446 lines
132 KiB
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446 lines
132 KiB
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<mods:title>The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence</mods:title>
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<emphasis bold="true" box="[882,1138,1132,1156]" pageId="8" pageNumber="1086">CERASOMMATIDIIDAE</emphasis>
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<emphasis bold="true" pageId="8" pageNumber="1086">STAT. RESTORED</emphasis>
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(
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<figureCitation box="[1077,1193,1202,1227]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-10="Figure 11" captionStart-11="Figure 12" captionStart-12="Figure 13" captionStart-13="Figure 14" captionStart-14="Figure 15" captionStart-15="Figure 16" captionStart-16="Figure 17" captionStart-17="Figure 18" captionStart-18="Figure 19" captionStart-19="Figure 20" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="5.[145,226,1413,1435]" captionStartId-1="6.[163,243,1744,1766]" captionStartId-10="20.[164,243,1731,1753]" captionStartId-11="24.[165,243,1357,1379]" captionStartId-12="25.[145,223,937,959]" captionStartId-13="26.[166,246,1040,1062]" captionStartId-14="27.[146,224,1623,1645]" captionStartId-15="28.[163,244,1465,1487]" captionStartId-16="31.[145,225,1505,1527]" captionStartId-17="32.[163,242,1451,1473]" captionStartId-18="33.[146,224,1461,1483]" captionStartId-19="34.[162,240,1081,1103]" captionStartId-2="10.[163,245,1463,1485]" captionStartId-3="11.[146,225,1546,1568]" captionStartId-4="12.[163,243,1601,1623]" captionStartId-5="15.[145,223,1742,1764]" captionStartId-6="16.[164,242,1280,1302]" captionStartId-7="17.[146,226,1180,1202]" captionStartId-8="18.[164,244,1685,1707]" captionStartId-9="19.[145,224,1601,1623]" captionTargetBox-0="[148,1422,212,1353]" captionTargetBox-1="[246,1361,198,1697]" captionTargetBox-10="[244,1364,194,1692]" captionTargetBox-11="[326,1284,202,1297]" captionTargetBox-12="[261,1302,205,895]" captionTargetBox-13="[329,1281,201,987]" captionTargetBox-14="[225,1345,196,1579]" captionTargetBox-15="[243,1363,195,1426]" captionTargetBox-16="[231,1339,197,1457]" captionTargetBox-17="[227,1379,194,1412]" captionTargetBox-18="[145,1425,195,1421]" captionTargetBox-19="[162,1442,200,1042]" captionTargetBox-2="[328,1277,202,1416]" captionTargetBox-3="[145,1425,195,1506]" captionTargetBox-4="[163,1443,196,1562]" captionTargetBox-5="[321,1262,200,1690]" captionTargetBox-6="[323,1279,211,1237]" captionTargetBox-7="[325,1255,199,1114]" captionTargetBox-8="[164,1444,196,1646]" captionTargetBox-9="[145,1425,195,1561]" captionTargetId-0="figure-168@5.[145,1425,194,1373]" captionTargetId-1="figure-25@6.[242,1364,194,1704]" captionTargetId-10="figure-26@20.[244,1364,194,1692]" captionTargetId-11="figure-271@24.[320,1287,196,1317]" captionTargetId-12="figure-455@25.[254,1316,198,898]" captionTargetId-13="figure-481@26.[323,1289,197,1001]" captionTargetId-14="figure-96@27.[225,1345,196,1584]" captionTargetId-15="figure-180@28.[243,1363,195,1426]" captionTargetId-16="figure-172@31.[222,1348,194,1466]" captionTargetId-17="figure-157@32.[227,1379,194,1412]" captionTargetId-18="figure-185@33.[145,1425,195,1421]" captionTargetId-19="figure-327@34.[162,1442,200,1042]" captionTargetId-2="figure-192@10.[319,1286,195,1423]" captionTargetId-3="figure-143@11.[145,1425,195,1506]" captionTargetId-4="figure-124@12.[163,1443,196,1562]" captionTargetId-5="figure-73@15.[302,1268,197,1702]" captionTargetId-6="figure-333@16.[319,1285,197,1240]" captionTargetId-7="figure-337@17.[302,1268,195,1140]" captionTargetId-8="figure-85@18.[164,1444,196,1646]" captionTargetId-9="figure-122@19.[145,1425,195,1561]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="20" captionTargetPageId-11="24" captionTargetPageId-12="25" captionTargetPageId-13="26" captionTargetPageId-14="27" captionTargetPageId-15="28" captionTargetPageId-16="31" captionTargetPageId-17="32" captionTargetPageId-18="33" captionTargetPageId-19="34" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="15" captionTargetPageId-6="16" captionTargetPageId-7="17" captionTargetPageId-8="18" captionTargetPageId-9="19" captionText-0="Figure 1. Results of the phylogenetic analysis of molecular data under Bayesian inference showing the position of Cerasommatidiidae status restored within Coccinelloidea. A summary tree of our molecular phylogenetic analyses using 236 taxa. Clades for each family/subfamily are collapsed. Statistical support for branches show posterior probabilities. The full-resolution tree is provided as in Supporting Information, File S11. Habitus pictures of the Cerasommatidiidae: MahaƲelo madagasus sp. nov. (lateral) and Yamuy marginatus sp. nov. (dorsal)." captionText-1="Figure 2. Results of the phylogenetic analyses of the combined, morphological and molecular dataset under Bayesian inference. Statistical support for branches show posterior probabilities. Branch lengths are kept and shown in the same proportion all along the tree. Apomorphies tracked in WINCLADA are shown upon each branch. DNA helix graphic after taxon name indicates taxa for which molecular data were available; asterisk ‘*’ indicates interspecific chimeras (combined data from two different species of the same genus)." captionText-10="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." captionText-11="Figure 12. Yamuy gen. nov. A, B, Yamuy constratus sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Yamuy marginatus sp. nov.: C, habitus dorsal. D, habitus lateral. Scale bars: 1mm." captionText-12="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." captionText-13="Figure 14. Terminalia of Yamuy gen. nov. A–C, Yamuy constratus sp. nov.: A, abdominal segment VIII and male genital segment, ventral. B, aedeagus, ventral. C, female genitalia. D–F, Yamuy marginatus sp. nov.: D, male genital segment, ventral. E, aedeagus, ventral. F, female genitalia." captionText-14="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." captionText-15="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." captionText-16="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." captionText-17="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." captionText-18="Figure 19. Habitats where type specimens of Cerasommatidiidae spp. were collected. A, track near the point where Yamuy constratus sp. nov. female paratype was collected. B, flight intercept trap in El Yunque National Forest where Y. constratus sp. nov. female paratype was collected. C, road-side habitat near Ifotaka, the type locality of MahaƲelo madagasus sp. nov. D, campsite at Forêt de Mahavelo, near the point where M. madagasus sp. nov. type series was collected." captionText-19="Figure 20. Geographical distribution of Cerasommatidiidae spp. A, distribution of species in eastern Caribbean region. B, distribution of species in southeastern Brazil. C, distribution of species in Madagascar." captionText-2="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." captionText-3="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." captionText-4="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." captionText-5="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." captionText-6="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." captionText-7="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." captionText-8="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." captionText-9="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi-0="http://doi.org/10.5281/zenodo.7814129" figureDoi-1="http://doi.org/10.5281/zenodo.7814131" figureDoi-10="http://doi.org/10.5281/zenodo.7814155" figureDoi-11="http://doi.org/10.5281/zenodo.7814161" figureDoi-12="http://doi.org/10.5281/zenodo.7814165" figureDoi-13="http://doi.org/10.5281/zenodo.7814169" figureDoi-14="http://doi.org/10.5281/zenodo.7814171" figureDoi-15="http://doi.org/10.5281/zenodo.7814175" figureDoi-16="http://doi.org/10.5281/zenodo.7814177" figureDoi-17="http://doi.org/10.5281/zenodo.7814181" figureDoi-18="http://doi.org/10.5281/zenodo.7814183" figureDoi-19="http://doi.org/10.5281/zenodo.7814187" figureDoi-2="http://doi.org/10.5281/zenodo.7814133" figureDoi-3="http://doi.org/10.5281/zenodo.7814135" figureDoi-4="http://doi.org/10.5281/zenodo.7814139" figureDoi-5="http://doi.org/10.5281/zenodo.7814141" figureDoi-6="http://doi.org/10.5281/zenodo.7814143" figureDoi-7="http://doi.org/10.5281/zenodo.7814145" figureDoi-8="http://doi.org/10.5281/zenodo.7814149" figureDoi-9="http://doi.org/10.5281/zenodo.7814151" httpUri-0="https://zenodo.org/record/7814129/files/figure.png" httpUri-1="https://zenodo.org/record/7814131/files/figure.png" httpUri-10="https://zenodo.org/record/7814155/files/figure.png" httpUri-11="https://zenodo.org/record/7814161/files/figure.png" httpUri-12="https://zenodo.org/record/7814165/files/figure.png" httpUri-13="https://zenodo.org/record/7814169/files/figure.png" httpUri-14="https://zenodo.org/record/7814171/files/figure.png" httpUri-15="https://zenodo.org/record/7814175/files/figure.png" httpUri-16="https://zenodo.org/record/7814177/files/figure.png" httpUri-17="https://zenodo.org/record/7814181/files/figure.png" httpUri-18="https://zenodo.org/record/7814183/files/figure.png" httpUri-19="https://zenodo.org/record/7814187/files/figure.png" httpUri-2="https://zenodo.org/record/7814133/files/figure.png" httpUri-3="https://zenodo.org/record/7814135/files/figure.png" httpUri-4="https://zenodo.org/record/7814139/files/figure.png" httpUri-5="https://zenodo.org/record/7814141/files/figure.png" httpUri-6="https://zenodo.org/record/7814143/files/figure.png" httpUri-7="https://zenodo.org/record/7814145/files/figure.png" httpUri-8="https://zenodo.org/record/7814149/files/figure.png" httpUri-9="https://zenodo.org/record/7814151/files/figure.png" pageId="8" pageNumber="1086">FIGS 1–20</figureCitation>
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)
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Cerasommatidiidae
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|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="1086" type="type_taxon">
|
||
<paragraph blockId="8.[827,1443,1242,1418]" pageId="8" pageNumber="1086">
|
||
<typeStatus box="[1301,1357,1243,1264]" pageId="8" pageNumber="1086">Type</typeStatus>
|
||
genus:
|
||
<taxonomicName authority="Brethes, 1925" authorityName="Brethes" authorityYear="1925" box="[859,1261,1273,1295]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1086" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[859,1047,1273,1294]" italics="true" pageId="8" pageNumber="1086">Cerasommatidia</emphasis>
|
||
<bibRefCitation author="Brethes J" box="[1080,1261,1273,1295]" pageId="8" pageNumber="1086" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural " Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">Brèthes, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
by original designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="1086" type="reference_group">
|
||
<paragraph blockId="8.[827,1443,1242,1418]" pageId="8" pageNumber="1086">
|
||
As synonym of
|
||
<taxonomicName authority="Casey, 1895: 452" authorityName="Casey" authorityPageNumber="452" authorityYear="1895" box="[1036,1424,1334,1356]" class="Bacillariophyceae" family="Plagiogrammaceae" genus="Eupsilobiinae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Triceratiales" pageId="8" pageNumber="1086" phylum="Ochrophyta" rank="genus">
|
||
Eupsilobiinae
|
||
<bibRefCitation author="Casey TL" box="[1211,1424,1334,1356]" pageId="8" pageNumber="1086" pagination="435 - 838" refId="ref22235" refString="Casey TL. 1895. Coleopterological notices VI. Annals of the Neae York Academy of Science 8: 435 - 838." type="journal article" year="1895">Casey, 1895: 452</bibRefCitation>
|
||
</taxonomicName>
|
||
.–
|
||
<bibRefCitation author="Pakaluk J & Slipinski SA & Lawrence JF" box="[859,1146,1365,1387]" pageId="8" pageNumber="1086" pagination="223 - 268" refId="ref22691" refString="Pakaluk J, Slipinski SA, Lawrence JF. 1994. Current classification and family-group names in Cucujoidea [Coleoptera]. Genus. International Journal of InVertebrate Taxonomy 5: 223 - 268." type="journal article" year="1994">
|
||
Pakaluk
|
||
<emphasis box="[962,1019,1365,1387]" italics="true" pageId="8" pageNumber="1086">et al.</emphasis>
|
||
, 1994: 228
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Shockley FW & Tomaszewska KW & McHugh JV" box="[1159,1439,1365,1387]" pageId="8" pageNumber="1086" pagination="1 - 113" refId="ref23149" refString="Shockley FW, Tomaszewska KW, McHugh JV. 2009. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113." type="journal article" year="2009">
|
||
Shockley
|
||
<emphasis box="[1268,1326,1365,1387]" italics="true" pageId="8" pageNumber="1086">et al.</emphasis>
|
||
, 2009: 27
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Bouchard P & Bousquet Y & Davies AE & Alonso-Zarazaga MA & Lawrence JF & Lyal CH & Newton AF & Reid CA & Schmitt M & Slipinski SA & Smith AB" box="[859,1153,1396,1418]" pageId="8" pageNumber="1086" pagination="1 - 972" refId="ref22151" refString="Bouchard P, Bousquet Y, Davies AE, Alonso-Zarazaga MA, Lawrence JF, Lyal CH, Newton AF, Reid CA, Schmitt M, Slipinski SA, Smith AB. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1 - 972." type="journal article" year="2011">
|
||
Bouchard
|
||
<emphasis box="[974,1022,1396,1417]" italics="true" pageId="8" pageNumber="1086">et al</emphasis>
|
||
., 2011: 372
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="9" lastPageNumber="1087" pageId="8" pageNumber="1086" type="diagnosis">
|
||
<paragraph blockId="8.[827,1444,1480,1900]" lastBlockId="9.[145,762,197,1905]" lastPageId="9" lastPageNumber="1087" pageId="8" pageNumber="1086">
|
||
<emphasis box="[827,1132,1480,1501]" italics="true" pageId="8" pageNumber="1086">Diagnostic combination:</emphasis>
|
||
Body short oval (
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.45" metricValueMin="0.8" pageId="8" pageNumber="1086" unit="mm" value="1.125" valueMax="1.45" valueMin="0.8">0.80– 1.45 mm</quantity>
|
||
) and convex (
|
||
<figureCitation box="[1101,1175,1510,1532]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="8" pageNumber="1086">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation box="[1190,1204,1510,1532]" captionStart="Figure 6" captionStartId="15.[145,223,1742,1764]" captionTargetBox="[321,1262,200,1690]" captionTargetId="figure-73@15.[302,1268,197,1702]" captionTargetPageId="15" captionText="Figure 6. Karumbe gen. nov. A, B, Karumbe brethesi sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Karumbe geiseri sp. nov.: C, habitus dorsal. D, habitus lateral. E, F, Karumbe pakaluki sp. nov.: E, habitus dorsal. F, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814141" httpUri="https://zenodo.org/record/7814141/files/figure.png" pageId="8" pageNumber="1086">6</figureCitation>
|
||
,
|
||
<figureCitation box="[1220,1248,1511,1532]" captionStart="Figure 12" captionStartId="24.[165,243,1357,1379]" captionTargetBox="[326,1284,202,1297]" captionTargetId="figure-271@24.[320,1287,196,1317]" captionTargetPageId="24" captionText="Figure 12. Yamuy gen. nov. A, B, Yamuy constratus sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Yamuy marginatus sp. nov.: C, habitus dorsal. D, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814161" httpUri="https://zenodo.org/record/7814161/files/figure.png" pageId="8" pageNumber="1086">12</figureCitation>
|
||
,
|
||
<figureCitation box="[1264,1292,1511,1532]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="8" pageNumber="1086">17</figureCitation>
|
||
). Head with frontoclypeal suture present (
|
||
<figureCitation box="[1165,1253,1541,1563]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="8" pageNumber="1086">Figs 9A</figureCitation>
|
||
,
|
||
<figureCitation box="[1264,1309,1541,1563]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="8" pageNumber="1086">15B</figureCitation>
|
||
,
|
||
<figureCitation box="[1321,1367,1541,1562]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="8" pageNumber="1086">18B</figureCitation>
|
||
). Eyes coarsely faceted (
|
||
<figureCitation box="[1026,1114,1572,1594]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="8" pageNumber="1086">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation box="[1128,1160,1572,1594]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="8" pageNumber="1086">5A</figureCitation>
|
||
,
|
||
<figureCitation box="[1173,1205,1572,1593]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="8" pageNumber="1086">7A</figureCitation>
|
||
,
|
||
<figureCitation box="[1219,1251,1572,1593]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="8" pageNumber="1086">9A</figureCitation>
|
||
,
|
||
<figureCitation box="[1264,1308,1572,1594]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="8" pageNumber="1086">15B</figureCitation>
|
||
,
|
||
<figureCitation box="[1322,1367,1572,1594]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="8" pageNumber="1086">16A</figureCitation>
|
||
,
|
||
<figureCitation box="[1381,1428,1572,1593]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="8" pageNumber="1086">17D</figureCitation>
|
||
). Antennal sockets visible from above (
|
||
<figureCitation box="[1258,1346,1602,1624]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="8" pageNumber="1086">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation box="[1360,1391,1602,1624]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="8" pageNumber="1086">5A</figureCitation>
|
||
,
|
||
<figureCitation box="[1405,1437,1603,1624]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="8" pageNumber="1086">9A</figureCitation>
|
||
,
|
||
<figureCitation box="[827,872,1633,1655]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="8" pageNumber="1086">16A</figureCitation>
|
||
). Ventral antennal grooves deep between eyes and mouthparts (
|
||
<figureCitation box="[975,1059,1664,1686]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="8" pageNumber="1086">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation box="[1070,1115,1664,1685]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="8" pageNumber="1086">18B</figureCitation>
|
||
). Gular sutures short, widely separated, weakly convergent anteriorly (
|
||
<figureCitation box="[1342,1436,1694,1716]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="8" pageNumber="1086">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation box="[827,872,1725,1747]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="8" pageNumber="1086">15A</figureCitation>
|
||
). Tentorium with anterior arms fused medially and widely divergent anteriorly, posterior arms connected throughout their length to posterior weakly sinuate transverse bridge/corpotentorium (
|
||
<figureCitation box="[1233,1321,1817,1839]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="8" pageNumber="1086">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation box="[1335,1367,1817,1838]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="8" pageNumber="1086">7A</figureCitation>
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,
|
||
<figureCitation box="[1381,1428,1817,1838]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="8" pageNumber="1086">17D</figureCitation>
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||
). Antenna composed of 11 antennomeres with distinct two-antennomere club (
|
||
<figureCitation box="[1095,1181,1878,1900]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="8" pageNumber="1086">Figs 3C</figureCitation>
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||
,
|
||
<figureCitation box="[1192,1224,1878,1900]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="8" pageNumber="1086">5E</figureCitation>
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,
|
||
<figureCitation box="[1235,1267,1879,1900]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="8" pageNumber="1086">7A</figureCitation>
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,
|
||
<figureCitation box="[1278,1323,1879,1900]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="8" pageNumber="1086">11A</figureCitation>
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,
|
||
<figureCitation box="[1335,1379,1879,1900]" captionStart="Figure 12" captionStartId="24.[165,243,1357,1379]" captionTargetBox="[326,1284,202,1297]" captionTargetId="figure-271@24.[320,1287,196,1317]" captionTargetPageId="24" captionText="Figure 12. Yamuy gen. nov. A, B, Yamuy constratus sp. nov.: A, habitus dorsal. B, habitus lateral. C, D, Yamuy marginatus sp. nov.: C, habitus dorsal. D, habitus lateral. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.7814161" httpUri="https://zenodo.org/record/7814161/files/figure.png" pageId="8" pageNumber="1086">12C</figureCitation>
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,
|
||
<figureCitation box="[1391,1437,1879,1900]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="8" pageNumber="1086">13A</figureCitation>
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||
,
|
||
<figureCitation box="[145,191,197,219]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15A</figureCitation>
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,
|
||
<figureCitation box="[208,254,197,218]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18A</figureCitation>
|
||
). Labrum large, sparsely setose, rounded anteriorly with or without small, median emargination (
|
||
<figureCitation box="[153,238,258,280]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">Figs 7B</figureCitation>
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||
,
|
||
<figureCitation box="[249,294,259,280]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10A</figureCitation>
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||
,
|
||
<figureCitation box="[305,350,259,280]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13B</figureCitation>
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||
,
|
||
<figureCitation box="[361,405,258,280]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15B</figureCitation>
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,
|
||
<figureCitation box="[416,462,259,280]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18B</figureCitation>
|
||
). Maxillary palpi (
|
||
<figureCitation box="[670,755,258,280]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation box="[145,175,289,311]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5B</figureCitation>
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||
,
|
||
<figureCitation box="[188,219,289,310]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7C</figureCitation>
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||
,
|
||
<figureCitation box="[232,263,289,310]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9B</figureCitation>
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||
,
|
||
<figureCitation box="[275,321,289,310]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10A</figureCitation>
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||
,
|
||
<figureCitation box="[334,378,289,310]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13C</figureCitation>
|
||
,
|
||
<figureCitation box="[391,437,289,311]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15A</figureCitation>
|
||
,
|
||
<figureCitation box="[449,495,289,310]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17E</figureCitation>
|
||
) with four palpomeres: palpomere 1 small; palpomere 2 weakly transverse and somewhat bulbous; palpomere 3 transverse; terminal palpomere distinctly elongate, swollen near base and then narrowing/flattened laterally toward apex; apex with short, sclerotized sensilla. Mandible stout (
|
||
<figureCitation box="[219,307,473,495]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">Figs 7E</figureCitation>
|
||
,
|
||
<figureCitation box="[321,367,473,494]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13E</figureCitation>
|
||
,
|
||
<figureCitation box="[380,425,473,494]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17F</figureCitation>
|
||
), sclerotized, with two apical teeth and two small subapical teeth; prostheca large, membranous with fringe of stout setae along its inner margin and with a series of long sclerotized, teeth-like projections apically; mola large, strongly sclerotized, with distinct transverse ridges. Labium with mentum irregularly hexagonal, with anterior margin straight, surface flat; prementum weakly elongate, sclerotized with ligula submembranous, lobed at sides (
|
||
<figureCitation box="[155,257,749,771]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">Figs 5B</figureCitation>
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,
|
||
<figureCitation box="[278,312,750,771]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7D</figureCitation>
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,
|
||
<figureCitation box="[334,367,749,770]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9B</figureCitation>
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,
|
||
<figureCitation box="[388,437,749,770]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10A</figureCitation>
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,
|
||
<figureCitation box="[459,508,749,771]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15A</figureCitation>
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||
). Palpi with three palpomeres, widely separated by a distance more than width of palpiger: palpomere 1 smallest, palpomere 2 bulbous, palpomere 3 sometimes slightly bulbous.
|
||
</paragraph>
|
||
<paragraph blockId="9.[145,762,197,1905]" pageId="9" pageNumber="1087">
|
||
Pronotum strongly transverse, widest at or near base and strongly convergent anteriorly (
|
||
<figureCitation box="[536,619,902,924]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3A</figureCitation>
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||
,
|
||
<figureCitation box="[630,659,902,923]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">4B</figureCitation>
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||
,
|
||
<figureCitation box="[671,701,902,924]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5C</figureCitation>
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,
|
||
<figureCitation box="[712,755,902,923]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10C</figureCitation>
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,
|
||
<figureCitation box="[145,190,933,954]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="9" pageNumber="1087">11B</figureCitation>
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,
|
||
<figureCitation box="[205,251,933,954]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18C</figureCitation>
|
||
). Front angles weakly produced anteriorly, obtuse; hind angles often with small, oblique indentation to receive humeral corner of elytron. Prosternum large, in form of chin piece covering mouthparts (
|
||
<figureCitation box="[588,673,1025,1047]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">Figs 5D</figureCitation>
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||
,
|
||
<figureCitation box="[685,712,1025,1046]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7F</figureCitation>
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||
,
|
||
<figureCitation box="[724,755,1025,1046]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9D</figureCitation>
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||
,
|
||
<figureCitation box="[145,188,1056,1077]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10B</figureCitation>
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||
,
|
||
<figureCitation box="[201,244,1056,1077]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="9" pageNumber="1087">11C</figureCitation>
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||
,
|
||
<figureCitation box="[257,302,1056,1078]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15D</figureCitation>
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,
|
||
<figureCitation box="[314,358,1056,1078]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16C</figureCitation>
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,
|
||
<figureCitation box="[370,416,1056,1077]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18D</figureCitation>
|
||
); prosternal process narrowest in about its half-length, continuously widened apically, provided with raised, paired lateral carinae or single median carina. Hypomeron with deep and long antennal grooves (
|
||
<figureCitation box="[254,344,1178,1200]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">Figs 7F</figureCitation>
|
||
,
|
||
<figureCitation box="[360,393,1178,1199]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9D</figureCitation>
|
||
,
|
||
<figureCitation box="[410,457,1178,1199]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10B</figureCitation>
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,
|
||
<figureCitation box="[473,520,1178,1200]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15D</figureCitation>
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,
|
||
<figureCitation box="[537,585,1178,1199]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18D</figureCitation>
|
||
). Mesoventral intercoxal process widely separates mesocoxae. Mesocoxa circular in outline, its cavity outwardly open; trochantin concealed (
|
||
<figureCitation box="[261,343,1270,1292]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation box="[352,383,1270,1291]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">4E</figureCitation>
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,
|
||
<figureCitation box="[392,425,1270,1292]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5H</figureCitation>
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,
|
||
<figureCitation box="[434,457,1271,1292]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7I</figureCitation>
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,
|
||
<figureCitation box="[467,492,1270,1291]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9F</figureCitation>
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,
|
||
<figureCitation box="[502,546,1270,1291]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10E</figureCitation>
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,
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<figureCitation box="[555,594,1270,1291]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="9" pageNumber="1087">11F</figureCitation>
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,
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<figureCitation box="[604,648,1270,1291]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13D</figureCitation>
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,
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||
<figureCitation box="[657,701,1270,1292]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15G</figureCitation>
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,
|
||
<figureCitation box="[711,754,1270,1292]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16D</figureCitation>
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,
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||
<figureCitation box="[145,189,1301,1322]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17G</figureCitation>
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,
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||
<figureCitation box="[200,243,1301,1322]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18E</figureCitation>
|
||
). Meso-metaventral junction arcuate anteriorly. Metaventrite strongly transverse, weakly convex, without postcoxal lines. Metanepisternum with small, outer blunt projection near anterior margin (
|
||
<figureCitation box="[666,755,1393,1415]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3D</figureCitation>
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||
,
|
||
<figureCitation box="[145,168,1424,1445]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7I</figureCitation>
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,
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||
<figureCitation box="[181,227,1424,1445]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17G</figureCitation>
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||
). Metacoxae transverse, widely separated (
|
||
<figureCitation captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3D</figureCitation>
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,
|
||
<figureCitation box="[190,214,1455,1476]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7I</figureCitation>
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,
|
||
<figureCitation box="[227,254,1454,1475]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9F</figureCitation>
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,
|
||
<figureCitation box="[268,309,1454,1475]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10F</figureCitation>
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,
|
||
<figureCitation box="[322,367,1454,1476]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16D</figureCitation>
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,
|
||
<figureCitation box="[381,426,1454,1475]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18E</figureCitation>
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||
). Metendosternite with short stalk and widely separated anterior arms and anterior tendons (
|
||
<figureCitation box="[247,320,1516,1538]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">Fig. 7J</figureCitation>
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,
|
||
<figureCitation box="[333,378,1516,1537]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13D</figureCitation>
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||
). Epipleura not foveate. Legs with trochanterofemoral attachment subheteromeroid (
|
||
<figureCitation captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation box="[191,226,1577,1599]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5H</figureCitation>
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,
|
||
<figureCitation box="[240,273,1577,1598]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7K</figureCitation>
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,
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||
<figureCitation box="[287,314,1577,1598]" captionStart="Figure 9" captionStartId="18.[164,244,1685,1707]" captionTargetBox="[164,1444,196,1646]" captionTargetId="figure-85@18.[164,1444,196,1646]" captionTargetPageId="18" captionText="Figure 9. Morphology of Karumbe brethesi sp. nov. A, head, frontal. B, head, ventral. C, right half of pronotum (black arrow: crenulate anterior margin; white arrow: indentation at posterolateral angle of pronotum). D, prosternum. E, detail of elytron, dorsal (arrow: peculiar punctures). F, metaventrite (arrow: large punctures at the centre of metaventrite)." figureDoi="http://doi.org/10.5281/zenodo.7814149" httpUri="https://zenodo.org/record/7814149/files/figure.png" pageId="9" pageNumber="1087">9F</figureCitation>
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,
|
||
<figureCitation box="[329,370,1577,1598]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10F</figureCitation>
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,
|
||
<figureCitation box="[384,426,1577,1598]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="9" pageNumber="1087">11F</figureCitation>
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,
|
||
<figureCitation box="[440,486,1577,1598]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13D</figureCitation>
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,
|
||
<figureCitation box="[500,547,1577,1599]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15D</figureCitation>
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,
|
||
<figureCitation box="[560,606,1577,1599]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16D</figureCitation>
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||
,
|
||
<figureCitation box="[621,667,1577,1598]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18E</figureCitation>
|
||
). Tibiae slender, gradually weakly widening towards tarsus; surrounded by short, stout spines. Tarsal formula 4-4-
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="1087" unit="in" value="4.0">4 in</quantity>
|
||
both sexes with tarsomere 1 weakly and tarsomere 2 distinctly lobed ventrally, tarsomere 3 short and tarsomere 4 about as long as tarsomere 1 (
|
||
<figureCitation box="[632,714,1730,1752]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">Figs 4F</figureCitation>
|
||
,
|
||
<figureCitation box="[728,754,1730,1752]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5F</figureCitation>
|
||
,
|
||
<figureCitation box="[145,174,1761,1782]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7L</figureCitation>
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||
,
|
||
<figureCitation box="[186,230,1761,1782]" captionStart="Figure 10" captionStartId="19.[145,224,1601,1623]" captionTargetBox="[145,1425,195,1561]" captionTargetId="figure-122@19.[145,1425,195,1561]" captionTargetPageId="19" captionText="Figure 10. Morphology of Karumbe geiseri sp. nov. A, mouthparts, ventral. B, prosternum. C, pronotum (arrow: crenulate anterior margin). D, protarsus. E, meso- and metaventrite (arrow: fine punctures at the centre of metaventrite). F, metacoxa and metatrochanter, lateral, first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814151" httpUri="https://zenodo.org/record/7814151/files/figure.png" pageId="9" pageNumber="1087">10D</figureCitation>
|
||
,
|
||
<figureCitation box="[242,287,1761,1782]" captionStart="Figure 11" captionStartId="20.[164,243,1731,1753]" captionTargetBox="[244,1364,194,1692]" captionTargetId="figure-26@20.[244,1364,194,1692]" captionTargetPageId="20" captionText="Figure 11. Morphology of Karumbe pakaluki sp. nov. A, antenna, dorsal. B, pronotum [arrows: pronotal anterior margin (pr am), pronotal internal lateral carina (pr car), pronotal posterolateral angle (pr ang)]. C, prosternum. D, detail of elytron, dorsal [arrows: setiferous puncture (set pun), foveate puncture (fov pun)]. E, metatarsus, dorsal. F, meso- metaventrite and first abdominal ventrite [arrow: abdominal postcoxal line (abd lin)]." figureDoi="http://doi.org/10.5281/zenodo.7814155" httpUri="https://zenodo.org/record/7814155/files/figure.png" pageId="9" pageNumber="1087">11E</figureCitation>
|
||
,
|
||
<figureCitation box="[298,342,1761,1783]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15E</figureCitation>
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||
,
|
||
<figureCitation box="[354,399,1761,1782]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18G</figureCitation>
|
||
). Claws toothed at base (
|
||
<figureCitation box="[675,755,1761,1783]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">Figs 5F</figureCitation>
|
||
,
|
||
<figureCitation box="[145,183,1792,1813]" captionStart="Figure 13" captionStartId="25.[145,223,937,959]" captionTargetBox="[261,1302,205,895]" captionTargetId="figure-455@25.[254,1316,198,898]" captionTargetPageId="25" captionText="Figure 13. Morphology of Yamuy gen. nov.A, Yamuy marginatus sp. nov. B–F, Yamuy constratus sp. nov.: B, labrum, dorsal. C, maxilla, ventral. D, meso- and metaventrite. E, mandible, ventral. F, terminal metatarsomere and claws." figureDoi="http://doi.org/10.5281/zenodo.7814165" httpUri="https://zenodo.org/record/7814165/files/figure.png" pageId="9" pageNumber="1087">13F</figureCitation>
|
||
,
|
||
<figureCitation box="[196,239,1791,1813]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15E</figureCitation>
|
||
,
|
||
<figureCitation box="[251,295,1791,1813]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[145,762,197,1905]" lastBlockId="9.[809,1426,197,617]" pageId="9" pageNumber="1087">
|
||
Abdomen with six ventrites in both sexes (
|
||
<figureCitation box="[665,754,1822,1844]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">Figs 4G</figureCitation>
|
||
,
|
||
<figureCitation box="[145,182,1853,1874]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="9" pageNumber="1087">8M</figureCitation>
|
||
,
|
||
<figureCitation box="[196,244,1853,1875]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">15H</figureCitation>
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||
,
|
||
<figureCitation box="[258,305,1853,1874]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18G</figureCitation>
|
||
). Ventrite 1 with (
|
||
<figureCitation box="[518,607,1853,1875]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation box="[621,653,1853,1874]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">4G</figureCitation>
|
||
,
|
||
<figureCitation box="[667,702,1853,1875]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5H</figureCitation>
|
||
,
|
||
<figureCitation box="[715,752,1853,1874]" captionStart="Figure 7" captionStartId="16.[164,242,1280,1302]" captionTargetBox="[323,1279,211,1237]" captionTargetId="figure-333@16.[319,1285,197,1240]" captionTargetPageId="16" captionText="Figure 7. Morphology of Karumbe gen. nov. A–E, G, J–K, M, Karumbe pakaluki sp. nov. F, I, L, Karumbe geiseri sp. nov. H, Karumbe brethesi sp. nov. A, head without mouthparts, ventral. B, labrum, dorsal. C, maxilla, ventral. D, labium, ventral. E, mandible, ventral. F, prothorax, ventral, G, metanotum. H, hindwing. I, meso- and metaventrite. J, details of metaventrite showing metendosternite. K, hindleg, ventral. L, metatarsus. M, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814143" httpUri="https://zenodo.org/record/7814143/files/figure.png" pageId="9" pageNumber="1087">7M</figureCitation>
|
||
) or without distinctly developed, rounded postcoxal lines (
|
||
<figureCitation box="[878,982,197,219]" captionStart="Figure 15" captionStartId="27.[146,224,1623,1645]" captionTargetBox="[225,1345,196,1579]" captionTargetId="figure-96@27.[225,1345,196,1584]" captionTargetPageId="27" captionText="Figure 15. Morphology of Yamuy constratus sp. nov. A, head, ventral. B, head, dorso-frontal. C, left half of pronotum (arrow: reduced pronotal internal carina). D, prothorax, ventral (arrow: protrochanter). E, mesotarsus, ventral. F, left elytron, dorsal. G, meso- and metaventrite, ventral (arrow: mesotrochanter). H, abdomen, ventral." figureDoi="http://doi.org/10.5281/zenodo.7814171" httpUri="https://zenodo.org/record/7814171/files/figure.png" pageId="9" pageNumber="1087">Figs 15H</figureCitation>
|
||
,
|
||
<figureCitation box="[995,1036,197,219]" captionStart="Figure 16" captionStartId="28.[163,244,1465,1487]" captionTargetBox="[243,1363,195,1426]" captionTargetId="figure-180@28.[243,1363,195,1426]" captionTargetPageId="28" captionText="Figure 16. Morphology of Yamuy marginatus sp. nov. A, head, dorsal. B, pronotum. C, prosternum (arrow: narrowed prosternal process). D, meso- and metaventrite (arrow: short postcoxal longitudinal carinae). E, protarsus, ventral. F, abdomen, ventral (arrow: short postcoxal longitudinal carinae)." figureDoi="http://doi.org/10.5281/zenodo.7814175" httpUri="https://zenodo.org/record/7814175/files/figure.png" pageId="9" pageNumber="1087">16F</figureCitation>
|
||
,
|
||
<figureCitation box="[1049,1096,197,218]" captionStart="Figure 18" captionStartId="32.[163,242,1451,1473]" captionTargetBox="[227,1379,194,1412]" captionTargetId="figure-157@32.[227,1379,194,1412]" captionTargetPageId="32" captionText="Figure 18. Morphology of MahaƲelo madagasus sp. nov. A, antenna. B, head, dorso-frontal (arrow: antennal grooves). C, left half of pronotum (arrow: indentation on postero-lateral corner of pronotum). D, prosternum (arrow: median carina). E, pterothorax and first abdominal ventrite (white arrow: epipleural groove; black arrows: meso- and metatrochanters). F, left elytron, dorsal. G, left half of abdomen, ventral (arrow: anterior bordering carina)." figureDoi="http://doi.org/10.5281/zenodo.7814181" httpUri="https://zenodo.org/record/7814181/files/figure.png" pageId="9" pageNumber="1087">18G</figureCitation>
|
||
); at midline about as long as three next ventrites together. Ventrite 5 with posterolateral angles strongly produced backwards and somewhat overlapping antero-lateral parts of ventrite 6 (anterior margin of ventrite 6 much narrower than posterior margin of ventrite 5); ventrite 6 arcuate apically, surrounded by anterior margin of tergite VIII bent downwards, appearing as narrow, false ‘ventrite 7’. Male genital segment (IX) reduced but with strongly developed, sclerotized, long, narrow, single apophysis (
|
||
<figureCitation box="[817,907,503,525]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation box="[922,988,504,525]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="9" pageNumber="1087">8A, D</figureCitation>
|
||
,
|
||
<figureCitation box="[1002,1082,504,525]" captionStart="Figure 14" captionStartId="26.[166,246,1040,1062]" captionTargetBox="[329,1281,201,987]" captionTargetId="figure-481@26.[323,1289,197,1001]" captionTargetPageId="26" captionText="Figure 14. Terminalia of Yamuy gen. nov. A–C, Yamuy constratus sp. nov.: A, abdominal segment VIII and male genital segment, ventral. B, aedeagus, ventral. C, female genitalia. D–F, Yamuy marginatus sp. nov.: D, male genital segment, ventral. E, aedeagus, ventral. F, female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814169" httpUri="https://zenodo.org/record/7814169/files/figure.png" pageId="9" pageNumber="1087">14A, D</figureCitation>
|
||
,
|
||
<figureCitation box="[1096,1134,504,525]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17I</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation box="[1287,1373,503,525]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="9" pageNumber="1087">8B, E</figureCitation>
|
||
,
|
||
<figureCitation box="[842,921,534,556]" captionStart="Figure 14" captionStartId="26.[166,246,1040,1062]" captionTargetBox="[329,1281,201,987]" captionTargetId="figure-481@26.[323,1289,197,1001]" captionTargetPageId="26" captionText="Figure 14. Terminalia of Yamuy gen. nov. A–C, Yamuy constratus sp. nov.: A, abdominal segment VIII and male genital segment, ventral. B, aedeagus, ventral. C, female genitalia. D–F, Yamuy marginatus sp. nov.: D, male genital segment, ventral. E, aedeagus, ventral. F, female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814169" httpUri="https://zenodo.org/record/7814169/files/figure.png" pageId="9" pageNumber="1087">14B, E</figureCitation>
|
||
,
|
||
<figureCitation box="[936,985,534,555]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17H</figureCitation>
|
||
) with penis resting on its side when retracted.Ovipositor of female genitalia without coxites (
|
||
<figureCitation box="[817,935,595,617]" captionStart="Figure 8" captionStartId="17.[146,226,1180,1202]" captionTargetBox="[325,1255,199,1114]" captionTargetId="figure-337@17.[302,1268,195,1140]" captionTargetPageId="17" captionText="Figure 8. Terminalia of Karumbe gen. nov. A–C, Karumbe geiseri sp. nov.: A, male genital segment, ventro-lateral view. B, aedeagus, ventral. C, female genitalia. D–F, Karumbe pakaluki sp. nov.: D, male genital segment, ventro-lateral view. E, aedeagus, ventral. F, apex of bursa copulatrix and spermatheca. G, Karumbe brethesi sp. nov., female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814145" httpUri="https://zenodo.org/record/7814145/files/figure.png" pageId="9" pageNumber="1087">Figs 8C, G</figureCitation>
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,
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<figureCitation box="[948,1019,596,617]" captionStart="Figure 14" captionStartId="26.[166,246,1040,1062]" captionTargetBox="[329,1281,201,987]" captionTargetId="figure-481@26.[323,1289,197,1001]" captionTargetPageId="26" captionText="Figure 14. Terminalia of Yamuy gen. nov. A–C, Yamuy constratus sp. nov.: A, abdominal segment VIII and male genital segment, ventral. B, aedeagus, ventral. C, female genitalia. D–F, Yamuy marginatus sp. nov.: D, male genital segment, ventral. E, aedeagus, ventral. F, female genitalia." figureDoi="http://doi.org/10.5281/zenodo.7814169" httpUri="https://zenodo.org/record/7814169/files/figure.png" pageId="9" pageNumber="1087">14C, F</figureCitation>
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,
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<figureCitation box="[1033,1074,596,617]" captionStart="Figure 17" captionStartId="31.[145,225,1505,1527]" captionTargetBox="[231,1339,197,1457]" captionTargetId="figure-172@31.[222,1348,194,1466]" captionTargetPageId="31" captionText="Figure 17. Morphology of MahaƲelo madagasus sp. nov. A, habitus dorsal. B, habitus frontal. C, habitus lateral. Scale bars: 1 mm. D, head with mouthparts removed, ventral. E, maxilla, ventral. F, mandible, ventral. G, meso- and metathorax, ventral. H, aedeagus, ventral. I, male genital segment, ventral. J, female genitalia, ventral. K, details of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7814177" httpUri="https://zenodo.org/record/7814177/files/figure.png" pageId="9" pageNumber="1087">17J</figureCitation>
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).
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |