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<document id="513E26C19092270E02C305965143D4AF" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF08FF78FDF2B9FAA50AFC7C" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus carita Noyes 2023, sp.nov." docType="treatment" docUuid="D9AD837B-B5DF-4DB5-9690-36009296248C" docUuidSource="ZooBank" docVersion="2" lastPageNumber="222" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="220" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="374FBACB17AB69074625367451F7FA9C">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="2A49F1890757817D6BAC81DF7B75B92C">Noyes, John Stuart</mods:namePart>
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<mods:date id="65F994A58527FCC046C4FB5B3BE5AC74">2023</mods:date>
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<subSubSection id="C309653AFF08FF7AFDF2B9FAA522FA32" box="[528,818,1502,1527]" pageId="221" pageNumber="220" type="nomenclature">
<paragraph id="8BAC36B1FF08FF7AFDF2B9FAA522FA32" blockId="221.[528,818,1502,1553]" box="[528,818,1502,1527]" pageId="221" pageNumber="220">
<heading id="D0E481DDFF08FF7AFDF2B9FAA522FA32" bold="true" box="[528,818,1502,1527]" centered="true" fontSize="10" level="2" pageId="221" pageNumber="220" reason="2">
<emphasis id="B967EAA3FF08FF7AFDF2B9FAA522FA32" bold="true" box="[528,818,1502,1527]" pageId="221" pageNumber="220">
<taxonomicName id="4C134D32FF08FF7AFDF2B9FAA4F1FA33" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[528,737,1502,1526]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="221" pageNumber="220" phylum="Arthropoda" rank="species" species="carita" status="sp.nov.">
<emphasis id="B967EAA3FF08FF7AFDF2B9FAA4F1FA33" bold="true" box="[528,737,1502,1526]" italics="true" pageId="221" pageNumber="220">Cheiloneurus carita</emphasis>
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<taxonomicNameLabel id="A25457D8FF08FF7AFD05B9FBA522FA32" box="[743,818,1503,1527]" pageId="221" pageNumber="220" rank="species">sp.nov.</taxonomicNameLabel>
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<subSubSection id="C309653AFF08FF7AFDFBB9DDA539F9D4" box="[537,809,1529,1553]" pageId="221" pageNumber="220" type="description">
<paragraph id="8BAC36B1FF08FF7AFDFBB9DDA539F9D4" blockId="221.[528,818,1502,1553]" box="[537,809,1529,1553]" pageId="221" pageNumber="220">
(
<figureCitation id="13282A34FF08FF7AFDC2B9DDA4BCF9D4" box="[544,684,1529,1553]" captionStart-0="Figs 524-537" captionStart-1="Figs 538-552" captionStartId-0="774.[181,226,1526,1550]" captionStartId-1="775.[181,226,1526,1550]" captionTargetBox-0="[185,1163,226,1505]" captionTargetBox-1="[181,1159,229,1508]" captionTargetId-0="figure-9@774.[181,1163,223,1508]" captionTargetId-1="figure-10@775.[181,1163,223,1508]" captionTargetPageId-0="774" captionTargetPageId-1="775" captionText-0="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." captionText-1="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="221" pageNumber="220">Figs 533-539</figureCitation>
; Hab. E 83)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF08FF78FF57BA0BA771FD25" lastPageId="223" lastPageNumber="222" pageId="221" pageNumber="220" type="diagnosis">
<paragraph id="8BAC36B1FF08FF7AFF57BA0BA29DF953" blockId="221.[181,1165,1582,1687]" pageId="221" pageNumber="220">
DIAGNOSIS.
<emphasis id="B967EAA3FF08FF7AFEACBA0AA78AF983" box="[334,410,1582,1606]" italics="true" pageId="221" pageNumber="220">Female</emphasis>
(overall length about
<quantity id="4CEB9B54FF08FF7AFD6BBA0BA4C5F982" box="[649,725,1583,1607]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="221" pageNumber="220" unit="mm" value="0.9">0.9mm</quantity>
): body generally dark brown mostly matt but with weakly metallic areas; mesoscutum with uniform brown setae; fore wing (
<figureCitation id="13282A34FF08FF7AFBC9BA6DA295F9A4" box="[1067,1157,1609,1633]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="221" pageNumber="220">Fig. 535</figureCitation>
) hyaline; fore coxa white; head (
<figureCitation id="13282A34FF08FF7AFE1BBA40A45FF9B9" box="[505,591,1636,1660]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="221" pageNumber="220">Fig. 537</figureCitation>
) about 3.2X as wide as frontovertex; antenna (
<figureCitation id="13282A34FF08FF7AFBCCBA40A295F9B9" box="[1070,1157,1636,1660]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="221" pageNumber="220">Fig. 534</figureCitation>
) with pedicel nearly as long as F1-F3 combined; F1 and F6 very slightly longer than broad, F2-F5
</paragraph>
<paragraph id="8BAC36B1FF0BFF79FBBCBCA6A29CFF5A" blockId="222.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="222" pageNumber="221">221</paragraph>
<paragraph id="8BAC36B1FF0BFF79FF57BCFAA4F3FE09" blockId="222.[181,1165,222,460]" pageId="222" pageNumber="221">
quadrate or transverse; linear sensilla on F6; clava slightly longer than funicle, apex narrowly rounded; mandible with 3 teeth, upper tooth narrowly truncate; mesoscutum (
<figureCitation id="13282A34FF0BFF79FC10BCDDA25DFED4" box="[1010,1101,249,273]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="222" pageNumber="221">Fig. 539</figureCitation>
) with longitudinally elongate, lineolate sculpture, some imbricate to polygonally reticulate sculpture near posterior margin; scutellum with longitudinally elongate, lineolate sculpture, some enclosed cells near axillae and medially; scutellum without a tuft of setae medially; fore wing (
<figureCitation id="13282A34FF0BFF79FBCFBD6DA295FEA4" box="[1069,1157,329,353]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="222" pageNumber="221">Fig. 535</figureCitation>
) about 2.5X as long as broad; parastigma hardly swollen, not downcurved; stigmal vein (
<figureCitation id="13282A34FF0BFF79FBCCBD40A295FEB9" box="[1070,1157,356,380]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="222" pageNumber="221">Fig. 533</figureCitation>
) with 3 apical campaniform sensilla; gaster with “glands” on Gt1 and Gt5, those on Gt1 separated by slightly more than 2X their own lengths, those on Gt5 separated by about 0.25X their own lengths; syntergum longer than broad.
<emphasis id="B967EAA3FF0BFF79FDDEBD90A460FE09" box="[572,624,436,460]" italics="true" pageId="222" pageNumber="221">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF0BFF79FF57BDCDA293FDC4" blockId="222.[181,1165,489,1687]" box="[181,1155,489,513]" pageId="222" pageNumber="221">
<materialsCitation id="3B7B3CECFF0BFF79FF57BDCDA26EFDC4" box="[181,1150,489,513]" collectionCode="CPD" pageId="222" pageNumber="221" specimenCount="1" typeStatus="holotype">
Female
<emphasis id="B967EAA3FF0BFF79FEE4BDCDA71EFDC4" bold="true" box="[262,270,489,513]" pageId="222" pageNumber="221">(</emphasis>
<typeStatus id="54A88813FF0BFF79FEEDBDCDA778FDC4" box="[271,360,489,513]" pageId="222" pageNumber="221">holotype</typeStatus>
<emphasis id="B967EAA3FF0BFF79FE85BDCDA77FFDC4" bold="true" box="[359,367,489,513]" pageId="222" pageNumber="221">)</emphasis>
: length, including ovipositor,
<quantity id="4CEB9B54FF0BFF79FD40BDCDA4E8FDC4" box="[674,760,489,513]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="222" pageNumber="221" unit="mm" value="0.94">0.94mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF0BFF79FC3DBDCDA223FDC4" box="[991,1075,489,513]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="222" pageNumber="221" unit="mm" value="0.9">0.90mm</quantity>
(
<collectionCode id="ED02AE74FF0BFF79FBA3BDCDA268FDC4" box="[1089,1144,489,513]" pageId="222" pageNumber="221">CPD</collectionCode>
)
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.
</paragraph>
<paragraph id="8BAC36B1FF0BFF79FF39BE20A24EFB59" blockId="222.[181,1165,489,1687]" pageId="222" pageNumber="221">Head dark brown to black with a weak metallic lustre; frontovertex coppery purple behind posterior ocelli, occipital margin medially metallic very dark blue, area around anterior ocellus metallic dark blue, area above scrobes coppery and purple; temple and gena largely metallic dark blue, posteriorly broadly coppery, eye margin narrowly coppery; scrobal area coppery purple, area between eye and scrobe metallic dark blue, interantennal prominence metallic blue between lower parts of toruli, mouth margin coppery; a line of translucent, silvery setae along inner eye margin continuing below eye to gena and along position of malar sulcus; radicle pale orange; scape dark brown with apical one-quarter white; pedicel dark brown, apical one-third white; funicle with F1-F5 very slightly dusky, very pale orange, F6 very pale yellow; clava pale brown; thorax dark brown to black; pronotum with a slight coppery sheen; mesoscutum fairly matt, but slightly silky in appearance with a very weak coppery hue, posterior margin slightly more coppery; axilla with a slight coppery purple sheen; scutellum generally similar in appearance to mesoscutum but extreme apex and side with a fairly strong metallic dark green sheen; dorsum of thorax mostly with dark brown setae but posterior half of mesoscutum clothed in sparse, conspicuous, silvery setae; tegula dark brown with a slight coppery sheen; mesopleuron with a moderate coppery purple sheen; fore coxa white, mid and hind coxa dark brown; fore femur mostly dark brown, with apices white, tibia dark brown with apices white, tarsus pale testaceous; mid femur white with a broad, subapical brown band in distal half, tibia white with a broad, subbasal brown band in basal two-fifths, spur and tarsus very pale yellow; hind femur dark brown, extreme apex white, tibia mostly dark brown with extreme base and apical one-third white, tarsus white with apical tarsomere brown; wings hyaline, venation pale yellow, marginal vein dusky orange; propodeum dark brown with a weak brassy and purple sheen, side with a strong metallic blue-green sheen, outside spiracle with only 1 or 2 silvery setae; gaster dark purple-brown, dorsally with a coppery purple sheen mixed slightly brassy and green, Gt1 with a distinct dark metallic dark blue sheen, side and venter with coppery, blue, brassy and green reflections; gonostylus dark brown, extreme apex pale orange-brown.</paragraph>
<paragraph id="8BAC36B1FF0BFF79FF39B8BBA79EF952" blockId="222.[181,1165,489,1687]" pageId="222" pageNumber="221">
Head (
<figureCitation id="13282A34FF0BFF79FEC2B8BBA767FB72" box="[288,375,1183,1207]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="222" pageNumber="221">Fig. 537</figureCitation>
) about 3.2X as wide as frontovertex, in profile about 1.35X as high as deep, tangent to frontovertex above scrobes forming an angle of about 90° with face below top of scrobes; a small, broadly oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 60°; frontovertex narrowest about level with posterior ocelli, relatively dull, with relatively deep, polygonally reticulate sculpture of mesh size mainly subequal eye facet, deepest and more polygonally reticulate in ocellar area; temple and gena with slightly shallower, longitudinally elongate imbricate-reticulate to polygonally reticulate sculpture; scrobes with relatively shallow, irregular, elongate sculpture; interantennal prominence and mouth margin with shallow, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, narrowly ∩-shaped; interantennal prominence dorsally broadly rounded; antenna as in
<figureCitation id="13282A34FF0BFF79FC0BB9ABA255FA62" box="[1001,1093,1423,1447]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="222" pageNumber="221">Fig. 534</figureCitation>
; scape hardly broadened and flattened, about 4X as long as broad; funicle with F1-F5 smaller, subequal, F1 very slightly longer than broad, F2-F5 slightly transverse, F6 clearly largest and broadest but slightly longer than broad; linear sensilla only on F6; clava slightly longer than pedicel and funicle combined, sutures parallel, sensory area small, at apex only giving it a pointed or slightly rounded appearance; eye not quite reaching occipital margin, separated by less than diameter of facet, more or less naked, clothed with extremely inconspicuous setae, each much less than 0.5X diameter of facet; malar sulcus absent; mandible tridentate, upper tooth narrowly truncate. Relative measurements: HW 61,
<collectingRegion id="49D7F853FF0BFF79FDE6BA41A439F9B9" box="[516,553,1637,1660]" country="Swaziland" name="Hhohho" pageId="222" pageNumber="221">HH</collectingRegion>
52, FV 19, POL 8, OOL 1, OCL 6, AOL 7.5, EL 34, EW 28, MS 25, SL 24, SW 6.
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF57BCA6A6F3FF5A" blockId="223.[181,227,130,159]" box="[181,227,130,159]" pageId="223" pageNumber="222">
<pageNumber id="BBB83D67FF0AFF78FF57BCA6A6F3FF5A" ambiguity="8" box="[181,227,130,159]" fuzzyness="0" pageId="223" pageNumber="222" score="12" value="222">
<pageTitle id="CB8CEED6FF0AFF78FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="223" pageNumber="222">222</pageTitle>
</pageNumber>
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FEB5BCA4A5FBFF5D" blockId="223.[343,1003,128,152]" box="[343,1003,128,152]" pageId="223" pageNumber="222">
<pageTitle id="CB8CEED6FF0AFF78FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="223" pageNumber="222">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF0AFF78FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="223" pageNumber="222" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF39BCFAA7C0FE2E" blockId="223.[181,1165,222,682]" pageId="223" pageNumber="222">
Thorax (
<figureCitation id="13282A34FF0AFF78FED6BCFAA79CFF33" box="[308,396,222,246]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="223" pageNumber="222">Fig. 539</figureCitation>
) with imbricate-reticulate to polygonally reticulate sculpture on pronotum that is about as deep as that in ocellar area, posterior margin quite smooth; mesoscutum throughout with fine, longitudinally elongate, striate-reticulate sculpture that is about as deep as sculpture on pronotum, anteriorly much finer and of narrower cells than posterior one-fifth; scutellum with similar, but much finer, slightly longitudinally elongate, polygonally reticulate sculpture, extreme side and apex completely smooth; setae on mesoscutum fairly sparse but quite evenly distributed; scutellum with a weak subapical tuft of only 4 or 5 setae; visible part of mesoscutum about 1.7X as broad as long; scutellum about 1.2X as broad as long; fore wing with venation and setation as in
<figureCitation id="13282A34FF0AFF78FF57BD9CA752FE15" box="[181,322,440,464]" captionStart="Figs 524-537" captionStartId="774.[181,226,1526,1550]" captionTargetBox="[185,1163,226,1505]" captionTargetId="figure-9@774.[181,1163,223,1508]" captionTargetPageId="774" captionText="Figs 524-537. Cheiloneurus argentipes (524) ovipositor, variant; (525) ovipositor, variant. Cheiloneurus sophocles, HT (526) forewing; (527) apex of fore wing venation; (528) antenna; (529) hypopygium; (530) mesosoma, dorsal view; (531) ovipositor; (532) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus carita, HT (533) apex of fore wing venation; (534) antenna; (535) fore wing; (536) hypopygium; (537) head, facial view." pageId="223" pageNumber="222">Figs 533, 535</figureCitation>
; costal cell with 5 or 6 dorsal setae at apex. Relative measurements: FWL 62, FWW 24.5; HWL 45, HWW 10.5.
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF39BDCBA755FD4A" blockId="223.[181,1165,222,682]" pageId="223" pageNumber="222">
Gaster with hypopygium reaching about 0.6X; syntergum about 1.4X as long as broad and 1.2X as long as mid tibia, with apex acute; “glands” (
<figureCitation id="13282A34FF0AFF78FD34BE2EA53DFDE7" box="[726,813,522,546]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="223" pageNumber="222">Fig. 551</figureCitation>
) present on Gt1 and Gt5, those on Gt1 separated by slightly more than 2X their own lengths, those on Gt5 separated by about 0.25X their own lengths; hypopygium
<figureCitation id="13282A34FF0AFF78FE1ABE64A440FD9D" box="[504,592,576,600]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="223" pageNumber="222">Fig. 549</figureCitation>
; ovipositor (
<figureCitation id="13282A34FF0AFF78FD2CBE64A534FD9D" box="[718,804,576,600]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="223" pageNumber="222">Fig. 538</figureCitation>
) slightly exserted, the exserted part about 0.5X length of mid tibial spur or about 0.18X mid tibia. Relative measurements: OL 80, GL 18.5 [MT 41].
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF39BEB6A457FD6F" blockId="223.[181,1165,222,682]" box="[219,583,658,682]" pageId="223" pageNumber="222">
Variation. Only
<typeStatus id="54A88813FF0AFF78FE9CBEB6A7C7FD6F" box="[382,471,658,682]" pageId="223" pageNumber="222">holotype</typeStatus>
examined.
</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF57BEECA771FD25" blockId="223.[181,353,712,736]" box="[181,353,712,736]" pageId="223" pageNumber="222">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0AFF78FF57BEDBA7F9FCD3" box="[181,489,766,791]" pageId="223" pageNumber="222" type="distribution">
<paragraph id="8BAC36B1FF0AFF78FF57BEDBA7F9FCD3" blockId="223.[181,489,766,791]" box="[181,489,766,791]" pageId="223" pageNumber="222">
DISTRIBUTION.
<collectingCountry id="F3047621FF0AFF78FE91BEDBA7F5FCD3" box="[371,485,766,791]" name="Costa Rica" pageId="223" pageNumber="222">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0AFF78FF57BF11A76EFC88" box="[181,382,821,845]" pageId="223" pageNumber="222" type="biology_ecology">
<paragraph id="8BAC36B1FF0AFF78FF57BF11A76EFC88" blockId="223.[181,382,821,845]" box="[181,382,821,845]" pageId="223" pageNumber="222">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0AFF78FF57BF49A50AFC7C" pageId="223" pageNumber="222" type="materials_examined">
<paragraph id="8BAC36B1FF0AFF78FF57BF49A7BDFC47" blockId="223.[181,1164,877,953]" box="[181,429,877,898]" pageId="223" pageNumber="222">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF0AFF78FF39BFADA50AFC7C" blockId="223.[181,1164,877,953]" pageId="223" pageNumber="222">
Type material.
<materialsCitation id="3B7B3CECFF0AFF78FE81BFACA453FC7C" collectingDate="2016-02-15" collectingDateMax="2016-02-17" collectingDateMin="2016-02-15" collectorName="J. S. Noyes" country="Costa Rica" elevation="770" latitude="10.8" location="Guanacaste" longLatPrecision="1297" longitude="-85.4" pageId="223" pageNumber="222" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF0AFF78FE81BFACA7A7FC5B" box="[355,439,904,926]" pageId="223" pageNumber="222">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF0AFF78FE38BFACA474FC5B" bold="true" box="[474,612,904,926]" pageId="223" pageNumber="222">
<collectingCountry id="F3047621FF0AFF78FE38BFACA474FC5B" box="[474,612,904,926]" name="Costa Rica" pageId="223" pageNumber="222">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF0AFF78FD8DBFACA4F0FC5B" bold="true" box="[623,736,904,926]" pageId="223" pageNumber="222">
<collectingRegion id="49D7F853FF0AFF78FD8DBFACA4F0FC5B" box="[623,736,904,926]" country="Costa Rica" name="Guanacaste" pageId="223" pageNumber="222">Guanacaste</collectingRegion>
</emphasis>
,
<quantity id="4CEB9B54FF0AFF78FD09BFACA50FFC5B" box="[747,799,904,926]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="223" pageNumber="222" unit="km" value="18.0">18km</quantity>
<taxonomicName id="4C134D32FF0AFF78FCC6BFADA225FC5B" authority=", Buena Vista Lodge" authorityName="Buena Vista Lodge" box="[804,1077,904,926]" class="Insecta" family="Encyrtidae" genus="Neococcidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="223" pageNumber="222" phylum="Arthropoda" rank="species" species="liberia">N Liberia, Buena Vista Lodge</taxonomicName>
,
<geoCoordinate id="EE275076FF0AFF78FBDCBFACA29CFC58" box="[1086,1164,904,925]" degrees="10" direction="north" minutes="48" orientation="latitude" pageId="223" pageNumber="222" precision="925" value="10.8">10°48N</geoCoordinate>
<geoCoordinate id="EE275076FF0AFF78FF57BF87A718FC7C" box="[181,264,931,953]" degrees="85" direction="west" minutes="24" orientation="longitude" pageId="223" pageNumber="222" precision="925" value="-85.4">85°24W</geoCoordinate>
,
<quantity id="4CEB9B54FF0AFF78FEF0BF80A757FC7C" box="[274,327,932,953]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="223" pageNumber="222" unit="m" value="770.0">
<elevation id="003ED182FF0AFF78FEF0BF80A757FC7C" box="[274,327,932,953]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="223" pageNumber="222" unit="m" value="770.0">770m</elevation>
</quantity>
,
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<collectingDate id="EFE9E999FF0AFF78FEB3BF87A7DCFC7C" box="[337,460,931,953]" pageId="223" pageNumber="222" value="2016-02-15" valueMax="2016-02-17" valueMin="2016-02-15">15-17.ii.2016</collectingDate>
</date>
(
<collectorName id="26E65367FF0AFF78FE3BBF80A42DFC7C" box="[473,573,932,953]" pageId="223" pageNumber="222">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF0AFF78FDAEBF87A506FC7C" box="[588,790,931,953]" collectionCode="NHMUK" country="Costa Rica" pageId="223" pageNumber="222" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF0AFF78FDAEBF87A4B0FC7C" box="[588,672,931,953]" pageId="223" pageNumber="222">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF0AFF78FD5FBF80A506FC7C" box="[701,790,932,953]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="223" pageNumber="222" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>