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<document id="444F68C89D5B71484767DDE8A8AC1254" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FFB3FFCFFE1FB957A4CBFC24" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Coccidencyrtus cyathus Noyes 2023, sp.nov." docType="treatment" docVersion="3" lastPageNumber="103" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="101" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="A916091D4887E951923D5755C38ACA5F">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="D091E259ED768E1381899A388FB0854A">Noyes, John Stuart</mods:namePart>
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<mods:title id="87DF2DF4514EF826B7244470DA9C1527">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="55290BA80C76A2FF4C5CD14D0A25C7CA">2023</mods:date>
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<treatment id="03BA87A7FFB3FFCFFE1FB957A4CBFC24" LSID="urn:lsid:plazi:treatment:03BA87A7FFB3FFCFFE1FB957A4CBFC24" httpUri="http://treatment.plazi.org/id/03BA87A7FFB3FFCFFE1FB957A4CBFC24" lastPageId="104" lastPageNumber="103" pageId="102" pageNumber="101">
<subSubSection id="C309653AFFB3FFC1FE1FB957A555FA49" box="[509,837,1395,1420]" pageId="102" pageNumber="101" type="nomenclature">
<paragraph id="8BAC36B1FFB3FFC1FE1FB957A555FA49" blockId="102.[509,837,1395,1447]" box="[509,837,1395,1420]" pageId="102" pageNumber="101">
<heading id="D0E481DDFFB3FFC1FE1FB957A555FA49" bold="true" box="[509,837,1395,1420]" centered="true" fontSize="10" level="2" pageId="102" pageNumber="101" reason="2">
<emphasis id="B967EAA3FFB3FFC1FE1FB957A555FA49" bold="true" box="[509,837,1395,1420]" pageId="102" pageNumber="101">
<taxonomicName id="4C134D32FFB3FFC1FE1FB957A4E4FA4E" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[509,756,1395,1419]" class="Insecta" family="Encyrtidae" genus="Coccidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="101" phylum="Arthropoda" rank="species" species="cyathus" status="sp. nov.">
<emphasis id="B967EAA3FFB3FFC1FE1FB957A4E4FA4E" bold="true" box="[509,756,1395,1419]" italics="true" pageId="102" pageNumber="101">Coccidencyrtus cyathus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457D8FFB3FFC1FD18B950A555FA49" box="[762,837,1396,1420]" pageId="102" pageNumber="101" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFB3FFC1FDFBB9AAA539FA63" box="[537,809,1422,1447]" pageId="102" pageNumber="101" type="description">
<paragraph id="8BAC36B1FFB3FFC1FDFBB9AAA539FA63" blockId="102.[509,837,1395,1447]" box="[537,809,1422,1447]" pageId="102" pageNumber="101">
(
<figureCitation id="13282A34FFB3FFC1FDC2B9AAA4BCFA62" box="[544,684,1422,1447]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="102" pageNumber="101">Figs 219-222</figureCitation>
; Hab. E 35)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFB3FFCFFF57B9E0A771FDA9" lastPageId="104" lastPageNumber="103" pageId="102" pageNumber="101" type="diagnosis">
<paragraph id="8BAC36B1FFB3FFC0FF57B9E0A70BFE53" blockId="102.[181,1165,1476,1686]" lastBlockId="103.[181,1165,222,406]" lastPageId="103" lastPageNumber="102" pageId="102" pageNumber="101">
DIAGNOSIS.
<emphasis id="B967EAA3FFB3FFC1FEAEB9E0A788FA19" box="[332,408,1476,1500]" italics="true" pageId="102" pageNumber="101">Female</emphasis>
(overall length about
<quantity id="4CEB9B54FFB3FFC1FD9CB9E0A4E2FA19" box="[638,754,1476,1500]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="102" pageNumber="101" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6-0.9mm</quantity>
): scape largely dark brown; pedicel and funicle yellow, clava pale brown; fore coxa pale yellow, mid and hind coxa dark brown; fore femur and tibia yellow, tibia with a very faint, pale brown subbasal band; mid femur white with a dark brown subapical band; hind femur mostly dark brown; mid and hind tibiae white with a dark brown subbasal band; gaster largely dark brown, but Gt1 medially pale orange; hypopygium brown; frontovertex about one-third to half head width; antenna (
<figureCitation id="13282A34FFB3FFC1FCB1BA6DA5BBF9A4" box="[851,939,1609,1633]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="102" pageNumber="101">Fig. 219</figureCitation>
) with scape about 3X as long as broad, funicle with F1 narrower than pedicel, funicle with F1-F5 slightly transverse, F6 largest and slightly longer than broad; linear sensilla only on F6; clava about as long as pedicel and funicle combined, about 3.5X as long as broad; palp formula 3-2; scutellum with 3 pairs of setae and fine, longitudinally elongate striate sculpture, about as deep as that on mesoscutum; mid basitarsus with 3 or 4 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with stigmal vein (
<figureCitation id="13282A34FFB2FFC0FE91BD0AA7DAFE82" box="[371,458,302,327]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="103" pageNumber="102">Fig. 221</figureCitation>
) slightly longer than postmarginal vein and about as long as marginal vein; hypopygium reaching about 0.7X towards apex of gaster; ovipositor, about 3.5X as long as gonostylus and slightly exserted with exserted part a little shorter than mid tibial spur.
<emphasis id="B967EAA3FFB2FFC0FBB3BD40A295FEB9" box="[1105,1157,356,380]" italics="true" pageId="103" pageNumber="102">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FFB2FFC0FF57BD90A293FE09" blockId="103.[181,1165,435,1687]" box="[181,1155,435,460]" pageId="103" pageNumber="102">
Female
<emphasis id="B967EAA3FFB2FFC0FEE4BD97A71EFE0E" bold="true" box="[262,270,435,459]" pageId="103" pageNumber="102">(</emphasis>
<typeStatus id="54A88813FFB2FFC0FEEDBD90A778FE09" box="[271,360,436,460]" pageId="103" pageNumber="102" type="holotype">holotype</typeStatus>
<emphasis id="B967EAA3FFB2FFC0FE85BD97A77FFE0E" bold="true" box="[359,367,435,459]" pageId="103" pageNumber="102">)</emphasis>
: length, including ovipositor,
<quantity id="4CEB9B54FFB2FFC0FD40BD90A4E8FE09" box="[674,760,436,460]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="103" pageNumber="102" unit="mm" value="0.84">0.84mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FFB2FFC0FC3DBD90A223FE09" box="[991,1075,436,460]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="103" pageNumber="102" unit="mm" value="0.79">0.79mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FFB2FFC0FF39BDEAA29DFBD2" blockId="103.[181,1165,435,1687]" pageId="103" pageNumber="102">Head dark brown to black with a weak to moderate metallic lustre; occiput with a weak brassy sheen; frontovertex generally metallic dark green, area between posterior ocelli and occipital margin coppery; area above scrobes coppery; scrobal area coppery to coppery purple; temple and gena relatively dull with a very slight brassy sheen; frontovertex with a few, inconspicuous, scattered, brown setae; gena and interantennal prominence with a few scattered, very inconspicuous, brown setae; maxillary palpus white; radicle dark brown; scape dark brown with extreme apex yellow; pedicel and funicle yellow, base of pedicel and F1-F5 slightly dusky; clava pale brown, distal part of basal segment slightly paler; thorax dark brown to black with a metallic lustre; pronotum with a weak coppery purple and brassy sheen; mesoscutum virtually matt with a very weak coppery purple sheen; tegula dark purple-brown with a weak brassy sheen; axilla with a weak coppery purple and brassy sheen; scutellum dorsally similar to mesoscutum, extreme side and apex shining and slightly brassy; mesopleuron with a moderate purple and brassy sheen; fore coxa very pale yellow; fore femur very pale yellow, tibia very pale yellow, with indication of a very faint, interrupted, pale brown, subbasal ring, tarsus pale orange with apical tarsomere slightly dusky; mid coxa dark brown; mid femur white in proximal 0.5X, a broad, dark brown subapical band, extreme apex white, tibia white to very pale orange and a narrow, subbasal dark, brown band, spur and tarsus very pale orange; hind coxa dark brown; hind femur with proximal 0.25X white, extreme apex white, otherwise dark brown, tibia white at extreme base, a narrow, subbasal, brown band, otherwise very pale orange, tarsus very pale orange, apical tarsomere slightly dusky; wings hyaline; propodeum dark brown with a weak brassy and purple sheen; gaster dark purple-brown, dorsally with a coppery purple sheen, proximal part of Gt1 with a distinct pale orange area medially, side and venter with coppery, blue, brassy and green reflections; gonostylus dark brown.</paragraph>
<paragraph id="8BAC36B1FFB2FFC0FF39B83DA5A6FA32" blockId="103.[181,1165,435,1687]" pageId="103" pageNumber="102">
Head about 2.2X as wide as frontovertex, in profile about 1.3X as high as deep, tangent to frontovertex forming an angle of about 80° with that of face; occipital margin more or less sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 90°; frontovertex hardly widening in front of anterior ocellus, with distinct polygonally reticulate sculpture of mesh size a little larger than eye facet, shallower above top of scrobes; temple and gena with relatively shallow, longitudinally elongate, fine polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture; interantennal prominence and mouth margin with shallow, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally rounded; antenna as in
<figureCitation id="13282A34FFB2FFC0FE40B92DA7EAFAE7" box="[418,506,1289,1314]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="103" pageNumber="102">Fig. 219</figureCitation>
; scape very slightly broadened and flattened, about 2.9X as long as broad; funicle with F1-F5 small, subequal and slightly transverse, F6 conspicuously larger but also slightly transverse; clava moderately slender, about 3.5X as long as broad, about 1.6X as wide as F6, not downcurved ventrally, about as long as pedicel and funicle combined, sutures parallel, sensory area relatively small, at apex only giving it a rounded or weak obliquely truncate appearance; eye slightly overreaching occipital margin, clothed with sparse, short inconspicuous setae, each not longer than diameter of facet; malar sulcus present, but very shallow and indistinct; clypeal margin evenly very slightly concave. Relative measurements: HW 42.5,
<collectingRegion id="49D7F853FFB2FFC0FC13B9E1A206FA19" box="[1009,1046,1477,1500]" country="Swaziland" name="Hhohho" pageId="103" pageNumber="102">HH</collectingRegion>
32, FV 19, POL 10, OOL 1.5, OCL 3.5, AOL 7, EL 23, EW 21, MS 14, SL 19, SW 6.5.
</paragraph>
<paragraph id="8BAC36B1FFB2FFCFFF39B9DDA7BFFEBE" blockId="103.[181,1165,435,1687]" lastBlockId="104.[181,1164,222,567]" lastPageId="104" lastPageNumber="103" pageId="103" pageNumber="102">
Thorax (
<figureCitation id="13282A34FFB2FFC0FED6B9DDA79DF9D7" box="[308,397,1529,1554]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="103" pageNumber="102">Fig. 220</figureCitation>
) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum with polygonally reticulate sculpture that is about as deep as that in ocellar area, and of similar mesh; mesoscutum with very fine, longitudinally elongate, polygonally reticulate to striate-reticulate sculpture that is very slightly deeper than that on pronotum and quite silky in appearance; axilla with similar sculpture to pronotum, but slightly more transversely elongate; scutellum dorsally with very similar sculpture to mesoscutum, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.9X as broad as long; scutellum about 1.25X as broad as long, apex with a pair of slender, elongate setae; mid basitarsus with 3 or 4 ventral pegs; mid tibial spur about as long as basitarsus; hind femur about 3.6X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FFBDFFCFFC61BD0AA21FFE82" box="[899,1039,302,327]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="104" pageNumber="103">Figs 221, 222</figureCitation>
; propodeum (
<figureCitation id="13282A34FFBDFFCFFF5FBD6DA705FEA4" box="[189,277,329,353]" captionStart="Figs 209-222" captionStartId="752.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@752.[181,1163,223,1508]" captionTargetPageId="752" captionText="Figs 209-222. Coccidencyrtus procris (209) mesosoma, dorsal view; (210) hypopygium (posterior margin slightly curled inwards). Coccidencyrtus umbro (211) head, facial view; (212) fore wing; (213) apex of fore wing venation; (214) mesosoma, dorsal view); (215) antenna; (216) hypopygium (posterior margin partially curled inwards); (217) antenna, G; (218) genitalia, G. Coccidencyrtus cyathus (219) antenna; (220) mesosoma, dorsal view; (221) apex of fore wing venation; (222) fore wing." pageId="104" pageNumber="103">Fig. 220</figureCitation>
) medially about 0.07X as long as scutellum. Relative measurements: FWL 44, FWW 18.5; HWL 29, HWW 7.
</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF39BD5BA7CCFE09" blockId="104.[181,1164,222,567]" pageId="104" pageNumber="103">Gaster with hypopygium reaching about 0.7X to apex; last tergite about 0.7X as long as mid tibia, with apex rounded; ovipositor exserted, the exserted part about 0.6X as long as mid tibial spur or about 0.2X mid tibia.</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF39BDEBA26BFDC4" blockId="104.[181,1164,222,567]" pageId="104" pageNumber="103">
<typeStatus id="54A88813FFBDFFCFFF39BDEBA724FE22" box="[219,308,463,487]" pageId="104" pageNumber="103" type="paratype">Paratype</typeStatus>
. Mandible with two sharp teeth and an upper, rounded tooth; palp formula 3-2; funicle with linear sensilla present only on F6. Relative measurements: OL 92, GL 25 [MT 81].
</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF39BE20A26BFDF3" blockId="104.[181,1164,222,567]" pageId="104" pageNumber="103">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FFBDFFCFFD5BBE20A554FDD9" box="[697,836,516,540]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="8.8" metricValueMin="5.6" pageId="104" pageNumber="103" unit="mm" value="0.72" valueMax="0.88" valueMin="0.56">0.56-0.88mm</quantity>
, the head varies from about 2.0- 2.7X as wide as the frontovertex, with the frontovertex relatively narrower in larger specimens.
</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF57BE70A771FDA9" blockId="104.[181,353,596,620]" box="[181,353,596,620]" pageId="104" pageNumber="103">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFFBDFFCFFF57BEADA459FD64" box="[181,585,649,673]" pageId="104" pageNumber="103" type="distribution">
<paragraph id="8BAC36B1FFBDFFCFFF57BEADA459FD64" blockId="104.[181,585,649,673]" box="[181,585,649,673]" pageId="104" pageNumber="103">
DISTRIBUTION.
<collectingCountry id="F3047621FFBDFFCFFE91BEADA7F4FD64" box="[371,484,649,673]" name="Costa Rica" pageId="104" pageNumber="103">Costa Rica</collectingCountry>
,
<collectingCountry id="F3047621FFBDFFCFFE0DBEADA455FD64" box="[495,581,649,673]" name="Ecuador" pageId="104" pageNumber="103">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFBDFFCFFF57BE9BA76EFD13" box="[181,382,702,727]" pageId="104" pageNumber="103" type="biology_ecology">
<paragraph id="8BAC36B1FFBDFFCFFF57BE9BA76EFD13" blockId="104.[181,382,702,727]" box="[181,382,702,727]" pageId="104" pageNumber="103">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFFBDFFCFFF57BED2A47BFC6E" pageId="104" pageNumber="103" type="materials_examined">
<paragraph id="8BAC36B1FFBDFFCFFF57BED2A7BDFCCE" blockId="104.[181,1165,758,939]" box="[181,429,758,779]" pageId="104" pageNumber="103">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF39BF35A5C9FCB3" blockId="104.[181,1165,758,939]" pageId="104" pageNumber="103">
Type material.
<materialsCitation id="3B7B3CECFFBDFFCFFE80BF34A73CFC85" collectingDate="1996-01-24" collectingDateMax="1996-02-27" collectingDateMin="1996-01-24" collectionCode="ACG" collectorName="J. S. Noyes" country="Costa Rica" elevation="700" location="Pitilla" pageId="104" pageNumber="103" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FFBDFFCFFE80BF34A7A6FCE3" box="[354,438,784,806]" pageId="104" pageNumber="103" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FFBDFFCFFE3ABF34A472FCE3" bold="true" box="[472,610,784,806]" pageId="104" pageNumber="103">
<collectingCountry id="F3047621FFBDFFCFFE3ABF34A472FCE3" box="[472,610,784,806]" name="Costa Rica" pageId="104" pageNumber="103">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FFBDFFCFFD8EBF34A4CDFCE3" bold="true" box="[620,733,784,806]" pageId="104" pageNumber="103">
<collectingRegion id="49D7F853FFBDFFCFFD8EBF34A4CDFCE3" box="[620,733,784,806]" country="Costa Rica" name="Guanacaste" pageId="104" pageNumber="103">Guanacaste</collectingRegion>
</emphasis>
,
<location id="8ECC606AFFBDFFCFFD0ABF34A50EFCE3" LSID="urn:lsid:plazi:treatment:03BA87A7FFB3FFCFFE1FB957A4CBFC24:8ECC606AFFBDFFCFFD0ABF34A50EFCE3" box="[744,798,784,806]" country="Costa Rica" name="Pitilla" pageId="104" pageNumber="103" stateProvince="Guanacaste">Pitilla</location>
(
<collectionCode id="ED02AE74FFBDFFCFFCCBBF34A54DFCE3" box="[809,861,784,806]" pageId="104" pageNumber="103">ACG</collectionCode>
),
<quantity id="4CEB9B54FFBDFFCFFC8EBF34A5B1FCE0" box="[876,929,784,805]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="104" pageNumber="103" unit="m" value="700.0">
<elevation id="003ED182FFBDFFCFFC8EBF34A5B1FCE0" box="[876,929,784,805]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="104" pageNumber="103" unit="m" value="700.0">700m</elevation>
</quantity>
, MT/YPT,
<date id="FFAD1071FFBDFFCFFBE4BF34A29CFCE3" box="[1030,1164,784,806]" pageId="104" pageNumber="103" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">
<collectingDate id="EFE9E999FFBDFFCFFBE4BF34A29CFCE3" box="[1030,1164,784,806]" pageId="104" pageNumber="103" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">24.i-27.ii.1996</collectingDate>
</date>
(
<collectorName id="26E65367FFBDFFCFFF5FBF0FA736FC85" box="[189,294,811,832]" pageId="104" pageNumber="103">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFFBDFFCFFEDBBF0FA423FCB3" collectingDate="2003-02-19" collectingDateMax="2003-02-20" collectingDateMin="2003-02-19" collectionCode="LN" collectorName="J. S. Noyes &amp; Alajuela &amp; PN Arenal &amp; Sendero Mena &amp; Sect. Catarata &amp; G. Carballo" country="Costa Rica" latitude="10.883333" location="Reserva Rincon Forestal" longLatPrecision="1296" longitude="-85.3" pageId="104" pageNumber="103" specimenCode="LN 454170" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="54A88813FFBDFFCFFEDBBF0FA785FC85" box="[313,405,811,832]" pageId="104" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FFBDFFCFFE43BF0FA420FC84" bold="true" box="[417,560,811,833]" pageId="104" pageNumber="103">
<collectingCountry id="F3047621FFBDFFCFFE43BF0FA420FC84" box="[417,560,811,833]" name="Costa Rica" pageId="104" pageNumber="103">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FFBDFFCFFD8FBF0FA4AFFC84" bold="true" box="[621,703,811,833]" pageId="104" pageNumber="103">
<collectingRegion id="49D7F853FFBDFFCFFD8FBF0FA4AFFC84" box="[621,703,811,833]" country="Costa Rica" name="Alajuela" pageId="104" pageNumber="103">Alajuela</collectingRegion>
</emphasis>
,
<location id="8ECC606AFFBDFFCFFD2CBF0FA5A7FC84" LSID="urn:lsid:plazi:treatment:03BA87A7FFB3FFCFFE1FB957A4CBFC24:8ECC606AFFBDFFCFFD2CBF0FA5A7FC84" box="[718,951,811,833]" country="Costa Rica" latitude="10.883333" longLatPrecision="1296" longitude="-85.3" name="Reserva Rincon Forestal" pageId="104" pageNumber="103" stateProvince="Alajuela">Reserva Rincon Forestal</location>
, Est. Caribe,
<geoCoordinate id="EE275076FFBDFFCFFBDCBF0FA29CFC84" box="[1086,1164,811,833]" degrees="10" direction="north" minutes="53" orientation="latitude" pageId="104" pageNumber="103" precision="925" value="10.883333">10°53N</geoCoordinate>
<geoCoordinate id="EE275076FFBDFFCFFF57BF61A718FC9E" box="[181,264,837,859]" degrees="85" direction="west" minutes="18" orientation="longitude" pageId="104" pageNumber="103" precision="925" value="-85.3">85°18W</geoCoordinate>
,
<date id="FFAD1071FFBDFFCFFEF1BF61A79DFC9E" box="[275,397,837,859]" pageId="104" pageNumber="103" value="2003-02-19" valueMax="2003-02-20" valueMin="2003-02-19">
<collectingDate id="EFE9E999FFBDFFCFFEF1BF61A79DFC9E" box="[275,397,837,859]" pageId="104" pageNumber="103" value="2003-02-19" valueMax="2003-02-20" valueMin="2003-02-19">19-20.ii.2003</collectingDate>
</date>
(
<collectorName id="26E65367FFBDFFCFFE79BF62A7EFFC9E" box="[411,511,838,859]" pageId="104" pageNumber="103">J.S. Noyes</collectorName>
); 1E,
<collectorName id="26E65367FFBDFFCFFDD5BF61A496FC9E" box="[567,646,837,859]" pageId="104" pageNumber="103">Alajuela</collectorName>
, [
<collectorName id="26E65367FFBDFFCFFD76BF62A4EBFC98" box="[660,763,838,861]" pageId="104" pageNumber="103">PN Arenal</collectorName>
,]
<collectorName id="26E65367FFBDFFCFFCE8BF61A580FC9E" box="[778,912,837,859]" pageId="104" pageNumber="103">Sendero Mena</collectorName>
,
<collectorName id="26E65367FFBDFFCFFC78BF62A20AFC9E" box="[922,1050,838,859]" pageId="104" pageNumber="103">Sect. Catarata</collectorName>
,
<specimenCode id="DBB59ECAFFBDFFCFFBC6BF62A29CFC9E" box="[1060,1164,837,859]" collectionCode="LN" pageId="104" pageNumber="103">LN 454170</specimenCode>
271800, #62078, iii-iv.2001 (
<collectorName id="26E65367FFBDFFCFFE5DBF44A43DFCB3" box="[447,557,864,886]" pageId="104" pageNumber="103">G. Carballo</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFFBDFFCFFDDEBF44A5C6FCB3" box="[572,982,864,886]" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="104" pageNumber="103" specimenCount="2" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="54A88813FFBDFFCFFDDEBF44A480FCB3" box="[572,656,864,886]" pageId="104" pageNumber="103" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FFBDFFCFFD4FBF44A510FCB0" box="[685,768,864,885]" pageId="104" pageNumber="103">MZUCR</collectionCode>
,
<typeStatus id="54A88813FFBDFFCFFCEBBF45A570FCB3" box="[777,864,865,886]" pageId="104" pageNumber="103" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FFBDFFCFFC9FBF45A5C6FCB3" box="[893,982,865,886]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="104" pageNumber="103" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36B1FFBDFFCFFF39BF5FA47BFC6E" blockId="104.[181,1165,758,939]" pageId="104" pageNumber="103">
<materialsCitation id="3B7B3CECFFBDFFCFFF39BF5FA477FC6E" collectingDate="1994-10-04" collectionCode="USNM" collectorName="T. L. Erwin &amp; Material" country="Ecuador" elevation="220" latitude="-0.65" location="Non" longLatPrecision="1308" longitude="-76.433334" pageId="104" pageNumber="103" specimenCount="1">
<location id="8ECC606AFFBDFFCFFF39BF5FA712FC55" LSID="urn:lsid:plazi:treatment:03BA87A7FFB3FFCFFE1FB957A4CBFC24:8ECC606AFFBDFFCFFF39BF5FA712FC55" box="[219,258,891,912]" country="Ecuador" latitude="-0.65" longLatPrecision="1308" longitude="-76.433334" name="Non" pageId="104" pageNumber="103">Non</location>
type material.
<emphasis id="B967EAA3FFBDFFCFFE68BF5FA7EDFC54" bold="true" box="[394,509,891,913]" pageId="104" pageNumber="103">
<collectingCountry id="F3047621FFBDFFCFFE68BF5FA7EDFC54" box="[394,509,891,913]" name="Ecuador" pageId="104" pageNumber="103">ECUADOR</collectingCountry>
</emphasis>
, 1E, Res. Eth. Waorani,
<geoCoordinate id="EE275076FFBDFFCFFD3DBF5FA53AFC55" box="[735,810,891,912]" degrees="00" direction="south" minutes="39" orientation="latitude" pageId="104" pageNumber="103" precision="925" value="-0.65">00°39S</geoCoordinate>
<geoCoordinate id="EE275076FFBDFFCFFCD2BF5FA593FC54" box="[816,899,891,913]" degrees="76" direction="west" minutes="26" orientation="longitude" pageId="104" pageNumber="103" precision="925" value="-76.433334">76°26W</geoCoordinate>
,
<quantity id="4CEB9B54FFBDFFCFFC6EBF5FA5D2FC55" box="[908,962,891,912]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="104" pageNumber="103" unit="m" value="220.0">
<elevation id="003ED182FFBDFFCFFC6EBF5FA5D2FC55" box="[908,962,891,912]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="104" pageNumber="103" unit="m" value="220.0">220m</elevation>
</quantity>
, canopy fog, lot #866,
<date id="FFAD1071FFBDFFCFFF57BFB2A715FC6E" box="[181,261,918,939]" pageId="104" pageNumber="103" value="1994-10-04">
<collectingDate id="EFE9E999FFBDFFCFFF57BFB2A715FC6E" box="[181,261,918,939]" pageId="104" pageNumber="103" value="1994-10-04">4.x.1994</collectingDate>
</date>
(
<collectorName id="26E65367FFBDFFCFFEF3BFB2A764FC6E" box="[273,372,917,939]" pageId="104" pageNumber="103">T.L. Erwin</collectorName>
et al.).
<collectorName id="26E65367FFBDFFCFFE5BBFB1A416FC6E" box="[441,518,917,939]" pageId="104" pageNumber="103">Material</collectorName>
in
<collectionCode id="ED02AE74FFBDFFCFFDC1BFB2A477FC6E" box="[547,615,918,939]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="104" pageNumber="103" type="Museum">USNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFFBDFFCFFF57BFEDA4CBFC24" box="[181,731,969,993]" pageId="104" pageNumber="103" type="discussion">
<paragraph id="8BAC36B1FFBDFFCFFF57BFEDA4CBFC24" blockId="104.[181,731,969,993]" box="[181,731,969,993]" pageId="104" pageNumber="103">
COMMENTS. See comments under
<taxonomicName id="4C134D32FFBDFFCFFDC9BFEDA49AFC24" authorityName="Noyes" authorityYear="2023" box="[555,650,969,993]" class="Insecta" family="Encyrtidae" genus="Coccidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="103" phylum="Arthropoda" rank="species" species="umbro">
<emphasis id="B967EAA3FFBDFFCFFDC9BFEDA49AFC24" box="[555,650,969,993]" italics="true" pageId="104" pageNumber="103">C. umbro</emphasis>
</taxonomicName>
(p. 99).
</paragraph>
</subSubSection>
</treatment>
</document>