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<document id="46509B3D6F6C279F8721DEF40D412B4F" ID-CLB-Dataset="9034" ID-DOI="10.11646/zootaxa.4881.1.5" ID-GBIF-Dataset="3a4a468d-2333-40c1-987a-ab7c5064c724" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425815" ID-ZooBank="E7296FA6-BF7A-4BC8-BDE3-F600A38F4135" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610043453082" checkinUser="plazi" docAuthor="Muadsub, Sopita &amp; Pinkaew, Nantasak" docDate="2020" docId="03C329786F13FFCE6BD492FF984088EE" docLanguage="en" docName="zootaxa.4881.1.5.pdf" docOrigin="Zootaxa 4881 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ukamenia thailandica Muadsub &amp; Pinkaew 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="87" masterDocId="FFFA51006F12FFCA6B4392689B31893F" masterDocTitle="Ukamenia Oku, 1981 (Lepidoptera: Tortricidae: Olethreutinae), new record for Thailand with descriptions of a new species" masterLastPageNumber="90" masterPageNumber="83" pageNumber="84" updateTime="1698923940840" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1696ECAA83A5047B6B032B11A9788BE0">Ukamenia Oku, 1981 (Lepidoptera: Tortricidae: Olethreutinae), new record for Thailand with descriptions of a new species</mods:title>
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<mods:namePart id="766490002E78D052D662A39BF24E574C">Muadsub, Sopita</mods:namePart>
<mods:nameIdentifier id="EC6E3EC8F4FE9046550CD28510716D4E" type="ORCID">0000-0002-9672-1878</mods:nameIdentifier>
<mods:affiliation id="DB11E20EB108F2CC8F3CDE118F92A03B">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Kamphaeng Saen Campus, Nakhon Pathom, 73140, Thailand. &amp; muadsub _ s @ hotmail. com; https: // orcid. org / 0000 - 0002 - 9672 - 1878</mods:affiliation>
<mods:nameIdentifier id="E77416F18CF1611A31894D16541859ED" type="email">muadsub_s@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="122E81BBAA77F2EB03F209E151B86890">Pinkaew, Nantasak</mods:namePart>
<mods:affiliation id="1D6D68794297B20C8B5559819BAF5463">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Kamphaeng Saen Campus, Nakhon Pathom, 73140, Thailand. &amp; Center for Advanced Studies in Tropical Natural Resources, NRU-KU, Kasetsart University, Chatuchak, Bangkok, 10900, Thailand</mods:affiliation>
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<mods:date id="2A658E474BC5BA05B5AA6F639BAAED6F">2020</mods:date>
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<taxonomicName id="4C6AE3ED6F13FFCB6BD492FF9A94898D" authority="Muadsub &amp; Pinkaew, 2020" authorityName="Muadsub &amp; Pinkaew" authorityYear="2020" box="[151,421,151,178]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B91E447C6F13FFCB6BD492FF9A94898D" bold="true" box="[151,421,151,178]" italics="true" pageId="1" pageNumber="84">Ukamenia thailandica</emphasis>
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<taxonomicNameLabel id="A22DF9076F13FFCB6AEF92FF9ADE898E" box="[428,495,151,177]" pageId="1" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C370CBE56F13FFCB6BD492B79A6989C5" box="[151,344,223,250]" pageId="1" pageNumber="84" type="description">
<paragraph id="8BD5986E6F13FFCB6BD492B79A6989C5" blockId="1.[151,981,151,250]" box="[151,344,223,250]" pageId="1" pageNumber="84">
(
<figureCitation id="135184EB6F13FFCB6BDC92B79BDC89C6" box="[159,237,223,249]" captionStart="FIGURES 12" captionStartId="1.[151,264,1687,1711]" captionTargetBox="[184,1403,453,1664]" captionTargetId="figure-89@1.[184,1403,453,1664]" captionTargetPageId="1" captionText="FIGURES 12. Living specimens in natural resting posture. 1. Ukamenia thailandica n. sp. 2. Gatesclarkeana idia." figureDoi="http://doi.org/10.5281/zenodo.4425819" httpUri="https://zenodo.org/record/4425819/files/figure.png" pageId="1" pageNumber="84">Figs. 1</figureCitation>
,
<figureCitation id="135184EB6F13FFCB6BB892B79A3B89C6" box="[251,266,223,249]" captionStart="FIGURES 34" captionStartId="2.[151,264,1651,1675]" captionTargetBox="[216,1369,193,1619]" captionTargetId="figure-27@2.[206,1382,181,1626]" captionTargetPageId="2" captionText="FIGURES 34. Adult specimens (scale bars = 2 mm). 3. Ukamenia thailandica n. sp. (holotype) 4. Gatesclarkeana idia." figureDoi="http://doi.org/10.5281/zenodo.4425821" httpUri="https://zenodo.org/record/4425821/files/figure.png" pageId="1" pageNumber="84">3</figureCitation>
,
<figureCitation id="135184EB6F13FFCB6A5492B79A7E89C5" box="[279,335,223,250]" captionStart-0="FIGURES 57" captionStart-1="FIGURES 810" captionStart-2="FIGURES 1115" captionStart-3="FIGURES 1618" captionStartId-0="3.[151,264,1581,1605]" captionStartId-1="4.[151,264,1960,1984]" captionStartId-2="5.[151,264,1940,1964]" captionStartId-3="6.[151,264,1277,1301]" captionTargetBox-0="[155,1431,597,1558]" captionTargetBox-1="[310,1277,489,1937]" captionTargetBox-2="[220,1367,181,1916]" captionTargetBox-3="[255,1338,181,1248]" captionTargetId-0="figure-226@3.[155,1431,597,1558]" captionTargetId-1="figure-159@4.[310,1277,489,1937]" captionTargetId-2="figure-17@5.[220,1367,181,1916]" captionTargetId-3="figure-27@6.[250,1338,181,1253]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 57. Head and hindwing of Ukamenia thailandica n. sp. (holotype). 5. Composition of head. 6.Underside of hindwing. 7. Curved up scales on the underside of hindwing." captionText-1="FIGURES 810. Hind tibia of Ukamenia thailandica n. sp. (holotype, left side). 8. Outer side. 9. Inner side. 10. Tuft of yellowish white hair pencils inside concavity." captionText-2="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." captionText-3="FIGURES 1618. Distinct characters on the abdomen of Ukamenia thailandica n. sp. (holotype). 16. Overall image of abdomen. 17. Small pits on tergum II. 18. Subcylindrical pouches on sternum III." figureDoi-0="http://doi.org/10.5281/zenodo.4425823" figureDoi-1="http://doi.org/10.5281/zenodo.4425825" figureDoi-2="http://doi.org/10.5281/zenodo.4425829" figureDoi-3="http://doi.org/10.5281/zenodo.4425831" httpUri-0="https://zenodo.org/record/4425823/files/figure.png" httpUri-1="https://zenodo.org/record/4425825/files/figure.png" httpUri-2="https://zenodo.org/record/4425829/files/figure.png" httpUri-3="https://zenodo.org/record/4425831/files/figure.png" pageId="1" pageNumber="84">518</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="B91E447C6F13FFCB6BD4934F9BE8887E" bold="true" box="[151,217,295,321]" pageId="1" pageNumber="84">Type.</emphasis>
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<typeStatus id="54D126CC6F13FFCB6BA3934F9A79887E" box="[224,328,295,321]" pageId="1" pageNumber="84" type="holotype">Holotype</typeStatus>
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.
<collectingCountry id="F37DD8FE6F13FFCB6A30934F9939887E" box="[371,520,295,321]" name="Thailand" pageId="1" pageNumber="84">THAILAND</collectingCountry>
:
<collectingRegion id="49AE568C6F13FFCB6950934F99D9887D" box="[531,744,295,322]" country="Thailand" name="Chaiyaphum" pageId="1" pageNumber="84">Chaiyaphum Prov.</collectingRegion>
:
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23ʹ13ʺN
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101
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35ʹ10ʺE
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, ca
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<collectorName id="269FFDB86F13FFCB6B9E93249A53885A" box="[221,354,331,358]" pageId="1" pageNumber="84">N. Pinkaew</collectorName>
; specimen no. np12599 (genitalia slide NP3904), deposited in
<collectionCode id="ED7B00AB6F13FFCB6F5993239F53885A" box="[1050,1122,331,357]" pageId="1" pageNumber="84">KKIC</collectionCode>
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.
<collectingCountry id="F37DD8FE6F13FFCB6E4793239EA8885A" box="[1284,1433,331,357]" name="Thailand" pageId="1" pageNumber="84">THAILAND</collectingCountry>
:
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:
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23ʹ13ʺN
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35ʹ10ʺE
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, ca
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<date id="FFD4BEAE6F13FFCB689593079F5888B6" box="[982,1129,367,394]" pageId="1" pageNumber="84" value="2018-12-11">
<collectingDate id="EF9047466F13FFCB689593079F5888B6" box="[982,1129,367,394]" pageId="1" pageNumber="84" value="2018-12-11">11 Dec 2018</collectingDate>
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<collectorName id="269FFDB86F13FFCB6F3793189FCD88B6" box="[1140,1276,367,394]" pageId="1" pageNumber="84">N. Pinkaew</collectorName>
; specimen no. np12600 (genitalia slide NP3905), deposited in
<collectionCode id="ED7B00AB6F13FFCB69EC93FB99C68892" box="[687,759,403,429]" pageId="1" pageNumber="84">KKIC</collectionCode>
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.
</paragraph>
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<caption id="DF15C8E66F13FFCB6BD494FF9E098F90" ID-DOI="http://doi.org/10.5281/zenodo.4425819" ID-Zenodo-Dep="4425819" box="[151,1336,1687,1712]" httpUri="https://zenodo.org/record/4425819/files/figure.png" pageId="1" pageNumber="84" startId="1.[151,264,1687,1711]" targetBox="[184,1403,453,1664]" targetPageId="1">
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<emphasis id="B91E447C6F13FFCB6BD494FF9A058F90" bold="true" box="[151,308,1687,1711]" pageId="1" pageNumber="84">FIGURES 12</emphasis>
. Living specimens in natural resting posture. 1.
<taxonomicName id="4C6AE3ED6F13FFCB685E94FF98CE8F90" authority="Muadsub &amp; Pinkaew, 2020" authorityName="Muadsub &amp; Pinkaew" authorityYear="2020" box="[797,1023,1687,1711]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B91E447C6F13FFCB685E94FF98CE8F90" box="[797,1023,1687,1711]" italics="true" pageId="1" pageNumber="84">Ukamenia thailandica</emphasis>
</taxonomicName>
<emphasis id="B91E447C6F13FFCB6F4694F09F0F8F8F" bold="true" box="[1029,1086,1688,1712]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A22DF9076F13FFCB6F4694F09F0F8F8F" box="[1029,1086,1688,1712]" pageId="1" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
2.
<taxonomicName id="4C6AE3ED6F13FFCB6F2794FF9E038F90" authorityName="Diakonoff" authorityYear="1973" box="[1124,1330,1687,1711]" class="Insecta" family="Tortricidae" genus="Gatesclarkeana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="idia">
<emphasis id="B91E447C6F13FFCB6F2794FF9E038F90" box="[1124,1330,1687,1711]" italics="true" pageId="1" pageNumber="84">Gatesclarkeana idia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C370CBE56F13FFCB6B8494B5981D8E5B" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="8BD5986E6F13FFCB6B8494B5981D8E5B" blockId="1.[151,1437,1757,2036]" pageId="1" pageNumber="84">
<emphasis id="B91E447C6F13FFCB6B8494B59A0B8FC8" bold="true" box="[199,314,1757,1783]" pageId="1" pageNumber="84">Diagnosis</emphasis>
. This species is most similar to
<taxonomicName id="4C6AE3ED6F13FFCB69E494B69FB98FC7" authority="(Matsumura, 1931)" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[679,1160,1758,1784]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B91E447C6F13FFCB69E494B698968FC8" box="[679,935,1758,1784]" italics="true" pageId="1" pageNumber="84">Ukamenia sapporensis</emphasis>
(
<bibRefCitation id="EFFBE59F6F13FFCB68F594B69F4E8FC7" author="Matsumura, S." box="[950,1151,1758,1784]" pageId="1" pageNumber="84" refId="ref2879" refString="Matsumura, S. (1931) 6000 Illustrated insects of Japan Empire. Toko Shoin, Tokyo, 1496 pp., 10 pls. [in Japanese]" type="book" year="1931">Matsumura, 1931</bibRefCitation>
)
</taxonomicName>
but male genitalia of
<taxonomicName id="4C6AE3ED6F13FFCB6EC294B69A268E23" authority="Muadsub &amp; Pinkaew, 2020" authorityName="Muadsub &amp; Pinkaew" authorityYear="2020" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B91E447C6F13FFCB6EC294B69A268E23" italics="true" pageId="1" pageNumber="84">U. thailandica</emphasis>
</taxonomicName>
<emphasis id="B91E447C6F13FFCB6A5D956A9A6C8E23" bold="true" box="[286,349,1794,1820]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A22DF9076F13FFCB6A5D956A9A6C8E23" box="[286,349,1794,1820]" pageId="1" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs by a pair of subelliptical pouch on abdominal sternum III (subcircular in
<taxonomicName id="4C6AE3ED6F13FFCB6FA6956A9EBD8E24" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[1253,1420,1794,1819]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B91E447C6F13FFCB6FA6956A9EBD8E24" box="[1253,1420,1794,1819]" italics="true" pageId="1" pageNumber="84">U. sapporensis</emphasis>
</taxonomicName>
), sacculus projecting lobe shorter than mid length of valva (longer than mid length of valva in
<taxonomicName id="4C6AE3ED6F13FFCB6FF7954E9E6D8E00" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[1204,1372,1830,1855]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B91E447C6F13FFCB6FF7954E9E6D8E00" box="[1204,1372,1830,1855]" italics="true" pageId="1" pageNumber="84">U. sapporensis</emphasis>
</taxonomicName>
), and subcone-shaped phallus (subcylindrical in
<taxonomicName id="4C6AE3ED6F13FFCB69369522982D8E5C" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[629,796,1866,1891]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="B91E447C6F13FFCB69369522982D8E5C" box="[629,796,1866,1891]" italics="true" pageId="1" pageNumber="84">U. sapporensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C370CBE56F13FFCE6B8495059BE58892" lastPageId="4" lastPageNumber="87" pageId="1" pageNumber="84" type="description">
<paragraph id="8BD5986E6F13FFCB6B8495059FA98ECB" blockId="1.[151,1437,1757,2036]" pageId="1" pageNumber="84">
<emphasis id="B91E447C6F13FFCB6B8495059A608EB8" bold="true" box="[199,337,1901,1927]" pageId="1" pageNumber="84">Description</emphasis>
. Head (
<figureCitation id="135184EB6F13FFCB6AEA95069ADC8EB7" box="[425,493,1901,1928]" captionStart="FIGURES 57" captionStartId="3.[151,264,1581,1605]" captionTargetBox="[155,1431,597,1558]" captionTargetId="figure-226@3.[155,1431,597,1558]" captionTargetPageId="3" captionText="FIGURES 57. Head and hindwing of Ukamenia thailandica n. sp. (holotype). 5. Composition of head. 6.Underside of hindwing. 7. Curved up scales on the underside of hindwing." figureDoi="http://doi.org/10.5281/zenodo.4425823" httpUri="https://zenodo.org/record/4425823/files/figure.png" pageId="1" pageNumber="84">Fig. 5</figureCitation>
): Lower frons light brown, upper frons light brown with dark brown laterally; vertex brown with brownish white tips; labial palpus porrect, first segment yellowish white, second segment with basal half narrow, yellowish white with a dark brown spot medially, apical half enlarged, light brown with a dark brown patch medially, apical segment small, greyish brown with white apex. Antenna light brown.
</paragraph>
<caption id="DF15C8E66F10FFC86BD4941B9E5B8FB4" ID-DOI="http://doi.org/10.5281/zenodo.4425821" ID-Zenodo-Dep="4425821" box="[151,1386,1651,1676]" httpUri="https://zenodo.org/record/4425821/files/figure.png" pageId="2" pageNumber="85" startId="2.[151,264,1651,1675]" targetBox="[216,1369,193,1619]" targetPageId="2">
<paragraph id="8BD5986E6F10FFC86BD4941B9E5B8FB4" blockId="2.[151,1386,1651,1676]" box="[151,1386,1651,1676]" pageId="2" pageNumber="85">
<emphasis id="B91E447C6F10FFC86BD4941B9A058FB4" bold="true" box="[151,308,1651,1675]" pageId="2" pageNumber="85">FIGURES 34</emphasis>
. Adult specimens (scale bars = 2 mm). 3.
<taxonomicName id="4C6AE3ED6F10FFC8699C941C98F18FB3" authority="Muadsub &amp; Pinkaew, 2020" authorityName="Muadsub &amp; Pinkaew" authorityYear="2020" box="[735,960,1652,1676]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B91E447C6F10FFC8699C941C98F18FB3" box="[735,960,1652,1676]" italics="true" pageId="2" pageNumber="85">Ukamenia thailandica</emphasis>
</taxonomicName>
<emphasis id="B91E447C6F10FFC86884941C9F318FB3" bold="true" box="[967,1024,1652,1676]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A22DF9076F10FFC86884941C9F318FB3" box="[967,1024,1652,1676]" pageId="2" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype) 4.
<taxonomicName id="4C6AE3ED6F10FFC86FD5941C9E558FB3" authorityName="Diakonoff" authorityYear="1973" box="[1174,1380,1652,1676]" class="Insecta" family="Tortricidae" genus="Gatesclarkeana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="idia">
<emphasis id="B91E447C6F10FFC86FD5941C9E558FB3" box="[1174,1380,1652,1676]" italics="true" pageId="2" pageNumber="85">Gatesclarkeana idia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD5986E6F10FFC96B8494D29BE38B01" blockId="2.[151,1437,1722,2036]" lastBlockId="3.[151,1437,151,574]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
Thorax: Pronotal collar brown; tegulae brown with basal 1/3 dark brown; mesonotum with anterior half brown, posterior half light brown, posterior crest brown with yellowish orange tips. Hind tibia enlarged with modified scales, outer surface covered with short appressed grey scales, inner surface shallow concavity covered with short, dark brown scales, basal half of concavity filled with tuft of very dense yellowish white hair pencils and top with elongate light brown scales (
<figureCitation id="135184EB6F10FFC86A9A952299628E5B" box="[473,595,1866,1892]" captionStart="FIGURES 810" captionStartId="4.[151,264,1960,1984]" captionTargetBox="[310,1277,489,1937]" captionTargetId="figure-159@4.[310,1277,489,1937]" captionTargetPageId="4" captionText="FIGURES 810. Hind tibia of Ukamenia thailandica n. sp. (holotype, left side). 8. Outer side. 9. Inner side. 10. Tuft of yellowish white hair pencils inside concavity." figureDoi="http://doi.org/10.5281/zenodo.4425825" httpUri="https://zenodo.org/record/4425825/files/figure.png" pageId="2" pageNumber="85">Figs. 810</figureCitation>
). Forewing broad subrectangular, length
<quantity id="4C92358B6F10FFC86F6395229F968E5B" box="[1056,1191,1866,1893]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.1" metricValueMin="5.9" pageId="2" pageNumber="85" unit="mm" value="6.0" valueMax="6.1" valueMin="5.9">5.96.1 mm</quantity>
in males (n = 2) (
<figureCitation id="135184EB6F10FFC86E2C95229B978EB7" captionStart="FIGURES 34" captionStartId="2.[151,264,1651,1675]" captionTargetBox="[216,1369,193,1619]" captionTargetId="figure-27@2.[206,1382,181,1626]" captionTargetPageId="2" captionText="FIGURES 34. Adult specimens (scale bars = 2 mm). 3. Ukamenia thailandica n. sp. (holotype) 4. Gatesclarkeana idia." figureDoi="http://doi.org/10.5281/zenodo.4425821" httpUri="https://zenodo.org/record/4425821/files/figure.png" pageId="2" pageNumber="85">Fig. 3</figureCitation>
), costa evenly curve; termen and wing apex rather round; costal strigulae well developed, pinkish red separated by blackish streaks; ground color light greyish brown mixed with scattered silvery spots and patches, costa with distinct pinkish red scales, slightly paler to wing apex and diffuse along termen, basal 1/5 with indistinct patch, brown to dark brown, with a moderately large silvery patch dorsobasally and small silvery spots scattered to dorsum, beyond with irregular transverse band, brown mixed with silvery patches, near margins with irregular, transverse narrow lines extending from costa to dorsum, dark brown, medially with large, irregular patch extending obliquely from middle of costa to near tornus, brown to dark brown mixed with silvery patches, outer margin edged with irregular transverse narrow line, dark brown, extending from base of R
<subScript id="17EE9A2B6F11FFC9681E937D9854881C" attach="left" box="[861,869,277,291]" fontSize="6" pageId="3" pageNumber="86">5</subScript>
to tornus, apical 1/4 with a silvery oblique patch extending from near costa at R
<subScript id="17EE9A2B6F11FFC96ABB935199318878" attach="left" box="[504,512,313,327]" fontSize="6" pageId="3" pageNumber="86">2</subScript>
to between R
<subScript id="17EE9A2B6F11FFC969E3935199998878" attach="left" box="[672,680,313,327]" fontSize="6" pageId="3" pageNumber="86">5</subScript>
and M
<subScript id="17EE9A2B6F11FFC969B8935198328878" attach="left" box="[763,771,313,327]" fontSize="6" pageId="3" pageNumber="86">1</subScript>
, continued with an irregular patch, dark brown, extending obliquely to near termen at M
<subScript id="17EE9A2B6F11FFC96AA993359AC38854" attach="left" box="[490,498,349,363]" fontSize="6" pageId="3" pageNumber="86">3</subScript>
, above with a small silvery patch near termen between R
<subScript id="17EE9A2B6F11FFC96F3E93359FB48854" attach="left" box="[1149,1157,349,363]" fontSize="6" pageId="3" pageNumber="86">5</subScript>
and M
<subScript id="17EE9A2B6F11FFC96F9093359FEA8854" attach="left" box="[1235,1243,349,363]" fontSize="6" pageId="3" pageNumber="86">2</subScript>
, tornal area with a large, transverse silvery patch extending obliquely from near base of M
<subScript id="17EE9A2B6F11FFC9689393E998E988B0" attach="left" box="[976,984,385,399]" fontSize="6" pageId="3" pageNumber="86">1</subScript>
to termen between CuA
<subScript id="17EE9A2B6F11FFC96FAF93E99FC588B0" attach="left" box="[1260,1268,385,399]" fontSize="6" pageId="3" pageNumber="86">2</subScript>
and CuP, wing apex with moderately subcircular patch, dark brown mixed with silvery scales, anal margin slight curve upwardly; fringe scales brown mixed with dark brown; underside light greyish brown, with yellowish white marks along costa. Hindwing (
<figureCitation id="135184EB6F11FFC96A5593B39A6688CA" box="[278,343,475,501]" captionStart="FIGURES 34" captionStartId="2.[151,264,1651,1675]" captionTargetBox="[216,1369,193,1619]" captionTargetId="figure-27@2.[206,1382,181,1626]" captionTargetPageId="2" captionText="FIGURES 34. Adult specimens (scale bars = 2 mm). 3. Ukamenia thailandica n. sp. (holotype) 4. Gatesclarkeana idia." figureDoi="http://doi.org/10.5281/zenodo.4425821" httpUri="https://zenodo.org/record/4425821/files/figure.png" pageId="3" pageNumber="86">Fig. 3</figureCitation>
) subtriangular, light brown, anal margin with fringe scales curve upwardly, underside light brown, slightly paler to apex, lower surface covered with diffuse curved up narrow scales between Sc+R
<subScript id="17EE9A2B6F11FFC96E4090799E3A8B20" attach="left" box="[1283,1291,529,543]" fontSize="6" pageId="3" pageNumber="86">1</subScript>
to 3A (
<figureCitation id="135184EB6F11FFC96E2793979BF48B01" captionStart="FIGURES 57" captionStartId="3.[151,264,1581,1605]" captionTargetBox="[155,1431,597,1558]" captionTargetId="figure-226@3.[155,1431,597,1558]" captionTargetPageId="3" captionText="FIGURES 57. Head and hindwing of Ukamenia thailandica n. sp. (holotype). 5. Composition of head. 6.Underside of hindwing. 7. Curved up scales on the underside of hindwing." figureDoi="http://doi.org/10.5281/zenodo.4425823" httpUri="https://zenodo.org/record/4425823/files/figure.png" pageId="3" pageNumber="86">Figs. 67</figureCitation>
).
</paragraph>
<caption id="DF15C8E66F11FFC96BD4944599E58F56" ID-DOI="http://doi.org/10.5281/zenodo.4425823" ID-Zenodo-Dep="4425823" httpUri="https://zenodo.org/record/4425823/files/figure.png" pageId="3" pageNumber="86" startId="3.[151,264,1581,1605]" targetBox="[155,1431,597,1558]" targetPageId="3">
<paragraph id="8BD5986E6F11FFC96BD4944599E58F56" blockId="3.[151,1436,1581,1641]" pageId="3" pageNumber="86">
<emphasis id="B91E447C6F11FFC96BD494459A058F7A" bold="true" box="[151,308,1581,1605]" pageId="3" pageNumber="86">FIGURES 57</emphasis>
. Head and hindwing of
<taxonomicName id="4C6AE3ED6F11FFC9696F9445983F8F7A" authority="Muadsub &amp; Pinkaew, 2020" authorityName="Muadsub &amp; Pinkaew" authorityYear="2020" box="[556,782,1581,1605]" class="Insecta" family="Tortricidae" genus="Ukamenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B91E447C6F11FFC9696F9445983F8F7A" box="[556,782,1581,1605]" italics="true" pageId="3" pageNumber="86">Ukamenia thailandica</emphasis>
</taxonomicName>
<emphasis id="B91E447C6F11FFC968569445987E8F7A" bold="true" box="[789,847,1581,1605]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A22DF9076F11FFC968569445987E8F7A" box="[789,847,1581,1605]" pageId="3" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype). 5. Composition of head. 6.Underside of hindwing. 7. Curved up scales on the underside of hindwing.
</paragraph>
</caption>
<paragraph id="8BD5986E6F11FFCE6B8494FF9BE58892" blockId="3.[151,1437,1687,2038]" lastBlockId="4.[151,1437,151,465]" lastPageId="4" lastPageNumber="87" pageId="3" pageNumber="86">
Abdomen. Tergum II with two small, crescentic pits on anterolateral corners (
<figureCitation id="135184EB6F11FFC96F6994FF9F4D8F8E" box="[1066,1148,1687,1713]" captionStart="FIGURES 1618" captionStartId="6.[151,264,1277,1301]" captionTargetBox="[255,1338,181,1248]" captionTargetId="figure-27@6.[250,1338,181,1253]" captionTargetPageId="6" captionText="FIGURES 1618. Distinct characters on the abdomen of Ukamenia thailandica n. sp. (holotype). 16. Overall image of abdomen. 17. Small pits on tergum II. 18. Subcylindrical pouches on sternum III." figureDoi="http://doi.org/10.5281/zenodo.4425831" httpUri="https://zenodo.org/record/4425831/files/figure.png" pageId="3" pageNumber="86">Fig. 17</figureCitation>
), sternum III anteromedially with a pair of large, subelliptic pouch contain dense scale sockets (
<figureCitation id="135184EB6F11FFC968FF94D39F218FEA" box="[956,1040,1723,1749]" captionStart="FIGURES 1618" captionStartId="6.[151,264,1277,1301]" captionTargetBox="[255,1338,181,1248]" captionTargetId="figure-27@6.[250,1338,181,1253]" captionTargetPageId="6" captionText="FIGURES 1618. Distinct characters on the abdomen of Ukamenia thailandica n. sp. (holotype). 16. Overall image of abdomen. 17. Small pits on tergum II. 18. Subcylindrical pouches on sternum III." figureDoi="http://doi.org/10.5281/zenodo.4425831" httpUri="https://zenodo.org/record/4425831/files/figure.png" pageId="3" pageNumber="86">Fig. 18</figureCitation>
), sternum VIII enlarged, forming a large subtriangular lobe, posterior margin with two small, pointed subtriangular lobes, tergum VIII enlarged, forming a large subtriangular lobe, dome-shaped (
<figureCitation id="135184EB6F11FFC969C9956B99ED8E21" box="[650,732,1795,1822]" captionStart="FIGURES 1618" captionStartId="6.[151,264,1277,1301]" captionTargetBox="[255,1338,181,1248]" captionTargetId="figure-27@6.[250,1338,181,1253]" captionTargetPageId="6" captionText="FIGURES 1618. Distinct characters on the abdomen of Ukamenia thailandica n. sp. (holotype). 16. Overall image of abdomen. 17. Small pits on tergum II. 18. Subcylindrical pouches on sternum III." figureDoi="http://doi.org/10.5281/zenodo.4425831" httpUri="https://zenodo.org/record/4425831/files/figure.png" pageId="3" pageNumber="86">Fig. 16</figureCitation>
). Male genitalia (
<figureCitation id="135184EB6F11FFC968DF956B9F138E21" box="[924,1058,1795,1822]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="3" pageNumber="86">Figs. 1115</figureCitation>
). Uncus absent; tegumen widened mid length, divided into two parts, upper half tapered to two rounded bilobed apex, lower half narrow to base, apical 1/4 of tegumen with roughly surface, with moderately dense setae; socii small, triangular, densely setose, arising at apical 1/4 of tegumen, pointing inwardly (
<figureCitation id="135184EB6F11FFC9693B950799FC8EB6" box="[632,717,1903,1929]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="3" pageNumber="86">Fig. 12</figureCitation>
); gnathos large, forming subpentagon, weakly sclerotized, arising from mid length of tegumen, lateroapical margins with two sclerotized arms, slant inwardly, becoming closer near blunted apices that bent outwardly, with sclerotized plus mark medially (
<figureCitation id="135184EB6F11FFC968BC95DF9F638EEE" box="[1023,1106,1975,2001]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="3" pageNumber="86">Fig. 12</figureCitation>
); vinculum large, moderately sclerotized; juxta subtriangular; caulis moderately long; phallus moderately long, cone-shaped, slightly curve medi- ally, basal half widened, abruptly narrow to mid length of phallus, then slightly tapered to pointed apex; valva large, costa process forming large, bean-shaped, protruding inwardly, with sparsely minute spines ventrally (
<figureCitation id="135184EB6F16FFCE6E7B92D39EBC89EA" box="[1336,1421,187,213]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="4" pageNumber="87">Fig. 13</figureCitation>
); sacculus rather small, subquadrate, near apicoventral corner with moderately long, subcylindrical projecting lobe, pointed apex, pointing outwardly, sparsely setose (
<figureCitation id="135184EB6F16FFCE6984936B98288822" box="[711,793,258,285]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="4" pageNumber="87">Fig. 15</figureCitation>
); cucullus large, elongate subelliptic, upper margin folded longitudinally, upper part with moderately dense, short mixed with long setae, ventral part sclerotized, smooth, apex of cucullus with a tuft of dense, long setae and one large strong spine, beneath with a row of strongly spines, varies in number (9 spines in
<typeStatus id="54D126CC6F16FFCE6A879307991788B6" box="[452,550,367,393]" pageId="4" pageNumber="87" type="holotype">holotype</typeStatus>
(
<figureCitation id="135184EB6F16FFCE6975930799B988B6" box="[566,648,367,393]" captionStart="FIGURES 1115" captionStartId="5.[151,264,1940,1964]" captionTargetBox="[220,1367,181,1916]" captionTargetId="figure-17@5.[220,1367,181,1916]" captionTargetPageId="5" captionText="FIGURES 1115. Male genitalia of Ukamenia thailandica n. sp. (holotype). 11. Male genital capsule with ex-situ aedeagus. 12. Tegumen, socii, and gnathos. 13. Costal process (right side). 14. Tuft of strongly setose with one strongly spine and a row of strongly 9 spines on terminal of left cucullus. 15. Subcylindrical projecting lobe on right sacculus." figureDoi="http://doi.org/10.5281/zenodo.4425829" httpUri="https://zenodo.org/record/4425829/files/figure.png" pageId="4" pageNumber="87">Fig. 14</figureCitation>
) and 10 spines in
<typeStatus id="54D126CC6F16FFCE681B9307988C88B6" box="[856,957,367,393]" pageId="4" pageNumber="87" type="paratype">paratype</typeStatus>
), length of these spines reduced orderly to apex.
</paragraph>
</subSubSection>
<subSubSection id="C370CBE56F16FFCE6B8493DF984088EE" box="[199,881,439,465]" pageId="4" pageNumber="87" type="etymology">
<paragraph id="8BD5986E6F16FFCE6B8493DF984088EE" blockId="4.[151,1437,151,465]" box="[199,881,439,465]" pageId="4" pageNumber="87">
<emphasis id="B91E447C6F16FFCE6B8493DF9A7988EE" bold="true" box="[199,328,439,465]" pageId="4" pageNumber="87">Etymology</emphasis>
. This species is named to
<collectingCountry id="F37DD8FE6F16FFCE692D93DF99E588EE" box="[622,724,439,465]" name="Thailand" pageId="4" pageNumber="87">Thailand</collectingCountry>
,
<typeStatus id="54D126CC6F16FFCE69A393DF982188EE" box="[736,784,439,465]" pageId="4" pageNumber="87">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>