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<document ID-DOI="10.1093/zoolinnean/zlac082" ID-GBIF-Dataset="f791f8ed-ae19-41a1-b5cc-b9b2cc89dec4" ID-ISSN="0024-4082" ID-Zenodo-Dep="7814127" ID-ZooBank="99842C3A-879F-4552-96B7-204D79CF3EDF" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="31" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="13" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="6" checkinTime="1680597737982" checkinUser="plazi" docAuthor="Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam &amp; Fikáček, Martin" docDate="2023" docId="03C2CE00FFF0FF963D8509BBFE1E4633" docLanguage="en" docName="zlac082.pdf" docOrigin="Zoological Journal of the Linnean Society 197 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlac082" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Cerasommatidia Brethes 1925" docType="treatment" docVersion="3" lastPageNumber="1091" masterDocId="FFFBB678FFF9FF9B3E190B0AFFBD416E" masterDocTitle="The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence" masterLastPageNumber="1115" masterPageNumber="1078" pageNumber="1087" updateTime="1681148941709" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Arriaga-Varela, Emmanuel</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tomaszewska, Wioletta</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Szawaryn, Karol</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Robertson, James</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ślipiński, Adam</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fikáček, Martin</mods:namePart>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-24</mods:number>
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<mods:detail type="volume">
<mods:number>197</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1093/zoolinnean/zlac082</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="ZooBank">99842C3A-879F-4552-96B7-204D79CF3EDF</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7814159" ID-Zenodo-Dep="7814159" LSID="urn:lsid:plazi:treatment:03C2CE00FFF0FF963D8509BBFE1E4633" httpUri="http://treatment.plazi.org/id/03C2CE00FFF0FF963D8509BBFE1E4633" lastPageId="13" lastPageNumber="1091" pageId="9" pageNumber="1087">
<subSubSection box="[924,1309,689,714]" pageId="9" pageNumber="1087" type="nomenclature">
<paragraph blockId="9.[924,1309,689,714]" box="[924,1309,689,714]" pageId="9" pageNumber="1087">
<heading box="[924,1309,689,714]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="1087" reason="2">
<taxonomicName authority="BRETHES, 1925" authorityName="Brethes" authorityYear="1925" box="[924,1309,689,714]" class="Insecta" family="Cerasommatidiidae" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[924,1128,689,712]" italics="true" pageId="9" pageNumber="1087">CERASOMMATIDIA</emphasis>
<bibRefCitation author="Brethes J" box="[1135,1309,689,714]" pageId="9" pageNumber="1087" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural &quot; Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">BRÈTHES, 1925</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1057,1176,727,752]" pageId="9" pageNumber="1087" type="description">
<paragraph blockId="9.[809,1426,727,966]" box="[1057,1176,727,752]" pageId="9" pageNumber="1087">
(
<figureCitation box="[1066,1168,727,752]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStartId-0="10.[163,245,1463,1485]" captionStartId-1="11.[146,225,1546,1568]" captionStartId-2="12.[163,243,1601,1623]" captionTargetBox-0="[328,1277,202,1416]" captionTargetBox-1="[145,1425,195,1506]" captionTargetBox-2="[163,1443,196,1562]" captionTargetId-0="figure-192@10.[319,1286,195,1423]" captionTargetId-1="figure-143@11.[145,1425,195,1506]" captionTargetId-2="figure-124@12.[163,1443,196,1562]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." captionText-1="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." captionText-2="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi-0="http://doi.org/10.5281/zenodo.7814133" figureDoi-1="http://doi.org/10.5281/zenodo.7814135" figureDoi-2="http://doi.org/10.5281/zenodo.7814139" httpUri-0="https://zenodo.org/record/7814133/files/figure.png" httpUri-1="https://zenodo.org/record/7814135/files/figure.png" httpUri-2="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">FIGS 3–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[809,1228,760,782]" pageId="9" pageNumber="1087" type="reference_group">
<paragraph blockId="9.[809,1426,727,966]" box="[809,1228,760,782]" pageId="9" pageNumber="1087">
<treatmentCitationGroup box="[809,1228,760,782]" pageId="9" pageNumber="1087">
<treatmentCitation author="Brethes J" box="[809,1224,760,782]" page="199" pageId="9" pageNumber="1087" year="1925">
<taxonomicName authority="Brethes, 1925: 199" authorityName="Brethes" authorityPageNumber="199" authorityYear="1925" box="[809,1224,760,782]" class="Insecta" family="Cerasommatidiidae" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[809,997,760,781]" italics="true" pageId="9" pageNumber="1087">Cerasommatidia</emphasis>
<bibRefCitation author="Brethes J" box="[1007,1224,760,782]" pageId="9" pageNumber="1087" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural &quot; Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">Brèthes, 1925: 199</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="1087" type="type_taxon">
<paragraph blockId="9.[809,1426,727,966]" pageId="9" pageNumber="1087">
<typeStatus box="[1237,1293,760,781]" pageId="9" pageNumber="1087">Type</typeStatus>
species, by monotypy,
<taxonomicName authority="Brethes, 1925" authorityName="Brethes" authorityYear="1925" box="[962,1397,790,812]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[962,1233,790,812]" italics="true" pageId="9" pageNumber="1087">Cerasommatidia arroaei</emphasis>
<bibRefCitation author="Brethes J" box="[1241,1397,790,812]" pageId="9" pageNumber="1087" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural &quot; Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">Brèthes, 1925</bibRefCitation>
</taxonomicName>
. –
<treatmentCitation author="Pakaluk J &amp; Slipinski SA &amp; Lawrence JF" box="[841,1119,821,843]" page="228" pageId="9" pageNumber="1087" year="1994">
<bibRefCitation author="Pakaluk J &amp; Slipinski SA &amp; Lawrence JF" box="[841,1119,821,843]" pageId="9" pageNumber="1087" pagination="223 - 268" refId="ref22691" refString="Pakaluk J, Slipinski SA, Lawrence JF. 1994. Current classification and family-group names in Cucujoidea [Coleoptera]. Genus. International Journal of InVertebrate Taxonomy 5: 223 - 268." type="journal article" year="1994">
Pakaluk
<emphasis box="[942,997,821,843]" italics="true" pageId="9" pageNumber="1087">et al.</emphasis>
, 1994: 228
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Shockley FW &amp; Tomaszewska KW &amp; McHugh JV" box="[1129,1401,821,843]" page="27" pageId="9" pageNumber="1087" year="2009">
<bibRefCitation author="Shockley FW &amp; Tomaszewska KW &amp; McHugh JV" box="[1129,1401,821,843]" pageId="9" pageNumber="1087" pagination="1 - 113" refId="ref23149" refString="Shockley FW, Tomaszewska KW, McHugh JV. 2009. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113." type="journal article" year="2009">
Shockley
<emphasis box="[1237,1292,821,843]" italics="true" pageId="9" pageNumber="1087">et al.</emphasis>
, 2009: 27
</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[809,1426,851,874]" pageId="9" pageNumber="1087" type="reference_group">
<paragraph blockId="9.[809,1426,727,966]" box="[809,1426,851,874]" pageId="9" pageNumber="1087">
<treatmentCitationGroup box="[809,1426,851,874]" pageId="9" pageNumber="1087">
<treatmentCitation author="Pakaluk J &amp; Slipinski SA" box="[809,1286,851,874]" page="717" pageId="9" pageNumber="1087" year="1990">
<taxonomicName authority="Pakaluk &amp; Slipinski, 1990: 717" authorityName="Pakaluk &amp; Slipinski" authorityPageNumber="717" authorityYear="1990" box="[809,1286,851,874]" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[809,917,852,873]" italics="true" pageId="9" pageNumber="1087">Ibicarella</emphasis>
<bibRefCitation author="Pakaluk J &amp; Slipinski SA" box="[927,1286,851,874]" pageId="9" pageNumber="1087" pagination="705 - 728" refId="ref22654" refString="Pakaluk J, Slipinski SA. 1990. Review of Eupsilobiinae (Coleoptera: Endomychidae) with descriptions of new genera and species from South America. ReVue Suisse de Zoologie 97: 705 - 728." type="journal article" year="1990">Pakaluk &amp; Ślipiński, 1990: 717</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, synon. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="1087" type="multiple">
<paragraph blockId="9.[809,1426,727,966]" pageId="9" pageNumber="1087">
<typeStatus box="[841,897,883,904]" pageId="9" pageNumber="1087">Type</typeStatus>
species, by original designation,
<taxonomicName authority="Pakaluk and Slipinski, 1990" authorityName="Pakaluk and Slipinski" authorityYear="1990" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis italics="true" pageId="9" pageNumber="1087">Ibicarella plaumanni</emphasis>
<bibRefCitation author="Pakaluk J &amp; Slipinski SA" box="[974,1294,912,935]" pageId="9" pageNumber="1087" pagination="705 - 728" refId="ref22654" refString="Pakaluk J, Slipinski SA. 1990. Review of Eupsilobiinae (Coleoptera: Endomychidae) with descriptions of new genera and species from South America. ReVue Suisse de Zoologie 97: 705 - 728." type="journal article" year="1990">Pakaluk and Ślipiński, 1990</bibRefCitation>
</taxonomicName>
. –
<treatmentCitation author="Pakaluk J &amp; Slipinski SA &amp; Lawrence JF" page="228" pageId="9" pageNumber="1087" year="1994">
<bibRefCitation author="Pakaluk J &amp; Slipinski SA &amp; Lawrence JF" pageId="9" pageNumber="1087" pagination="223 - 268" refId="ref22691" refString="Pakaluk J, Slipinski SA, Lawrence JF. 1994. Current classification and family-group names in Cucujoidea [Coleoptera]. Genus. International Journal of InVertebrate Taxonomy 5: 223 - 268." type="journal article" year="1994">
Pakaluk
<emphasis box="[841,896,944,965]" italics="true" pageId="9" pageNumber="1087">et al.</emphasis>
, 1994: 228
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Shockley FW &amp; Tomaszewska KW &amp; McHugh JV" box="[1029,1300,944,966]" page="27" pageId="9" pageNumber="1087" year="2009">
<bibRefCitation author="Shockley FW &amp; Tomaszewska KW &amp; McHugh JV" box="[1029,1300,944,966]" pageId="9" pageNumber="1087" pagination="1 - 113" refId="ref23149" refString="Shockley FW, Tomaszewska KW, McHugh JV. 2009. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113." type="journal article" year="2009">
Shockley
<emphasis box="[1136,1191,944,965]" italics="true" pageId="9" pageNumber="1087">et al.</emphasis>
, 2009: 27
</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="1087" type="diagnosis">
<paragraph blockId="9.[809,1426,1020,1808]" pageId="9" pageNumber="1087">
<emphasis box="[809,1152,1020,1041]" italics="true" pageId="9" pageNumber="1087">
Diagnosis:
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[955,1152,1020,1041]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">Cerasommatidia</taxonomicName>
</emphasis>
can be separated from other genera of
<taxonomicName authorityName="BRETHES" authorityYear="1925" box="[996,1227,1050,1072]" class="Insecta" family="Cerasommatidiidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="family">Cerasommatidiidae</taxonomicName>
by the following combination of characters: dorsal surface of the body covered by inconspicuous tiny decumbent hairs (often broken or missing) (
<figureCitation box="[1044,1164,1142,1164]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3A, B</figureCitation>
,
<figureCitation box="[1178,1210,1143,1164]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">4D</figureCitation>
,
<figureCitation box="[1224,1288,1142,1164]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5C, G</figureCitation>
), pronotum with anterior and lateral sides narrowly marginate (
<figureCitation box="[817,902,1204,1226]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">Figs 4B</figureCitation>
,
<figureCitation box="[912,944,1204,1226]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5C</figureCitation>
); prosternal process moderately wide (
<figureCitation captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">Fig. 5D</figureCitation>
); trochanters flattened and expanded posteriorly to cover the tibio-tarsal joint in repose (
<figureCitation box="[1292,1419,1265,1287]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">Figs 4E, G</figureCitation>
,
<figureCitation box="[809,844,1296,1318]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5H</figureCitation>
); abdominal ventrite 1 with complete, semicircular postcoxal lines (
<figureCitation box="[1015,1111,1326,1348]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Figs 3E</figureCitation>
,
<figureCitation box="[1129,1163,1327,1348]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="9" pageNumber="1087">4G</figureCitation>
,
<figureCitation box="[1181,1216,1326,1348]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="9" pageNumber="1087">5H</figureCitation>
); aedeagus with tegmen membranous, not fused with penis (
<figureCitation captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="9" pageNumber="1087">Fig. 3E, F</figureCitation>
).
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[896,1094,1388,1409]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[896,1094,1388,1409]" italics="true" pageId="9" pageNumber="1087">Cerasommatidia</emphasis>
</taxonomicName>
resembles
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1231,1346,1388,1409]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[1231,1346,1388,1409]" italics="true" pageId="9" pageNumber="1087">MahaƲelo</emphasis>
</taxonomicName>
in the body shape, coloration, short, inconspicuous dorsal vestiture and the pronotal lateral margins without internal carina. However, it can be distinguished from the later by having the prosternal process with a pair of lateral carinae extending anteriorly beyond level of anterior margin of procoxal cavities (a single, long, median, longitudinal carina present in
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[845,967,1633,1654]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[845,967,1633,1654]" italics="true" pageId="9" pageNumber="1087">MahaƲelo</emphasis>
</taxonomicName>
), hind angles of pronotum without indentation (in
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[1001,1118,1664,1685]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[1001,1118,1664,1685]" italics="true" pageId="9" pageNumber="1087">MahaƲelo</emphasis>
</taxonomicName>
hind angles of pronotum possess small indentation to receive humeral corner of elytron), scutellar shield visible (invisible/absent in
<taxonomicName authorityName="Arriaga-Varela &amp; Tomaszewska &amp; Szawaryn &amp; Robertson &amp; Seidel &amp; Ślipiński &amp; Fikáček" authorityYear="2023" box="[844,964,1756,1777]" class="Insecta" family="Eupsilobiidae" genus="Mahavelo" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1087" phylum="Arthropoda" rank="genus">
<emphasis box="[844,964,1756,1777]" italics="true" pageId="9" pageNumber="1087">MahaƲelo</emphasis>
</taxonomicName>
) and complete semicircular femoral lines in the abdominal ventrite 1.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="1091" pageId="9" pageNumber="1087" type="description">
<paragraph blockId="9.[809,1425,1851,1904]" lastBlockId="10.[163,780,1609,1907]" lastPageId="10" lastPageNumber="1088" pageId="9" pageNumber="1087">
<emphasis box="[809,946,1852,1873]" italics="true" pageId="9" pageNumber="1087">Description:</emphasis>
Length
<quantity box="[1055,1220,1851,1873]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.7" metricValueMin="1.15" pageId="9" pageNumber="1087" unit="mm" value="1.4249999999999998" valueMax="1.7" valueMin="1.15">1.15–1.70 mm</quantity>
. Body moderately short oval, convex, 1.3–1.4 times as long as wide, 1.8–1.9 times as long as high. Coloration homogeneous: dark brown to black (
<figureCitation box="[403,514,1640,1662]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="10" pageNumber="1088">Fig. 3A, B</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814133" ID-Zenodo-Dep="7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="10" pageNumber="1088" startId="10.[163,245,1463,1485]" targetBox="[328,1277,202,1416]" targetPageId="10">
<paragraph blockId="10.[163,1443,1463,1573]" pageId="10" pageNumber="1088">
<emphasis bold="true" box="[163,272,1463,1485]" pageId="10" pageNumber="1088">Figure 3.</emphasis>
Morphology of
<taxonomicName box="[450,711,1463,1485]" class="Insecta" genus="Cerasommatidia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1088" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis box="[450,632,1463,1485]" italics="true" pageId="10" pageNumber="1088">Cerasommatidia</emphasis>
spp. A
</taxonomicName>
,
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[722,986,1463,1485]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1088" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[722,986,1463,1485]" italics="true" pageId="10" pageNumber="1088">Cerasommatidia arroaei</emphasis>
</taxonomicName>
holotype, dorsal habitus of dismembered specimen. B,
<taxonomicName baseAuthorityName="Pakaluk &amp; Slipinski" baseAuthorityYear="1990" box="[302,602,1493,1515]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1088" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis box="[302,602,1493,1515]" italics="true" pageId="10" pageNumber="1088">Cerasommatidia plaumanni</emphasis>
</taxonomicName>
paratype, dorsal habitus. C–E,
<taxonomicName baseAuthorityName="Pakaluk &amp; Slipinski" baseAuthorityYear="1990" box="[938,1238,1493,1515]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1088" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis box="[938,1238,1493,1515]" italics="true" pageId="10" pageNumber="1088">Cerasommatidia plaumanni</emphasis>
</taxonomicName>
: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F,
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[292,542,1551,1573]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1088" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[292,542,1551,1573]" italics="true" pageId="10" pageNumber="1088">Cerasommatidia arroaei</emphasis>
</taxonomicName>
male genitalia. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph blockId="10.[163,780,1609,1907]" lastBlockId="10.[827,1443,1609,1907]" pageId="10" pageNumber="1088">
Head with dorsal surface uniformly covered with short fine setae (
<figureCitation box="[436,529,1701,1723]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="10" pageNumber="1088">Figs 4A</figureCitation>
,
<figureCitation box="[546,578,1701,1723]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="10" pageNumber="1088">5A</figureCitation>
). Clypeus large, rectangular. Frontoclypeal suture present but feebly marked, almost straight. Ventral antennal grooves short, reaching close to posterior eye level (
<figureCitation captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="10" pageNumber="1088">Fig. 5B</figureCitation>
). Antenna approximately 0.35 of length of body (
<figureCitation box="[172,297,1854,1876]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="10" pageNumber="1088">Figs 3A, C</figureCitation>
,
<figureCitation box="[312,344,1854,1876]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="10" pageNumber="1088">5E</figureCitation>
); antennal club one quarter of total antennal length; antennomeres 1–5 longer than wide, with antennomere 5 longer than antennomere 6; antennomeres 6–9 variable. Eyes comparatively small, moderately prominent (
<figureCitation box="[1102,1190,1670,1692]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="10" pageNumber="1088">Figs 3C</figureCitation>
,
<figureCitation box="[1203,1235,1670,1691]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="10" pageNumber="1088">4A</figureCitation>
,
<figureCitation box="[1249,1281,1670,1692]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="10" pageNumber="1088">5A</figureCitation>
). Galea large, densely setose apically. Lacinia weakly narrower than galea with dense setae at apex and inner margin. Labium (
<figureCitation box="[930,1011,1762,1784]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="10" pageNumber="1088">Fig. 5B</figureCitation>
) with mentum widest near midlength or basal third, palpomere 2 and 3 large, bulbous, somewhat oval and subrectangular respectively; terminal palpomere short and subtruncate to weakly elongate, acuminate, 1.2–1.5 times as long as wide.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814135" ID-Zenodo-Dep="7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="11" pageNumber="1089" startId="11.[146,225,1546,1568]" targetBox="[145,1425,195,1506]" targetPageId="11">
<paragraph blockId="11.[145,1425,1546,1598]" pageId="11" pageNumber="1089">
<emphasis bold="true" box="[146,251,1546,1568]" pageId="11" pageNumber="1089">Figure 4.</emphasis>
Morphology of
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[418,670,1546,1568]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1089" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[418,670,1546,1568]" italics="true" pageId="11" pageNumber="1089">Cerasommatidia arroaei</emphasis>
</taxonomicName>
. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line).
</paragraph>
</caption>
<paragraph blockId="11.[145,762,1665,1871]" lastBlockId="12.[163,780,1710,1885]" lastPageId="12" lastPageNumber="1090" pageId="11" pageNumber="1089">
Prothorax. Pronotum 2.2–2.5 times as wide as long, widest at base and strongly convergent anteriorly, 1.9–2.2 times wider at widest part than on front angles (
<figureCitation box="[672,754,1726,1748]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="11" pageNumber="1089">Figs 4B</figureCitation>
,
<figureCitation box="[145,175,1757,1779]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="11" pageNumber="1089">5C</figureCitation>
). Anterior margin (at least partly) and lateral margins narrowly bordered; base of pronotum regularly rounded, without distinct bordering line or with faint bordering line present medially. Pronotal sides scarcely rounded; hind angles without indentation to receive humeral corner of elytron. Pronotal disc moderately convex, smooth, with sparse fine punctures provided with thin, tiny hairs. Prosternum (
<figureCitation box="[1085,1167,1757,1779]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="11" pageNumber="1089">Fig. 5D</figureCitation>
) with anterior margin weakly arcuate posteriorly; prosternal process extending posteriorly to hind level of procoxae, comparatively wide, at apex about 0.7–0.8 of the width of procoxal cavity; with raised lateral carinae reaching anteriorly well beyond level of anterior margin of procoxal cavities; area between carinae weakly concave. Hypomeron with deep, long, straight antennal grooves (
<figureCitation box="[515,594,1802,1824]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="12" pageNumber="1090">Fig. 5D</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7814139" ID-Zenodo-Dep="7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="12" pageNumber="1090" startId="12.[163,243,1601,1623]" targetBox="[163,1443,196,1562]" targetPageId="12">
<paragraph blockId="12.[163,1443,1601,1682]" pageId="12" pageNumber="1090">
<emphasis bold="true" box="[163,271,1601,1623]" pageId="12" pageNumber="1090">Figure 5.</emphasis>
Morphology of
<taxonomicName baseAuthorityName="Pakaluk &amp; Slipinski" baseAuthorityYear="1990" box="[444,749,1602,1624]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1090" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis box="[444,749,1602,1624]" italics="true" pageId="12" pageNumber="1090">Cerasommatidia plaumanni</emphasis>
</taxonomicName>
. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line).
</paragraph>
</caption>
<paragraph blockId="12.[163,780,1710,1885]" lastBlockId="13.[145,762,197,1047]" lastPageId="13" lastPageNumber="1091" pageId="12" pageNumber="1090">
Pterothorax. Mesonotum with scutellar shield small, transverse, weakly rounded at apex (
<figureCitation box="[732,859,1710,1885]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="12" pageNumber="1090">Fig. 5C</figureCitation>
). Mesoventrite (
<figureCitation box="[1056,1148,1710,1732]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="12" pageNumber="1090">Figs 3D</figureCitation>
,
<figureCitation box="[1163,1195,1710,1731]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="12" pageNumber="1090">4E</figureCitation>
,
<figureCitation box="[1210,1245,1710,1732]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="12" pageNumber="1090">5H</figureCitation>
) with intercoxal process smooth, almost flat with anterior raised border incomplete medially; 1.2–1.4 times as wide as mesocoxal diameter. Metaventrite (
<figureCitation box="[1117,1210,1802,1824]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="12" pageNumber="1090">Figs 3D</figureCitation>
,
<figureCitation box="[1224,1256,1802,1823]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="12" pageNumber="1090">4E</figureCitation>
,
<figureCitation box="[1272,1307,1802,1824]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="12" pageNumber="1090">5H</figureCitation>
) as long as abdominal ventrites 1–3 together; with remnants of discrimen posteriorly; anterior margin with bordering carina widening towards lateral corners; central area with small to large and deep setiferous punctures. Anterior part of metanepisternum with small outer blunt like projection (
<figureCitation box="[402,488,289,311]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="13" pageNumber="1091">Fig. 3D</figureCitation>
). Elytra 0.85–1.00 mm long (
<figureCitation box="[211,301,320,342]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="13" pageNumber="1091">Figs 3A</figureCitation>
,
<figureCitation box="[316,348,320,341]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="13" pageNumber="1091">4C</figureCitation>
,
<figureCitation box="[362,396,320,342]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="13" pageNumber="1091">5G</figureCitation>
), about as long a wide, 2.9–3.3 times as long and 1.2 times as wide as pronotum, with lateral margins narrowly explanate. Surface with small setiferous punctures bearing tiny, thin setae. Epipleura almost reaching elytral apex but incomplete, narrow, with internal bordering line narrow, present from the level of mid coxae to apex. Hindwings reduced or absent.
</paragraph>
<paragraph blockId="13.[145,762,197,1047]" pageId="13" pageNumber="1091">
Legs (
<figureCitation box="[244,333,534,556]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="13" pageNumber="1091">Fig. 3D</figureCitation>
). Trochanters angulately produced posteriorly. Meso- and metatrochanters with posterior margin as a conspicuous laminar expansion that covers tibio-tarsal joint in repose. Femora flattened (mid and hind femora more distinctly than fore femora); with grooves for tibiae present throughout whole length. Claws with comparatively small blunt tooth at base (
<figureCitation box="[632,708,718,740]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="13" pageNumber="1091">Figs 4F</figureCitation>
,
<figureCitation box="[719,747,718,740]" captionStart="Figure 5" captionStartId="12.[163,243,1601,1623]" captionTargetBox="[163,1443,196,1562]" captionTargetId="figure-124@12.[163,1443,196,1562]" captionTargetPageId="12" captionText="Figure 5. Morphology of Cerasommatidia plaumanni. A, head, dorsal. B, head, ventral. C, pronotum (arrow: posterior margin of pronotum). D, prosternum. E, antenna. F, mesotarsus. G, left elytron, dorsal. H, meso- and metaventrite, and first abdominal ventrite (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814139" httpUri="https://zenodo.org/record/7814139/files/figure.png" pageId="13" pageNumber="1091">5F</figureCitation>
).
</paragraph>
<paragraph blockId="13.[145,762,197,1047]" pageId="13" pageNumber="1091">
Abdomen (
<figureCitation box="[297,388,749,771]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="13" pageNumber="1091">Figs 3E</figureCitation>
,
<figureCitation box="[404,437,749,770]" captionStart="Figure 4" captionStartId="11.[146,225,1546,1568]" captionTargetBox="[145,1425,195,1506]" captionTargetId="figure-143@11.[145,1425,195,1506]" captionTargetPageId="11" captionText="Figure 4. Morphology of Cerasommatidia arroaei. A, head, dorsal. B, pronotum (arrow: posterior margin of pronotum). C, left elytron, dorsal. D, details of elytral surface. E, metaventrite. F, metatarsus. G, abdomen, ventral (arrow: postcoxal line)." figureDoi="http://doi.org/10.5281/zenodo.7814135" httpUri="https://zenodo.org/record/7814135/files/figure.png" pageId="13" pageNumber="1091">4G</figureCitation>
) with ventrite 1 anteriorly with bordering carina forming distinct, rounded and complete laterally postcoxal lines, extending posteriorly beyond midlength of ventrite 1.
</paragraph>
<paragraph blockId="13.[145,762,197,1047]" pageId="13" pageNumber="1091">
Male genitalia (
<figureCitation box="[364,483,872,894]" captionStart="Figure 3" captionStartId="10.[163,245,1463,1485]" captionTargetBox="[328,1277,202,1416]" captionTargetId="figure-192@10.[319,1286,195,1423]" captionTargetPageId="10" captionText="Figure 3. Morphology of Cerasommatidia spp. A, Cerasommatidia arroaei holotype, dorsal habitus of dismembered specimen. B, Cerasommatidia plaumanni paratype, dorsal habitus. C–E, Cerasommatidia plaumanni: C, head, dorsal. D, meso- and metaventral structures. E, abdomen with male terminalia and genitalia partially visible through the abdominal ventrites. F, Cerasommatidia arroaei male genitalia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7814133" httpUri="https://zenodo.org/record/7814133/files/figure.png" pageId="13" pageNumber="1091">Fig. 3E, F</figureCitation>
). Aedeagus with penis long and narrow, sclerotized, curved or asymmetrically sinuate, not-ramificate apically. Tegmen: large, in form of submembranous ring; tegminal strut long, membranous, flattened.
</paragraph>
<paragraph blockId="13.[145,762,197,1047]" box="[169,500,1025,1047]" pageId="13" pageNumber="1091">Female genitalia not studied.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="1091" type="synonymic_list">
<paragraph blockId="13.[145,762,1101,1185]" pageId="13" pageNumber="1091">
<emphasis box="[145,348,1101,1122]" italics="true" pageId="13" pageNumber="1091">Species included:</emphasis>
<taxonomicName authority="Brethes, 1925" authorityName="Brethes" authorityYear="1925" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[370,653,1101,1123]" italics="true" pageId="13" pageNumber="1091">Cerasommatidia arroaei</emphasis>
<bibRefCitation author="Brethes J" pageId="13" pageNumber="1091" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural &quot; Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">Brèthes, 1925</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="(Pakaluk &amp; Slipinski, 1990)" baseAuthorityName="Pakaluk &amp; Slipinski" baseAuthorityYear="1990" box="[222,756,1131,1154]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis box="[222,393,1132,1153]" italics="true" pageId="13" pageNumber="1091">C. plaumanni</emphasis>
(
<bibRefCitation author="Pakaluk J &amp; Slipinski SA" box="[414,744,1131,1154]" pageId="13" pageNumber="1091" pagination="705 - 728" refId="ref22654" refString="Pakaluk J, Slipinski SA. 1990. Review of Eupsilobiinae (Coleoptera: Endomychidae) with descriptions of new genera and species from South America. ReVue Suisse de Zoologie 97: 705 - 728." type="journal article" year="1990">Pakaluk &amp; Ślipiński, 1990</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="(Pakaluk &amp; Slipinski, 1990)" baseAuthorityName="Pakaluk &amp; Slipinski" baseAuthorityYear="1990" box="[145,604,1162,1185]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="species" species="rotundata">
<emphasis box="[145,287,1162,1184]" italics="true" pageId="13" pageNumber="1091">C. rotundata</emphasis>
(
<bibRefCitation author="Pakaluk J &amp; Slipinski SA" box="[302,593,1162,1185]" pageId="13" pageNumber="1091" pagination="705 - 728" refId="ref22654" refString="Pakaluk J, Slipinski SA. 1990. Review of Eupsilobiinae (Coleoptera: Endomychidae) with descriptions of new genera and species from South America. ReVue Suisse de Zoologie 97: 705 - 728." type="journal article" year="1990">Pakaluk &amp; Ślipiński, 1990</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection box="[145,501,1241,1263]" pageId="13" pageNumber="1091" type="distribution">
<paragraph blockId="13.[145,501,1241,1263]" box="[145,501,1241,1263]" pageId="13" pageNumber="1091">
<emphasis box="[145,289,1241,1262]" italics="true" pageId="13" pageNumber="1091">Distribution:</emphasis>
<collectingCountry box="[308,377,1241,1263]" name="Brazil" pageId="13" pageNumber="1091">Brazil</collectingCountry>
(
<figureCitation box="[392,486,1241,1263]" captionStart="Figure 20" captionStartId="34.[162,240,1081,1103]" captionTargetBox="[162,1442,200,1042]" captionTargetId="figure-327@34.[162,1442,200,1042]" captionTargetPageId="34" captionText="Figure 20. Geographical distribution of Cerasommatidiidae spp. A, distribution of species in eastern Caribbean region. B, distribution of species in southeastern Brazil. C, distribution of species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.7814187" httpUri="https://zenodo.org/record/7814187/files/figure.png" pageId="13" pageNumber="1091">Fig. 20B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="1091" type="discussion">
<paragraph blockId="13.[145,762,1311,1885]" pageId="13" pageNumber="1091">
<emphasis box="[145,261,1312,1333]" italics="true" pageId="13" pageNumber="1091">Comment:</emphasis>
When
<bibRefCitation author="Pakaluk J &amp; Slipinski SA" box="[348,648,1311,1334]" pageId="13" pageNumber="1091" pagination="705 - 728" refId="ref22654" refString="Pakaluk J, Slipinski SA. 1990. Review of Eupsilobiinae (Coleoptera: Endomychidae) with descriptions of new genera and species from South America. ReVue Suisse de Zoologie 97: 705 - 728." type="journal article" year="1990">Pakaluk &amp; Ślipiński (1990)</bibRefCitation>
described
<taxonomicName authorityName="Pakaluk &amp; Slipinski" authorityYear="1990" box="[145,262,1342,1363]" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[145,262,1342,1363]" italics="true" pageId="13" pageNumber="1091">Ibicarella</emphasis>
</taxonomicName>
in Eupsilobiinae they were not aware of its resemblance to
<taxonomicName authority="Brethes, 1925" authorityName="Brethes" authorityYear="1925" box="[377,754,1373,1395]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[377,577,1373,1394]" italics="true" pageId="13" pageNumber="1091">Cerasommatidia</emphasis>
<bibRefCitation author="Brethes J" box="[587,754,1373,1395]" pageId="13" pageNumber="1091" pagination="195 - 214" refId="ref22201" refString="Brethes J. 1925. Coccinellides du British Museum (avec une nouvelle famille de Coleopteres). Anales del Museo Nacional de Historia Natural &quot; Bernardino RiVadaVia 33: 195 - 214." type="journal article" year="1925">Brèthes, 1925</bibRefCitation>
</taxonomicName>
. Subsequently, when the
<typeStatus box="[431,530,1404,1426]" pageId="13" pageNumber="1091">holotype</typeStatus>
of
<taxonomicName authorityName="Brethes" authorityYear="1925" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis italics="true" pageId="13" pageNumber="1091">Cerasommatidia arroaei</emphasis>
</taxonomicName>
was rediscovered its resemblance with
<taxonomicName authorityName="Pakaluk &amp; Slipinski" authorityYear="1990" box="[145,263,1465,1486]" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[145,263,1465,1486]" italics="true" pageId="13" pageNumber="1091">Ibicarella</emphasis>
</taxonomicName>
was noticed and served as basis for the synonymy of
<taxonomicName authorityName="BRETHES" authorityYear="1925" box="[295,520,1496,1518]" class="Insecta" family="Cerasommatidiidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="family">Cerasommatidiidae</taxonomicName>
under
<taxonomicName authority="(Pakaluk et al., 1994)" baseAuthorityName="Pakaluk" baseAuthorityYear="1994" class="Bacillariophyceae" family="Plagiogrammaceae" genus="Eupsilobiinae" higherTaxonomySource="GBIF" kingdom="Chromista" order="Triceratiales" pageId="13" pageNumber="1091" phylum="Ochrophyta" rank="genus">
Eupsilobiinae (
<bibRefCitation author="Pakaluk J &amp; Slipinski SA &amp; Lawrence JF" box="[154,396,1526,1548]" pageId="13" pageNumber="1091" pagination="223 - 268" refId="ref22691" refString="Pakaluk J, Slipinski SA, Lawrence JF. 1994. Current classification and family-group names in Cucujoidea [Coleoptera]. Genus. International Journal of InVertebrate Taxonomy 5: 223 - 268." type="journal article" year="1994">
Pakaluk
<emphasis box="[262,323,1526,1548]" italics="true" pageId="13" pageNumber="1091">et al.</emphasis>
, 1994
</bibRefCitation>
)
</taxonomicName>
. Nevertheless, in part due to the state of preservation of the
<typeStatus box="[512,611,1557,1579]" pageId="13" pageNumber="1091">holotype</typeStatus>
of
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[647,754,1557,1579]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="species" species="arroaei">
<emphasis box="[647,754,1557,1579]" italics="true" pageId="13" pageNumber="1091">C. arroaei</emphasis>
</taxonomicName>
, its morphology was not thoroughly studied and both
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[145,336,1618,1639]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[145,336,1618,1639]" italics="true" pageId="13" pageNumber="1091">Cerasommatidia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Pakaluk &amp; Slipinski" authorityYear="1990" box="[391,502,1618,1639]" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[391,502,1618,1639]" italics="true" pageId="13" pageNumber="1091">Ibicarella</emphasis>
</taxonomicName>
were retained as valid genera. In the present study, we carefully examined the morphology of the type material of all
<taxonomicName authorityName="Brethes" authorityYear="1925" box="[570,761,1680,1701]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[570,761,1680,1701]" italics="true" pageId="13" pageNumber="1091">Cerasommatidia</emphasis>
</taxonomicName>
and
<taxonomicName box="[194,392,1710,1732]" pageId="13" pageNumber="1091">
<emphasis box="[194,305,1710,1731]" italics="true" pageId="13" pageNumber="1091">Ibicarella</emphasis>
species
</taxonomicName>
and, following the results of our morphological (Supporting Information, File S12a–c) and combined (
<figureCitation box="[336,407,1772,1794]" captionStart="Figure 2" captionStartId="6.[163,243,1744,1766]" captionTargetBox="[246,1361,198,1697]" captionTargetId="figure-25@6.[242,1364,194,1704]" captionTargetPageId="6" captionText="Figure 2. Results of the phylogenetic analyses of the combined, morphological and molecular dataset under Bayesian inference. Statistical support for branches show posterior probabilities. Branch lengths are kept and shown in the same proportion all along the tree. Apomorphies tracked in WINCLADA are shown upon each branch. DNA helix graphic after taxon name indicates taxa for which molecular data were available; asterisk ‘*’ indicates interspecific chimeras (combined data from two different species of the same genus)." figureDoi="http://doi.org/10.5281/zenodo.7814131" httpUri="https://zenodo.org/record/7814131/files/figure.png" pageId="13" pageNumber="1091">Fig. 2</figureCitation>
) analyses, we consider both represent the same genus and
<taxonomicName authority="Pakaluk &amp; Slipinski" authorityName="Pakaluk &amp; Slipinski" authorityYear="1990" class="Insecta" family="Endomychidae" genus="Ibicarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[512,626,1802,1823]" italics="true" pageId="13" pageNumber="1091">Ibicarella</emphasis>
Pakaluk &amp; Ślipiński
</taxonomicName>
is considered a subjective junior synonym of
<taxonomicName authority="Arrow." box="[145,419,1864,1885]" class="Insecta" genus="Cerasommatidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1091" phylum="Arthropoda" rank="genus">
<emphasis box="[145,336,1864,1885]" italics="true" pageId="13" pageNumber="1091">Cerasommatidia</emphasis>
Arrow.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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