190 lines
31 KiB
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190 lines
31 KiB
XML
<document id="E6DE75363F4BB8ACC0D3673E6EEA9FA9" ID-DOI="10.5281/zenodo.199508" ID-GBIF-Dataset="98bc6420-5732-4771-af50-30adacbc87b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="199508" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460338355409" checkinUser="plazi" docAuthor="Johanson, Kjell Arne & Holzenthal, Ralph W." docDate="2010" docId="03C387C4FF8AB40D1EBF8FCBD1D8FF62" docLanguage="en" docName="zt02689p047.pdf" docOrigin="Zootaxa 2689" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Helicopsyche alajuela Johanson & Holzenthal, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="39" masterDocId="FFFAFFBCFF8BB40F1E288C6ED201FFB5" masterDocTitle="The snail-case caddisfly subgenus Helicopsyche (Feropsyche) in Costa Rica, with the description of 3 new species (Trichoptera: Helicopsychidae)" masterLastPageNumber="47" masterPageNumber="37" pageNumber="38" updateTime="1698260395609" updateUser="plazi">
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<mods:titleInfo id="E819C8B16FBA196A762BEA81EA7FAE22">
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<mods:title id="70CC085FC9AE3A682805A0970DE40C06">The snail-case caddisfly subgenus Helicopsyche (Feropsyche) in Costa Rica, with the description of 3 new species (Trichoptera: Helicopsychidae)</mods:title>
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<mods:namePart id="DA92A528D3FE2F6596262A3973228D22">Johanson, Kjell Arne</mods:namePart>
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<mods:name id="871459CF56F13822FBC680A41022DD4A" type="personal">
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<mods:roleTerm id="575EF28B66651A835DA09652F727B529">Author</mods:roleTerm>
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<mods:namePart id="D0C4CC82E456F22E23F87E7E20148340">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="DF2A7CDFE451B4F64307B409303C7868">text</mods:typeOfResource>
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<mods:titleInfo id="019C02558BAAD14DC1ABD682904393B4">
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<mods:title id="F3EF854CF9FA05DB430FA564A55DAB88">Zootaxa</mods:title>
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<mods:part id="D1B109D77564A7B3BFC49829D964C5A9">
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<mods:date id="ED10B99D0D940BB097DE94A94604C49B">2010</mods:date>
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<mods:detail id="C03975C813BEADECCD7793CDA17A4AD1" type="volume">
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<mods:number id="B55E13CDC35C83AE87C688081B73DC04">2689</mods:number>
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<mods:classification id="DB7B2FFD6B0796BCD93CFA18942E5765">journal article</mods:classification>
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<mods:identifier id="C9B318F46DBC63E4FB63CCC2AC08755B" type="DOI">10.5281/zenodo.199508</mods:identifier>
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<mods:identifier id="47C0CF244EC792017BB7AE2AF098E6A2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3B22CCE12187708B26476447194D29C4" type="Zenodo-Dep">199508</mods:identifier>
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<treatment id="03C387C4FF8AB40D1EBF8FCBD1D8FF62" ID-DOI="http://doi.org/10.5281/zenodo.6204556" ID-GBIF-Taxon="119405574" ID-Zenodo-Dep="6204556" LSID="urn:lsid:plazi:treatment:03C387C4FF8AB40D1EBF8FCBD1D8FF62" httpUri="http://treatment.plazi.org/id/03C387C4FF8AB40D1EBF8FCBD1D8FF62" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
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<subSubSection id="C3706559FF8AB40E1EBF8FCBD325FC51" pageId="1" pageNumber="38" type="nomenclature">
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<paragraph id="8BD536D2FF8AB40E1EBF8FCBD03EFC0A" blockId="1.[151,575,933,996]" box="[151,575,933,959]" pageId="1" pageNumber="38">
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<heading id="D09D81BEFF8AB40E1EBF8FCBD03EFC0A" bold="true" box="[151,575,933,959]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
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<emphasis id="B91EEAC0FF8AB40E1EBF8FCBD03EFC0A" bold="true" box="[151,575,933,959]" pageId="1" pageNumber="38">
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<taxonomicName id="4C6A4D51FF8AB40E1EBF8FCBD3A0FC0A" box="[151,417,933,959]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="alajuela" status="sp. nov.">
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<emphasis id="B91EEAC0FF8AB40E1EBF8FCBD3A0FC0A" bold="true" box="[151,417,933,959]" italics="true" pageId="1" pageNumber="38">Helicopsyche alajuela</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22D57BBFF8AB40E1F988FCBD03EFC0A" box="[432,575,933,959]" pageId="1" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD536D2FF8AB40E1EBF8FA4D325FC51" blockId="1.[151,575,933,996]" box="[151,292,970,996]" pageId="1" pageNumber="38">
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<figureCitation id="13512A57FF8AB40E1EBF8FA4D2FBFC51" box="[151,250,970,996]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 1–7</figureCitation>
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,
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<figureCitation id="13512A57FF8AB40E1F2F8FA4D325FC51" box="[263,292,970,996]" captionStart="FIGURE 22" captionStartId="6.[151,255,1739,1763]" captionText="FIGURE 22. Map of Costa Rica showing distributions of H. alajuela, new species (filled rectangle), H. golfitoensis, new species (filled circle), and H. dorsocurvata, new species (filled diamond)." pageId="1" pageNumber="38">22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706559FF8AB40E1EBF887BD6A8F8D1" pageId="1" pageNumber="38" type="reference_group">
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<paragraph id="8BD536D2FF8AB40E1EBF887BD64CFB2A" blockId="1.[151,1437,1045,2004]" pageId="1" pageNumber="38">
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<taxonomicName id="4C6A4D51FF8AB40E1EBF887BD39CFB9B" box="[151,413,1045,1070]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="alajuela">
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<emphasis id="B91EEAC0FF8AB40E1EBF887BD39CFB9B" box="[151,413,1045,1070]" italics="true" pageId="1" pageNumber="38">Helicopsyche alajuela</emphasis>
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</taxonomicName>
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is most similar to
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<emphasis id="B91EEAC0FF8AB40E1C51887BD08FFB9B" box="[633,654,1045,1070]" italics="true" pageId="1" pageNumber="38">H</emphasis>
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.
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<emphasis id="B91EEAC0FF8AB40E1CB6887BD11BFB9B" box="[670,794,1045,1070]" italics="true" pageId="1" pageNumber="38">chipoensis</emphasis>
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<bibRefCitation id="EFFB4B23FF8AB40E1D09887BD646FB9A" author="Johanson" box="[801,1095,1045,1071]" pageId="1" pageNumber="38" refString="Johanson, K. A. & Malm, T. (2006) Seven new Helicopsyche (Feropsyche) Johanson, 2002 from the Neotropical region and Nearctic Mexico (Insecta: Trichoptera: Helicopsychidae). Zootaxa, 1208, 1 - 24." type="journal article" year="2006">Johanson & Malm, 2006</bibRefCitation>
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,
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<taxonomicName id="4C6A4D51FF8AB40E1A7C887BD335FBE1" authority="Johanson & Malm, 2006" authorityName="Johanson & Malm" authorityYear="2006" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="chiriquensis">
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<emphasis id="B91EEAC0FF8AB40E1A7C887BD709FB9B" box="[1108,1288,1045,1070]" italics="true" pageId="1" pageNumber="38">H. chiriquensis</emphasis>
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<bibRefCitation id="EFFB4B23FF8AB40E1B38887BD335FBE1" author="Johanson" pageId="1" pageNumber="38" refString="Johanson, K. A. & Malm, T. (2006) Seven new Helicopsyche (Feropsyche) Johanson, 2002 from the Neotropical region and Nearctic Mexico (Insecta: Trichoptera: Helicopsychidae). Zootaxa, 1208, 1 - 24." type="journal article" year="2006">Johanson & Malm, 2006</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B91EEAC0FF8AB40E1F698854D3B5FBE6" box="[321,436,1082,1107]" italics="true" pageId="1" pageNumber="38">H. perija</emphasis>
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<bibRefCitation id="EFFB4B23FF8AB40E1F958854D139FBE1" author="Johanson" box="[445,824,1082,1108]" pageId="1" pageNumber="38" refString="Johanson, K. A. & Holzenthal, R. W. (2004) Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae). Zootaxa, 711, 1 - 40." type="journal article" year="2004">Johanson & Holzenthal, 2004</bibRefCitation>
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, and
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<emphasis id="B91EEAC0FF8AB40E1D548854D615FBE6" box="[892,1044,1082,1107]" italics="true" pageId="1" pageNumber="38">H. circulata</emphasis>
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<bibRefCitation id="EFFB4B23FF8AB40E1A358854D799FBE1" author="Johanson" box="[1053,1432,1082,1108]" pageId="1" pageNumber="38" refString="Johanson, K. A. & Holzenthal, R. W. (2004) Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae). Zootaxa, 711, 1 - 40." type="journal article" year="2004">Johanson & Holzenthal, 2004</bibRefCitation>
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, particularly due to the similar shape of the primary branch of the gonopods. The new species is easily distinguished from these by the absence of the basomesal lobes on the gonopods.
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</paragraph>
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<paragraph id="8BD536D2FF8AB40E1EEE88C4D127FB71" blockId="1.[151,1437,1045,2004]" box="[198,806,1194,1220]" pageId="1" pageNumber="38">
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Forewing length
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<quantity id="4C929B37FF8AB40E1FA688C4D3EAFB71" box="[398,491,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="38" unit="mm" value="3.4">3.4 mm</quantity>
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, hind wing length
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<quantity id="4C929B37FF8AB40E1CEC88C4D123FB71" box="[708,802,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="38" unit="mm" value="2.4">2.4 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BD536D2FF8AB40E1EEE88BED6A8F8D1" blockId="1.[151,1437,1045,2004]" pageId="1" pageNumber="38">
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Male abdomen and genitalia (
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<figureCitation id="13512A57FF8AB40E1C6888BED0ACFB5F" box="[576,685,1232,1258]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 1–7</figureCitation>
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). Sternal process VI long, straight, oriented posteroventrad; parallel-sided in lateral view (
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<figureCitation id="13512A57FF8AB40E1FD4889BD042FABA" box="[508,579,1269,1295]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
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); weakly tapering in ventral view (
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<figureCitation id="13512A57FF8AB40E1DC0889BD632FABA" box="[1000,1075,1269,1295]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 2</figureCitation>
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). Anterior lobe of segment IX (
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<figureCitation id="13512A57FF8AB40E1EB78974D2E7FA81" box="[159,230,1306,1332]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
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) narrowly ellipsoid in lateral view, oriented anterodorsad from midheight of segment IX; anterodorsal and anteroventral margins concave; ventral margin long, straight (
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<figureCitation id="13512A57FF8AB40E1DFC892ED620FAEF" box="[980,1057,1344,1370]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
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); in dorsal view, inner margin forming diamond-shaped cavity (
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<figureCitation id="13512A57FF8AB40E1C37890BD068FACA" box="[543,617,1381,1407]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 4</figureCitation>
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); in ventral view, with small central posterior process (
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<figureCitation id="13512A57FF8AB40E1ADB890BD73AFACA" box="[1267,1339,1381,1407]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 5</figureCitation>
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); lateral apodeme nearly horizontal (
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<figureCitation id="13512A57FF8AB40E1FC189E4D033FA11" box="[489,562,1418,1444]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
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), tapering anteriorly, meeting anterior margin, sub-marginal line absent; tergal transverse apodeme absent; sternal transverse apodemes present. Tergum X, in lateral view, oriented posteroventrad (
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<figureCitation id="13512A57FF8AB40E1F7689BBD3A8FA5A" box="[350,425,1493,1519]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
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), straight, apex rounded, weakly tapering along its length (
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<figureCitation id="13512A57FF8AB40E1AAA89BBD6CEFA5A" box="[1154,1231,1493,1519]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 4</figureCitation>
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); in dorsal view, tongue-shaped (
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<figureCitation id="13512A57FF8AB40E1F748994D3A8F9A1" box="[348,425,1530,1556]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 4</figureCitation>
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), apical part shallowly notched, with about 13 pairs of equally long megasetae in longitudinal group, starting basally on segment. Superior appendage tubular (
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<figureCitation id="13512A57FF8AB40E1A7E8A4ED6A5F98F" box="[1110,1188,1568,1594]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 4</figureCitation>
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||
). Primary branch of gonopods, in lateral view (
|
||
<figureCitation id="13512A57FF8AB40E1FF88A2BD016F9EA" box="[464,535,1605,1631]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
|
||
), strongly broadened distad, narrowest immediately behind posterior margin of segment IX; dorsal margins semi-circular, weakly undulating; ventral margin irregularly undulating (
|
||
<figureCitation id="13512A57FF8AB40E1B448A04D2A9F91F" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
|
||
); apices produced mesad. Basomesal lobes absent; posterobasal margins strongly undulating, with setae (
|
||
<figureCitation id="13512A57FF8AB40E1E888ADBD2EBF97A" box="[160,234,1717,1743]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 5</figureCitation>
|
||
). Basal plate, in lateral view (
|
||
<figureCitation id="13512A57FF8AB40E1C7D8ADBD09CF97A" box="[597,669,1717,1743]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 3</figureCitation>
|
||
), nearly straight, wide along its length; in ventral view, sharply triangular (
|
||
<figureCitation id="13512A57FF8AB40E1F358AB4D366F941" box="[285,359,1754,1780]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 5</figureCitation>
|
||
). Phallic apparatus, lateral view, wide proximally (
|
||
<figureCitation id="13512A57FF8AB40E1DF98AB4D61AF941" box="[977,1051,1754,1780]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 6</figureCitation>
|
||
), narrowest at mid-length, apex about as broad as base; slightly curving ventrad along its length; in ventral view, with basis narrow; (
|
||
<figureCitation id="13512A57FF8AB40E1B638B6ED793F8AF" box="[1355,1426,1792,1818]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 7</figureCitation>
|
||
) slightly wider proximally and distally; endotheca weakly produced, posteroventral part well sclerotized; sperm channel slender anteriorly before sharply widening at 1/3rd phallic length (
|
||
<figureCitation id="13512A57FF8AB40E1A7B8B24D69AF8D1" box="[1107,1179,1866,1892]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,270,1597,1621]" captionTargetBox="[154,1431,272,1553]" captionTargetId="figure@2.[151,1436,259,1573]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Helicopsyche alajuela, new species, holotype. 1 — sternum VI, lateral; 2 — sternum VI process, ventral; 3 — genitalia, lateral; 4 — genitalia, dorsal; 5 — genitalia, ventral; 6 — phallic apparatus, lateral; 7 — phallic apparatus, ventral. Scale bar (0.1 mm) refers to all figures." httpUri="https://zenodo.org/record/199509/files/figure.png" pageId="1" pageNumber="38">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706559FF8AB40E1EED8B01D641F861" pageId="1" pageNumber="38" type="description">
|
||
<paragraph id="8BD536D2FF8AB40E1EED8B01D6FDF81A" blockId="1.[151,1437,1045,2004]" pageId="1" pageNumber="38">
|
||
<emphasis id="B91EEAC0FF8AB40E1EED8B01D0CEF83C" bold="true" box="[197,719,1903,1929]" pageId="1" pageNumber="38">
|
||
Male
|
||
<typeStatus id="54D18870FF8AB40E1F268B01D37FF83C" box="[270,382,1903,1929]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
.
|
||
<collectingCountry id="F37D7642FF8AB40E1FA48B01D04DF83C" box="[396,588,1903,1929]" name="Costa Rica" pageId="1" pageNumber="38">COSTA RICA</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C6A4D51FF8AB40E1C718B01D0CBF83C" box="[601,714,1903,1929]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="43" phylum="Arthropoda" rank="species" species="alajuela" status="sp. nov.">Alajuela</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
Reserva Forestal San Ramón, Río San Lorencito and tribs,
|
||
<geoCoordinate id="EE5E5015FF8AB40E1EBF8BFBD30DF81A" box="[151,268,1941,1967]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="55" value="10.216">10.216°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E5015FF8AB40E1F3F8BFBD38EF81A" box="[279,399,1941,1967]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="55" value="-84.607">84.607°W</geoCoordinate>
|
||
,
|
||
<date id="FFD41012FF8AB40E1FB38BFBD046F81A" box="[411,583,1941,1967]" pageId="1" pageNumber="38" value="1988-06-13" valueMax="1988-06-16" valueMin="1988-06-13">
|
||
13–16.vi.
|
||
<quantity id="4C929B37FF8AB40E1C2F8BFBD046F81A" box="[519,583,1941,1967]" metricMagnitude="6" metricUnit="m" metricValue="1.98898" pageId="1" pageNumber="38" unit="m" value="1988980.0">1988</quantity>
|
||
</date>
|
||
, 980 m [C.M. & O.S. Flint, Holzenthal] (NMNH, alcohol).
|
||
</paragraph>
|
||
<paragraph id="8BD536D2FF8AB40E1EED8BD4D641F861" blockId="1.[151,1437,1045,2004]" box="[197,1088,1978,2004]" pageId="1" pageNumber="38">
|
||
<emphasis id="B91EEAC0FF8AB40E1EED8BD4D33FF861" bold="true" box="[197,318,1978,2004]" pageId="1" pageNumber="38">
|
||
<typeStatus id="54D18870FF8AB40E1EED8BD4D338F861" box="[197,313,1978,2004]" pageId="1" pageNumber="38" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
same data as
|
||
<typeStatus id="54D18870FF8AB40E1FC98BD4D04AF861" box="[481,587,1978,2004]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
, abdomen lost—
|
||
<specimenCount id="9D6CFD5BFF8AB40E1D3E8BD4D15EF861" box="[790,863,1978,2004]" pageId="1" pageNumber="38" type="male">1 male</specimenCount>
|
||
(NMNH, alcohol).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706559FF89B40D1EED8CF9D0E1FF07" box="[197,736,151,178]" pageId="2" pageNumber="39" type="distribution">
|
||
<paragraph id="8BD536D2FF89B40D1EED8CF9D0E1FF07" blockId="2.[197,985,151,215]" box="[197,736,151,178]" pageId="2" pageNumber="39">
|
||
<emphasis id="B91EEAC0FF89B40D1EED8CF9D364FF04" bold="true" box="[197,357,151,177]" pageId="2" pageNumber="39">Distribution.</emphasis>
|
||
<collectingCountry id="F37D7642FF89B40D1F448CF6D3EDFF07" box="[364,492,152,178]" name="Costa Rica" pageId="2" pageNumber="39">Costa Rica</collectingCountry>
|
||
(
|
||
<taxonomicName id="4C6A4D51FF89B40D1FD38CF6D063FF07" box="[507,610,152,178]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="alajuela" status="sp. nov.">Alajuela</taxonomicName>
|
||
) (
|
||
<figureCitation id="13512A57FF89B40D1C528CF6D0D3FF07" box="[634,722,152,178]" captionStart="FIGURE 22" captionStartId="6.[151,255,1739,1763]" captionText="FIGURE 22. Map of Costa Rica showing distributions of H. alajuela, new species (filled rectangle), H. golfitoensis, new species (filled circle), and H. dorsocurvata, new species (filled diamond)." pageId="2" pageNumber="39">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706559FF89B40D1EED8CD3D1D8FF62" box="[197,985,189,215]" pageId="2" pageNumber="39" type="etymology">
|
||
<paragraph id="8BD536D2FF89B40D1EED8CD3D1D8FF62" blockId="2.[197,985,151,215]" box="[197,985,189,215]" pageId="2" pageNumber="39">
|
||
<emphasis id="B91EEAC0FF89B40D1EED8CD3D350FF62" bold="true" box="[197,337,189,215]" pageId="2" pageNumber="39">Etymology.</emphasis>
|
||
<taxonomicName id="4C6A4D51FF89B40D1F718CD3D3BDFF63" box="[345,444,189,214]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="alajuela" status="sp. nov.">
|
||
<emphasis id="B91EEAC0FF89B40D1F718CD3D3BDFF63" box="[345,444,189,214]" italics="true" pageId="2" pageNumber="39">Alajuela</emphasis>
|
||
</taxonomicName>
|
||
, named after the province of the
|
||
<typeStatus id="54D18870FF89B40D1D698CD3D172FF62" box="[833,883,189,215]" pageId="2" pageNumber="39">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |