF9CFF6A8F0CFF61E832" box="[151,252,1839,1864]" pageId="22" pageNumber="223" type="holotype">holotype</typeStatus>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9844FF9CFEFA8F0CFEE5E832" box="[263,376,1839,1864]" direction="north" orientation="latitude" pageId="22" pageNumber="223" precision="1" value="1.11905">1.11905N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9844FF9CFE7E8F0CFD90E83D" box="[387,525,1839,1863]" direction="east" orientation="longitude" pageId="22" pageNumber="223" precision="1" value="110.22793">110.22793E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9844FF9CFDE58F0CFD07E832" box="[536,666,1839,1864]" pageId="22" pageNumber="223" value="2015-07-15">15 vii 2015</date>
|
||
, leg. R.A. Dow, in collection Dow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919844FF9CFF3A8F70FEB2E8EA" pageId="22" pageNumber="223" type="etymology">
|
||
<paragraph id="8BD5181A9844FF9CFF3A8F70FEB2E8EA" blockId="22.[151,1437,1695,2008]" pageId="22" pageNumber="223">
|
||
<emphasis id="B91EC4089844FF9CFF3A8F70FED3E816" bold="true" box="[199,334,1875,1900]" pageId="22" pageNumber="223">Etymology.</emphasis>
|
||
Schieli, a noun in the genitive case. Named for Franz-Josef Schiel in honour of his contributions to Odonatology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919844FF9DFF3A8FB8FD31EFAE" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="diagnosis">
|
||
<paragraph id="8BD5181A9844FF9DFF3A8FB8FD31EFAE" blockId="22.[151,1437,1695,2008]" lastBlockId="23.[151,1436,151,213]" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
|
||
<emphasis id="B91EC4089844FF9CFF3A8FB8FEDCE8CE" bold="true" box="[199,321,1947,1972]" pageId="22" pageNumber="223">Diagnosis.</emphasis>
|
||
The male is easily distinguished from those of all other
|
||
<taxonomicName id="4C6A63999844FF9CFC438FB8FBFEE8CE" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[958,1123,1947,1972]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EC4089844FF9CFC438FB8FBFEE8CE" box="[958,1123,1947,1972]" italics="true" pageId="22" pageNumber="223">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
known from Borneo by the convergent branches of the epiproct. The female is distinguished from those of all species known from Borneo except
|
||
<taxonomicName id="4C6A63999845FF9DFF1588BAFE17EFCA" authorityName="Laidlaw" authorityYear="1912" box="[232,394,151,176]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089845FF9DFF1588BAFE17EFCA" box="[232,394,151,176]" italics="true" pageId="23" pageNumber="224">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<emphasis id="B91EC4089845FF9DFE3088BAFE79EFCA" box="[461,484,153,176]" italics="true" pageId="23" pageNumber="224">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A63999845FF9DFDCE88B4FD2BEFCA" authorityName="Laidlaw" authorityYear="1912" box="[563,694,151,176]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089845FF9DFDCE88B4FD2BEFCA" box="[563,694,151,176]" italics="true" pageId="23" pageNumber="224">williamsoni</emphasis>
|
||
</taxonomicName>
|
||
) by the dorsal markings of the thorax, and from the female of
|
||
<taxonomicName id="4C6A63999845FF9DFA7888BAFE87EFAF" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089845FF9DFA7888BAFE87EFAF" italics="true" pageId="23" pageNumber="224">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
by the head lacking occipital spurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1548929845FF9DFF6A8D6EFEF3EADB" httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="23" pageNumber="224" targetBox="[151,1436,265,1336]" targetPageId="23">
|
||
<paragraph id="8BD5181A9845FF9DFF6A8D6EFEF3EADB" blockId="23.[151,1436,1357,1441]" pageId="23" pageNumber="224">
|
||
<emphasis id="B91EC4089845FF9DFF6A8D6EFECAEA19" bold="true" box="[151,343,1357,1380]" pageId="23" pageNumber="224">FIGURES 32–35.</emphasis>
|
||
Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32)
|
||
<taxonomicName id="4C6A63999845FF9DFAF68D6CFA01EA1E" authorityName="Steinhoff" authorityYear="2015" box="[1291,1436,1358,1380]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC4089845FF9DFAF68D6CFA01EA1E" box="[1291,1436,1358,1380]" italics="true" pageId="23" pageNumber="224">L pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
SAR16_GOM8; (33)
|
||
<taxonomicName id="4C6A63999845FF9DFE8A8D4DFE48EAF8" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[375,469,1388,1411]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089845FF9DFE8A8D4DFE48EAF8" box="[375,469,1388,1411]" italics="true" pageId="23" pageNumber="224">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
paratype SAR15_GOM6; (34)
|
||
<taxonomicName id="4C6A63999845FF9DFCE18D4DFC33EAF8" authorityName="Laidlaw" authorityYear="1912" box="[796,942,1388,1411]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089845FF9DFCE18D4DFC33EAF8" box="[796,942,1388,1411]" italics="true" pageId="23" pageNumber="224">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR13_14_GOM51; (35)
|
||
<emphasis id="B91EC4089845FF9DFB348D4DFB43EAF9" box="[1225,1246,1390,1411]" italics="true" pageId="23" pageNumber="224">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A63999845FF9DFADB8D4FFA01EAF8" authorityName="Laidlaw" authorityYear="1912" box="[1318,1436,1388,1410]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089845FF9DFADB8D4FFA01EAF8" box="[1318,1436,1388,1410]" italics="true" pageId="23" pageNumber="224">williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM59.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3704B919845FF90FF3A8DEEFC40EFCA" lastPageId="26" lastPageNumber="227" pageId="23" pageNumber="224" type="description">
|
||
<paragraph id="8BD5181A9845FF9DFF3A8DEEFA16E9E1" blockId="23.[151,1437,1485,2015]" pageId="23" pageNumber="224">
|
||
<emphasis id="B91EC4089845FF9DFF3A8DEEFDB3EA9C" bold="true" box="[199,558,1485,1510]" pageId="23" pageNumber="224">
|
||
Description of
|
||
<typeStatus id="54D1A6B89845FF9DFE808DEEFE79EA9C" box="[381,484,1485,1510]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
<emphasis id="B91EC4089845FF9DFDC48DEEFDEBEA9C" box="[569,630,1485,1510]" italics="true" pageId="23" pageNumber="224">Head</emphasis>
|
||
(
|
||
<figureCitation id="1351049F9845FF9DFD7A8DEEFD4EEA9C" box="[647,723,1485,1510]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="23" pageNumber="224">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9845FF9DFD1E8DEEFC9FEA9C" box="[739,770,1485,1510]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="23" pageNumber="224">16</figureCitation>
|
||
). Labium largely pale, very dark brown basally centrally, apical part median lobe, narrowly on apical outer margin of palps, and hooks. Labrum black, transverse basal pale stripe. Mandible bases pale, genae pale adjacent to mandible bases, otherwise black. Clypeus black with pair of widely separated yellow marks at anterior lateral corners postclypeus. Ante- and postfrons not very sharply divided, postfrons almost entirely pale bluish (
|
||
<figureCitation id="1351049F9845FF9DFD538E7EFC9EE90C" box="[686,771,1629,1654]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="23" pageNumber="224">Fig. 16</figureCitation>
|
||
). Pair of prominent, robust tubercles behind lateral ocelli, occupying almost all of vertex between ocelli and occipital plate; ocelli pale yellowish. Vertex and occiput black.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9845FF92FF3A8E86FB3AEE3A" blockId="23.[151,1437,1485,2015]" lastBlockId="24.[151,1437,151,1964]" lastPageId="24" lastPageNumber="225" pageId="23" pageNumber="224">
|
||
<emphasis id="B91EC4089845FF9DFF3A8E86FE85E9C4" box="[199,280,1701,1726]" italics="true" pageId="23" pageNumber="224">Thorax</emphasis>
|
||
. Prothorax mostly black, pale irregular transverse stripe anteriorly on anterior pronotal lobe, greenish yellow on lateral parts, brownish orange centrally. Pair of greenish-yellow lateral marks, small central posterior mark (divided centrally) on middle pronotal lobe. Propleuron mottled dark brown and pale. Synthorax black and dark brown with pale greenish markings as follows (
|
||
<figureCitation id="1351049F9845FF9DFD148F31FCDCE851" box="[745,833,1810,1835]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="23" pageNumber="224">Figs 58</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9845FF9DFCB08F31FCF0E851" box="[845,877,1810,1835]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="23" pageNumber="224">62</figureCitation>
|
||
): broad mesothoracic collar, divided at middorsal carina. Narrow dorsal thoracic stripes reaching level of apex of mesothoracic collar but separated from it, extending well past apex of antealar crest. Narrow antehumeral stripes just separated from mesokatepisternum. Mesepimeron with broad stripe, irregular along its lower margin. Metepisternum with large subrectangular mark near antealar carina, metepimeron with broad stripe. Mesokatepisternum yellow in line with antehumeral stripe, black above. Metakatepisternum largely yellowish. Metaposternum greyish and pale brown. Legs robust and relatively short. Coxae mottled pale, grey and brown, trochanters largely black. Rest of legs black except inner surface of anterior femora with pale stripe. Wings: sectors of arculus separated at origin with 5 cross veins up to and at first bifurcation of superior sector in Fw,
|
||
<quantity id="4C92B5FF984AFF92FD1C88C3FC8FEF82" box="[737,786,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="24" pageNumber="225" unit="in" value="3.0">3 in</quantity>
|
||
Hw. Discoidal field with 2 rows of cells from origin, transitioning to three rows before level of nodus in both wings. 14 (right) to 15 (left) Ax in Fw,
|
||
<quantity id="4C92B5FF984AFF92FB368927FA98EE67" box="[1227,1285,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="225" unit="in" value="10.0">10 in</quantity>
|
||
Hw, 10 (left) or 12 (right) Px in Fw, 14 (right) to 12 (left) in Hw. Pt brown, covering ca 4 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A896FFE2BECDF" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A896FFEACEE1F" box="[199,305,332,357]" italics="true" pageId="24" pageNumber="225">Abdomen</emphasis>
|
||
. Slender after base of S3, expanding moderately from base of S7, maximum width reached apical part of S8, then almost constant. Black with pale markings as follows: S1 large pale lateral mark, dorsal mark. S2 central lateral pale mark from base, including part of auricle, narrowly divided from posterior yellow mark, narrow apical middorsal stripe. S3–6 with small lateral markings at base, tiny, barely visible on S6. On left S8 with tiny apical lateral brown mark. S3–7 with very narrow, irregular middorsal line, only present in apical half S3, on S7 slight expanded apically. Cerci (
|
||
<figureCitation id="1351049F984AFF92FDF589DCFDEDED62" box="[520,624,511,536]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="24" pageNumber="225">Figs 103</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FD838A23FD34ED62" box="[638,681,512,536]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="24" pageNumber="225">109</figureCitation>
|
||
) black, broad at base in lateral view (
|
||
<figureCitation id="1351049F984AFF92FB9C89DCFB59ED62" box="[1121,1220,511,536]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="24" pageNumber="225">Fig. 109</figureCitation>
|
||
), tapering to sharp slightly upturned tip, along upper margin sloping steeply down from base to just before mid-length, then gently curved to apex, along lower margin sloping down from base to just before half length, then abruptly up to apex, forming a prominent corner, small teeth present on up-turned section. In dorsal view (
|
||
<figureCitation id="1351049F984AFF92FB6C8A4FFB69EDFE" box="[1169,1268,620,645]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="24" pageNumber="225">Fig. 103</figureCitation>
|
||
) outer margin running very slightly outward from base, then inward to tips at just after half length; prominent corner on lower margin just visible. Dorsally dark brown at apex. Epiproct (
|
||
<figureCitation id="1351049F984AFF92FCC88A97FC06EDB6" box="[821,923,692,717]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="24" pageNumber="225">Figs 103</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FC5A8A97FC4CEDB7" box="[935,977,692,717]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="24" pageNumber="225">115</figureCitation>
|
||
) mostly black, shorter than cerci, deeply divided, branches narrow, turned up and strongly inwards so rounded apices only narrowly separated. In lateral view (
|
||
<figureCitation id="1351049F984AFF92FF208ADFFEA0EC6E" box="[221,317,764,789]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="24" pageNumber="225">Fig. 109</figureCitation>
|
||
) strongly curved, apices before tips of cerci, turned almost to rear; upper margin obscured by cerci. Accessory genitalia as shown in
|
||
<figureCitation id="1351049F984AFF92FDF58B3CFD1FEC42" box="[520,642,799,824]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="24" pageNumber="225">Figures 91</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FD728B03FD30EC42" box="[655,685,800,824]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="24" pageNumber="225">97</figureCitation>
|
||
, with anterior hamule moderate sized, tapering toward sharp apex. Posterior hamule large, ca twice length of anterior hamule, directed postero-ventrally, anterior margin converging on posterior margin for much of length, then whole curving gently forward so directed down, tapering to sharp tip, directed slightly outward.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8B92FDADEB22" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A8B92FE9BECB2" box="[199,262,945,968]" italics="true" pageId="24" pageNumber="225">Penis</emphasis>
|
||
(
|
||
<figureCitation id="1351049F984AFF92FEEB8B8CFEF0ECB2" box="[278,365,943,968]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="24" pageNumber="225">Fig. 97</figureCitation>
|
||
). Penultimate segment extended to rear of join with terminal segment, bifurcated in this part, apices slender, curved down. Terminal segment extended beyond penultimate segment, approximately dumbbell shaped in lateral view, then shallow contracted before strongly expanding dorsally, apically with short rearward directed, curled, cornua; short, rearward directed flap just below cornua. Penis vesicle prominent, slightly more than half height of posterior hamule.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8C46FB1DEB06" blockId="24.[151,1437,151,1964]" box="[199,1152,1124,1149]" pageId="24" pageNumber="225">Measurements (mm). Hw 28, abdomen excluding anal appendages 32.5, cerci ca 1.4.</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8CA4FA9CEAE3" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A8CA4FD38EBDA" bold="true" box="[199,677,1159,1184]" pageId="24" pageNumber="225">Description of female (SAR15_GOM6).</emphasis>
|
||
As male except as noted.
|
||
<emphasis id="B91EC408984AFF92FC268CA4FB85EBDA" box="[987,1048,1159,1184]" italics="true" pageId="24" pageNumber="225">Head</emphasis>
|
||
(
|
||
<figureCitation id="1351049F984AFF92FBD78CA4FB19EBDA" box="[1066,1156,1159,1184]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="24" pageNumber="225">Figs 25</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FB6F8CABFB2DEBDA" box="[1170,1200,1160,1184]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="24" pageNumber="225">33</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FB428CABFB40EBDA" box="[1215,1245,1160,1184]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="24" pageNumber="225">43</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FB118CABFA91EBDA" box="[1260,1292,1160,1184]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="24" pageNumber="225">48</figureCitation>
|
||
). Transverse basal pale stripe of labium narrowly divided into large lateral parts and small central part. Yellow anterior lateral marks on post clypeus larger, small indistinct upper central mark. Stripe on postfrons greenish yellow. Pair of small tubercles immediately behind and inside of lateral ocelli, narrowly separated from slightly larger tubercles adjacent to occipital plate, these separated from occipital plate by narrow but deep depression. Rear tubercles green on their inner faces, area between them and onto rear of anterior pair of tubercles pale greyish green. Occipital plate black bearing pair of widely separated horns, directed up and slightly forward, pair of small inconspicuous tubercles between these at occipital ridge. No occipital spurs, but pair bulges on occiput below compound eye.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8D80FA05E95C" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A8D80FE85EAC6" box="[199,280,1443,1468]" italics="true" pageId="24" pageNumber="225">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="1351049F984AFF92FEDA8D80FEE3EAC6" box="[295,382,1443,1468]" captionStart="FIGURES 74–77" captionStartId="37.[151,264,1356,1379]" captionTargetBox="[151,1435,194,1335]" captionTargetId="figure@37.[151,1436,193,1335]" captionTargetPageId="37" captionText="FIGURES 74–77. Female synthorax dorsal: (74) L pendleburyi SAR16_GOM8; (75) L. schieli paratype SAR15_GOM6; (76) L. williamsoni SAR13_14_GOM51; (77) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068034/files/figure.png" pageId="24" pageNumber="225">Figs 75</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984AFF92FE768D80FE37EAC6" box="[395,426,1443,1468]" captionStart="FIGURES 84–87" captionStartId="39.[151,264,1427,1450]" captionTargetBox="[151,1435,194,1406]" captionTargetId="figure@39.[151,1436,193,1406]" captionTargetPageId="39" captionText="FIGURES 84–87. Female synthorax lateral: (84) L pendleburyi SAR16_GOM8; (85) L. schieli paratype SAR15_GOM6; (86) L. williamsoni SAR13_14_GOM51, right hand side, flipped for consistency;(87) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068038/files/figure.png" pageId="24" pageNumber="225">85</figureCitation>
|
||
). Prothorax with stripe on anterior pronotal lobe all yellow, tiny yellow marks on anterior dorsum middle pronotal lobe. Dorsal thoracic stripe just touching mesothoracic collar on left. Trochanters with some pale marks. Wings: sectors of arculus with 6 cross veins up to first bifurcation of superior sector in left Fw. 13 (right) or 14 (left) Ax in Fw,
|
||
<quantity id="4C92B5FF984AFF92FE218E2DFD89E95C" box="[476,532,1549,1574]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="225" unit="in" value="10.0">10 in</quantity>
|
||
Hw, 10 (left) or 13 (right) Px in Fw, 10 (right) or 11 (left) in Hw. Pt pale brown.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8E12FAE5E864" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A8E12FEACE930" box="[199,305,1585,1610]" italics="true" pageId="24" pageNumber="225">Abdomen</emphasis>
|
||
. Very gradually tapering from base to S8, S9 constricted at base then expanded again apically. Black with more extensive pale markings than male: S1 almost entirely yellowish. S2 with lateral yellow stripe running length of segment, including small, rounded auricle, stripe narrowly divided immediately after auricle. S3 with stripe running from base for most of segment, narrowly divided before its midpoint. S4–7 with basal lateral spot and small streak just after midpoint, the latter, diminishing in size on successive segments, tiny and very faint on S7. Middorsal stripe narrow but wider than in male, running S2–7. Cerci black, tapering to sharp slightly upturned tip, ca same length as S10. Vulvar scale (
|
||
<figureCitation id="1351049F984AFF92FD9F8F26FD5FE864" box="[610,706,1797,1822]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="24" pageNumber="225">Fig. 123</figureCitation>
|
||
) as long as sternum of S9, divided at ca three-quarters length.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8F09FB9AE838" blockId="24.[151,1437,151,1964]" box="[199,1031,1833,1858]" pageId="24" pageNumber="225">Measurements (mm). Hw 27.5, abdomen excluding anal appendages 29.5.</paragraph>
|
||
<paragraph id="8BD5181A984AFF92FF3A8F6FFB19E8D1" blockId="24.[151,1437,151,1964]" pageId="24" pageNumber="225">
|
||
<emphasis id="B91EC408984AFF92FF3A8F6FFEA2E81F" bold="true" box="[199,319,1868,1893]" pageId="24" pageNumber="225">Variation.</emphasis>
|
||
The second female
|
||
<typeStatus id="54D1A6B8984AFF92FDE38F6EFDE3E81F" box="[542,638,1869,1893]" pageId="24" pageNumber="225" type="paratype">paratype</typeStatus>
|
||
is very similar to the one described, but has no central pale mark on the postclypeus, the dorsal thoracic stripe is connected to mesothoracic collar on the right rather than left, the non-basal lateral streak on S7 is larger and brighter. 14–16 Ax in Fw, 12–13 Px in Fw,
|
||
<quantity id="4C92B5FF984AFF92FC108FB7FBD2E8D6" box="[1005,1103,1939,1964]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="24" pageNumber="225" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
|
||
Hw.
|
||
</paragraph>
|
||
<caption id="DF154892984BFF93FF6A8F96FAE8E890" httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="25" pageNumber="226" targetBox="[204,1387,183,1958]" targetPageId="25">
|
||
<paragraph id="8BD5181A984BFF93FF6A8F96FAE8E890" blockId="25.[151,1436,1973,2026]" pageId="25" pageNumber="226">
|
||
<emphasis id="B91EC408984BFF93FF6A8F96FEC4E8B1" bold="true" box="[151,345,1973,1996]" pageId="25" pageNumber="226">FIGURES 36–41.</emphasis>
|
||
Female head frontal: (36)
|
||
<taxonomicName id="4C6A6399984BFF93FD7E8F94FC99E8B6" authorityName="Laidlaw" authorityYear="1936" box="[643,772,1975,1996]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408984BFF93FD7E8F94FC99E8B6" box="[643,772,1975,1996]" italics="true" pageId="25" pageNumber="226">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM48; (37)
|
||
<taxonomicName id="4C6A6399984BFF93FBD78F94FB0FE8B6" box="[1066,1170,1975,1996]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408984BFF93FBD78F94FB0FE8B6" box="[1066,1170,1975,1996]" italics="true" pageId="25" pageNumber="226">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
holotype; (38)
|
||
<emphasis id="B91EC408984BFF93FAC28F94FAC9E8B6" box="[1343,1364,1975,1996]" italics="true" pageId="25" pageNumber="226">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A6399984BFF93FF6A8FF6FF65E890" authorityName="Laidlaw" authorityYear="1936" box="[151,248,2005,2026]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408984BFF93FF6A8FF6FF65E890" box="[151,248,2005,2026]" italics="true" pageId="25" pageNumber="226">coomansi</emphasis>
|
||
</taxonomicName>
|
||
PomS260; (39)
|
||
<taxonomicName id="4C6A6399984BFF93FE638FF6FE6DE890" authorityName="van Tol" authorityYear="1990" box="[414,496,2005,2026]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="pasia">
|
||
<emphasis id="B91EC408984BFF93FE638FF6FE6DE890" box="[414,496,2005,2026]" italics="true" pageId="25" pageNumber="226">L. pasia</emphasis>
|
||
</taxonomicName>
|
||
holotype; (40)
|
||
<taxonomicName id="4C6A6399984BFF93FD708FF6FD22E890" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[653,703,2005,2026]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="sii">
|
||
<emphasis id="B91EC408984BFF93FD708FF6FD22E890" box="[653,703,2005,2026]" italics="true" pageId="25" pageNumber="226">L. sii</emphasis>
|
||
</taxonomicName>
|
||
paratype SAR13_14_GOM44; (41)
|
||
<emphasis id="B91EC408984BFF93FBCE8FF6FBD5E890" box="[1075,1096,2005,2026]" italics="true" pageId="25" pageNumber="226">L.</emphasis>
|
||
species SAR11_12_GOM33.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A9848FF90FF3A88BBFC40EFCA" blockId="26.[151,1437,151,608]" box="[199,989,151,176]" pageId="26" pageNumber="227">Measurements (mm). Hw 28, abdomen excluding anal appendages 30.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919848FF90FF3A889FFDD6ED1A" pageId="26" pageNumber="227" type="discussion">
|
||
<paragraph id="8BD5181A9848FF90FF3A889FFB30EED7" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="227">
|
||
<emphasis id="B91EC4089848FF90FF3A889FFEA6EFAF" bold="true" box="[199,315,188,213]" pageId="26" pageNumber="227">Remarks.</emphasis>
|
||
The females are associated with the male by reasonable supposition; the seepage in which they were caught is very close to the location where the male was taken, and they resemble the male in the length of the dorsal thoracic stripes, longer than those in the most similar species,
|
||
<taxonomicName id="4C6A63999848FF90FCAF8926FC6EEE67" authorityName="Laidlaw" authorityYear="1912" box="[850,1011,260,285]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089848FF90FCAF8926FC6EEE67" box="[850,1011,260,285]" italics="true" pageId="26" pageNumber="227">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
, or the females treated as
|
||
<emphasis id="B91EC4089848FF90FAE78926FAACEE66" box="[1306,1329,261,284]" italics="true" pageId="26" pageNumber="227">L.</emphasis>
|
||
sp. sp. cf
|
||
<taxonomicName id="4C6A63999848FF90FF6A8904FE87EE3A" authorityName="Laidlaw" authorityYear="1912" box="[151,282,295,320]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089848FF90FF6A8904FE87EE3A" box="[151,282,295,320]" italics="true" pageId="26" pageNumber="227">williamsoni</emphasis>
|
||
</taxonomicName>
|
||
below, also from Gunung Penrissen. They also differ from both
|
||
<taxonomicName id="4C6A63999848FF90FC17890AFB16EE3A" authorityName="Laidlaw" authorityYear="1912" box="[1002,1163,295,320]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089848FF90FC17890AFB16EE3A" box="[1002,1163,295,320]" italics="true" pageId="26" pageNumber="227">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91EC4089848FF90FB3D890AFB4AEE3A" box="[1216,1239,297,320]" italics="true" pageId="26" pageNumber="227">L.</emphasis>
|
||
sp. cf
|
||
<emphasis id="B91EC4089848FF90FADC8904FA01EE3A" box="[1313,1436,295,320]" italics="true" pageId="26" pageNumber="227">wiliamsoni</emphasis>
|
||
in the structures on the occiput and vertex; lacking occipital spurs and with markedly different tubercles on the vertex, and with much wider spacing between the occipital horns than in
|
||
<emphasis id="B91EC4089848FF90FC238952FC68EEF2" box="[990,1013,369,392]" italics="true" pageId="26" pageNumber="227">L.</emphasis>
|
||
sp. cf
|
||
<emphasis id="B91EC4089848FF90FBB9894CFB22EEF2" box="[1092,1215,367,392]" italics="true" pageId="26" pageNumber="227">wiliamsoni</emphasis>
|
||
. The bulges on the rear occiput below the compound eyes are also not present on the taxa with similar markings.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9848FF90FF3A8994FD14EE8E" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="227">
|
||
<taxonomicName id="4C6A63999848FF90FF3A8994FE26EEAA" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[199,443,439,464]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089848FF90FF3A8994FE26EEAA" box="[199,443,439,464]" italics="true" pageId="26" pageNumber="227">Leptogomphus schieli</emphasis>
|
||
</taxonomicName>
|
||
was collected too late to include in the molecular analysis, but can be expected to cluster with
|
||
<taxonomicName id="4C6A63999848FF90FF3289FEFEEDEE8F" authorityName="Laidlaw" authorityYear="1912" box="[207,368,476,501]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089848FF90FF3289FEFEEDEE8F" box="[207,368,476,501]" italics="true" pageId="26" pageNumber="227">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91EC4089848FF90FE5B89FEFE20EE8E" box="[422,445,477,500]" italics="true" pageId="26" pageNumber="227">L.</emphasis>
|
||
sp. cf
|
||
<emphasis id="B91EC4089848FF90FDFA89FFFD1FEE8F" box="[519,642,476,501]" italics="true" pageId="26" pageNumber="227">wiliamsoni</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9848FF90FF3A8A23FDD6ED1A" blockId="26.[151,1437,151,608]" pageId="26" pageNumber="227">
|
||
At present
|
||
<taxonomicName id="4C6A63999848FF90FEBF8A22FE35ED62" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[322,424,511,536]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089848FF90FEBF8A22FE35ED62" box="[322,424,511,536]" italics="true" pageId="26" pageNumber="227">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
is only known from near the peak of Gunung Penrissen in south-west
|
||
<collectingCountry id="F37D588A9848FF90FB4389DCFA83ED62" box="[1214,1310,511,536]" name="Malaysia" pageId="26" pageNumber="227">
|
||
<collectingRegion id="49AED6F89848FF90FB4389DCFA83ED62" box="[1214,1310,511,536]" country="Malaysia" name="Sarawak" pageId="26" pageNumber="227">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
(
|
||
<figureCitation id="1351049F9848FF90FAD089DCFA12ED62" box="[1325,1423,511,536]" captionStart="FIGURE128" captionStartId="50.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1589]" captionTargetId="figure@50.[151,1436,193,1589]" captionTargetPageId="50" captionText="FIGURE128. Distributionof L. pendleburyi (yellowcircle), L. schieli (black square) and L. species from Pulong Tau National Park, Sarawak (white square)." httpUri="https://zenodo.org/record/1068058/files/figure.png" pageId="26" pageNumber="227">Fig. 128</figureCitation>
|
||
), at an altitude of approximately
|
||
<quantity id="4C92B5FF9848FF90FDF98A07FDFFED46" box="[516,610,548,572]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="227" unit="m" value="1.1">1,100 m</quantity>
|
||
a.s.l. It should be looked for in the nearby Bungo Range, and over the border in
|
||
<collectingRegion id="49AED6F89848FF90FEF98A64FE1AED1A" box="[260,391,583,608]" country="Indonesia" name="Kalimantan Utara" pageId="26" pageNumber="227">Kalimantan</collectingRegion>
|
||
on Gunung Niut.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |