192 lines
20 KiB
XML
192 lines
20 KiB
XML
<document id="DC0C99E7732B39FE6089532B082DDDC3" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FDACFDDDFE31BA35A26DFBEB" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Caldencyrtus acamas Noyes & Hanson 1996" docType="treatment" docVersion="2" lastPageNumber="633" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="632" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="9A423F8D4EB7299A5CF1E43D96018BC2">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="AE2446C811E6FD44CF3DE329CA0826EF">Noyes, John Stuart</mods:namePart>
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<mods:title id="4C314D5AB7071765BBD109509071D565">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="8A321840C902065A457AA437CAC54FAC">2023</mods:date>
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<mods:title id="3786E18893A38D4B5C2098613DF2D2A4">Oxford, England</mods:title>
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<mods:number id="81BBEC443EC7C44DFCBE5FB4F641638E">2023-06-30</mods:number>
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<mods:identifier id="00CE894FD95DFDCAD01FE75697A30B26" type="DOI">10.5281/zenodo.8074943</mods:identifier>
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<mods:identifier id="9724032A331943771DC4FF65C52579F4" type="ISSN">2754-9844</mods:identifier>
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<mods:identifier id="F283C6AFCF7AEED017DD3348797FDF2B" type="ZooBank">BCAD06E8-0AFE-46ED-B7FA-930983CD44C4</mods:identifier>
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</mods:mods>
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<treatment id="03BA87A7FDACFDDDFE31BA35A26DFBEB" LSID="urn:lsid:plazi:treatment:03BA87A7FDACFDDDFE31BA35A26DFBEB" httpUri="http://treatment.plazi.org/id/03BA87A7FDACFDDDFE31BA35A26DFBEB" lastPageId="634" lastPageNumber="633" pageId="633" pageNumber="632">
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<subSubSection id="C309653AFDACFDDEFE31BA35A571F981" pageId="633" pageNumber="632" type="nomenclature">
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<paragraph id="8BAC36B1FDACFDDEFE31BA35A57EF9EC" blockId="633.[467,878,1553,1604]" box="[467,878,1553,1577]" pageId="633" pageNumber="632">
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<heading id="D0E481DDFDACFDDEFE31BA35A57EF9EC" bold="true" box="[467,878,1553,1577]" centered="true" fontSize="10" level="2" pageId="633" pageNumber="632" reason="2">
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<taxonomicName id="4C134D32FDACFDDEFE31BA35A57EF9EC" ID-CoL="PHT8" authority="Noyes & Hanson" authorityName="Noyes & Hanson" authorityYear="1996" box="[467,878,1553,1577]" class="Insecta" family="Encyrtidae" genus="Caldencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="633" pageNumber="632" phylum="Arthropoda" rank="species" species="acamas">
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<emphasis id="B967EAA3FDACFDDEFE31BA35A57EF9EC" bold="true" box="[467,878,1553,1577]" pageId="633" pageNumber="632">
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<emphasis id="B967EAA3FDACFDDEFE31BA35A4A3F9ED" bold="true" box="[467,691,1553,1577]" italics="true" pageId="633" pageNumber="632">Caldencyrtus acamas</emphasis>
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Noyes & Hanson
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BAC36B1FDACFDDEFE02BA08A571F981" blockId="633.[467,878,1553,1604]" box="[480,865,1580,1604]" pageId="633" pageNumber="632">(Figs 1722-1732; Hab. E 253, G 254)</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFDACFDDDFF57BA45A29CFF5A" lastPageId="634" lastPageNumber="633" pageId="633" pageNumber="632" type="reference_group">
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<paragraph id="8BAC36B1FDACFDDEFF57BA45A77BF952" blockId="633.[181,1164,1633,1687]" pageId="633" pageNumber="632">
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<taxonomicName id="4C134D32FDACFDDEFF57BA45A49EF9BC" authority="Noyes & Hanson, 1996" authorityName="Noyes & Hanson" authorityYear="1996" box="[181,654,1633,1657]" class="Insecta" family="Encyrtidae" genus="Caldencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="633" pageNumber="632" phylum="Arthropoda" rank="species" species="acamas">
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<emphasis id="B967EAA3FDACFDDEFF57BA45A781F9BC" box="[181,401,1633,1657]" italics="true" pageId="633" pageNumber="632">Caldencyrtus acamas</emphasis>
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<materialsCitation id="3B7B3CECFDACFDDEFE78BA45A49EF9BC" box="[410,654,1633,1657]" collectionCode="NHMUK" country="Costa Rica" location="Costa Rica" pageId="633" pageNumber="632" specimenCount="1" typeStatus="holotype">
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<bibRefCitation id="EF824B40FDACFDDEFE78BA45A49EF9BC" author="Noyes, J. S. & Hanson, P." box="[410,654,1633,1657]" pageId="633" pageNumber="632" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. & Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes & Hanson, 1996</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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:116,117-118.
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<typeStatus id="54A88813FDACFDDEFCC2BA45A56EF9BC" box="[800,894,1633,1657]" pageId="633" pageNumber="632">Holotype</typeStatus>
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E,
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<collectingCountry id="F3047621FDACFDDEFC4ABA45A20AF9BC" box="[936,1050,1633,1657]" name="Costa Rica" pageId="633" pageNumber="632">Costa Rica</collectingCountry>
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, NHMUK, examined.
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</paragraph>
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<paragraph id="8BAC36B1FDAFFDDDFBBCBCA6A29CFF5A" blockId="634.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="634" pageNumber="633">633</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFDAFFDDDFF57BCFBA401FCC5" pageId="634" pageNumber="633" type="diagnosis">
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<paragraph id="8BAC36B1FDAFFDDDFF57BCFBA401FCC5" blockId="634.[181,1165,222,768]" pageId="634" pageNumber="633">
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DIAGNOSIS.
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<emphasis id="B967EAA3FDAFFDDDFEB2BCFAA78CFF33" box="[336,412,222,246]" italics="true" pageId="634" pageNumber="633">Female</emphasis>
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(length about
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<quantity id="4CEB9B54FDAFFDDDFDDCBCFAA4A2FF33" box="[574,690,222,246]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="634" pageNumber="633" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1-2.5mm</quantity>
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): antenna (Figs 1723-1725) with scape dark brown; scutellum in proximal half relatively dull coppery purple with some weak violet reflections laterally, in apical half contrasting shiny metallic blue-green; fore tibia more or less completely pale orange, mid and hind tibiae pale orange in apical half; eye separated from occipital margin by much less than diameter of facet, occipital margin sharp at nearest point to eye; vertex with a broadly oval, shiny bottomed depression adjacent to eye, slightly smaller than diameter of posterior ocellus and a little behind it; frontovertex (Fig. 1732) with shallow, small, inconspicuous piliferous punctures; antenna (Figs 1723-1725) with pedicel a little more than 2X as long as broad, F1 subequal in length and width to pedicel and about 1.5-2X as long as broad; F6 subquadrate, not more than 1.2X as long as broad; flagellum very slightly shorter than head width; fore wing with venation and distribution of setae as in Figs 1730, 1731; posterolateral angle of propodeum (Fig. 1726) with a short, broadly truncate tooth above hind coxa, area between tooth and spiracle convex not depressed; gaster very slightly shorter than thorax; last tergite about 0.75X as long as mid tibia; hypopygium (Fig. 1727) with depth of posterior invagination about 0.5-0.8X mid tibial spur; ovipositor (Fig,. 1728) about 1.7-1.9X as long as syntergum and about 3.6-3.9X as long as gonostylus.
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<emphasis id="B967EAA3FDAFFDDDFED0BE5EA776FD57" box="[306,358,634,658]" italics="true" pageId="634" pageNumber="633">Male</emphasis>
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(length about
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<quantity id="4CEB9B54FDAFFDDDFE1CBE5EA45AFD57" box="[510,586,634,658]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="634" pageNumber="633" unit="mm" value="1.8">1.8mm</quantity>
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): antenna (Fig. 1729) with F1 at most about 2X as long as broad, as wide as pedicel, with at most 25 longitudinal sensilla, each of which is about 0.25X as long as segment; aedeagus (Fig 1722) with pores separated from apex by about 2X their own diameters, apex acute; setae on each side of phallobase relatively close, separated by not more than 1.5X the diameter of sockets.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFDAFFDDDFF57BF3BA7F9FCF2" box="[181,489,799,823]" pageId="634" pageNumber="633" type="distribution">
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<paragraph id="8BAC36B1FDAFFDDDFF57BF3BA7F9FCF2" blockId="634.[181,489,799,823]" box="[181,489,799,823]" pageId="634" pageNumber="633">
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DISTRIBUTION.
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<collectingCountry id="F3047621FDAFFDDDFE91BF3BA7F5FCF2" box="[371,485,799,823]" name="Costa Rica" pageId="634" pageNumber="633">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFDAFFDDDFF57BF72A6E8FC60" pageId="634" pageNumber="633" type="biology_ecology">
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<paragraph id="8BAC36B1FDAFFDDDFF57BF72A6E8FC60" blockId="634.[181,1164,854,933]" pageId="634" pageNumber="633">
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HOSTS. Reared from galls formed by
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<taxonomicName id="4C134D32FDAFFDDDFDACBF73A4F0FCAB" authority="Forster" authorityName="Forster" box="[590,736,855,878]" class="Insecta" family="Triozidae" genus="Trioza" kingdom="Animalia" order="Hemiptera" pageId="634" pageNumber="633" phylum="Arthropoda" rank="genus">
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<emphasis id="B967EAA3FDAFFDDDFDACBF73A49FFCAB" box="[590,655,855,878]" italics="true" pageId="634" pageNumber="633">Trioza</emphasis>
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Förster
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</taxonomicName>
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<emphasis id="B967EAA3FDAFFDDDFD0ABF72A543FCAB" box="[744,851,854,878]" italics="true" pageId="634" pageNumber="633">magnoliae</emphasis>
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group (
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<taxonomicName id="4C134D32FDAFFDDDFC45BF72A208FCAB" box="[935,1048,854,878]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="634" pageNumber="633" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
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:
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<taxonomicName id="4C134D32FDAFFDDDFBC6BF72A296FCAB" box="[1060,1158,854,878]" class="Insecta" family="Triozidae" kingdom="Animalia" order="Hemiptera" pageId="634" pageNumber="633" phylum="Arthropoda" rank="family">Triozidae</taxonomicName>
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) on fruit of
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<taxonomicName id="4C134D32FDAFFDDDFEC7BF56A4F1FC4F" authority="(W. C. Burger)" baseAuthorityName="W. C. Burger" box="[293,737,881,906]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" order="Laurales" pageId="634" pageNumber="633" phylum="Tracheophyta" rank="species" species="hammelianum">
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<emphasis id="B967EAA3FDAFFDDDFEC7BF56A45AFC4C" box="[293,586,881,905]" italics="true" pageId="634" pageNumber="633">Cinnamomum hammelianum</emphasis>
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(W.C. Burger)
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</taxonomicName>
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(
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<taxonomicName id="4C134D32FDAFFDDDFD0DBF55A55CFC4C" box="[751,844,881,905]" class="Magnoliopsida" kingdom="Plantae" order="Laurales" pageId="634" pageNumber="633" phylum="Tracheophyta" rank="order">Laurales</taxonomicName>
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:
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<taxonomicName id="4C134D32FDAFFDDDFCB4BF56A5D0FC4F" box="[854,960,882,906]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="634" pageNumber="633" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
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) (
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<bibRefCitation id="EF824B40FDAFFDDDFC36BF56A6FCFC60" author="Noyes, J. S. & Hanson, P." pageId="634" pageNumber="633" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. & Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes & Hanson, 1996</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFDAFFDDDFF57BFE2A26DFBEB" pageId="634" pageNumber="633" type="materials_examined">
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<paragraph id="8BAC36B1FDAFFDDDFF57BFE2A7BDFC1E" blockId="634.[181,1165,966,1070]" box="[181,429,966,987]" pageId="634" pageNumber="633">MATERIAL EXAMINED.</paragraph>
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<paragraph id="8BAC36B1FDAFFDDDFF39BFC6A26DFBEB" blockId="634.[181,1165,966,1070]" pageId="634" pageNumber="633">
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Type material.
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<materialsCitation id="3B7B3CECFDAFFDDDFE81BFC5A79AFBD7" collectingDate="1991-09-22" collectorName="P. Hanson" country="Costa Rica" elevation="2500" location="Volcan Poas" pageId="634" pageNumber="633" specimenCount="1" stateProvince="Alajuela" typeStatus="holotype">
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<typeStatus id="54A88813FDAFFDDDFE81BFC5A7A7FC32" box="[355,439,993,1015]" pageId="634" pageNumber="633">Holotype</typeStatus>
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E:
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<emphasis id="B967EAA3FDAFFDDDFE39BFC5A476FC32" bold="true" box="[475,614,993,1015]" pageId="634" pageNumber="633">
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<collectingCountry id="F3047621FDAFFDDDFE39BFC5A476FC32" box="[475,614,993,1015]" name="Costa Rica" pageId="634" pageNumber="633">COSTA RICA</collectingCountry>
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</emphasis>
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,
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<emphasis id="B967EAA3FDAFFDDDFD93BFC5A4D3FC32" bold="true" box="[625,707,993,1015]" pageId="634" pageNumber="633">
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<collectingRegion id="49D7F853FDAFFDDDFD93BFC5A4D3FC32" box="[625,707,993,1015]" country="Costa Rica" name="Alajuela" pageId="634" pageNumber="633">Alajuela</collectingRegion>
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</emphasis>
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,
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<location id="8ECC606AFDAFFDDDFD2FBFC5A52FFC32" LSID="urn:lsid:plazi:treatment:03BA87A7FDACFDDDFE31BA35A26DFBEB:8ECC606AFDAFFDDDFD2FBFC5A52FFC32" box="[717,831,993,1015]" country="Costa Rica" name="Volcan Poas" pageId="634" pageNumber="633" stateProvince="Alajuela">Volcan Poas</location>
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,
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<quantity id="4CEB9B54FDAFFDDDFCAABFC5A599FC32" box="[840,905,993,1015]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="634" pageNumber="633" unit="m" value="2500.0">
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<elevation id="003ED182FDAFFDDDFCAABFC5A599FC32" box="[840,905,993,1015]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="634" pageNumber="633" unit="m" value="2500.0">2500m</elevation>
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</quantity>
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,
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<taxonomicName id="4C134D32FDAFFDDDFC70BFC6A5FFFC32" box="[914,1007,994,1015]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="634" pageNumber="633" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
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psyllid fruit gall,
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<date id="FFAD1071FDAFFDDDFF57BFD9A706FBD6" box="[181,278,1021,1043]" pageId="634" pageNumber="633" value="1991-09-22">
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<collectingDate id="EFE9E999FDAFFDDDFF57BFD9A706FBD6" box="[181,278,1021,1043]" pageId="634" pageNumber="633" value="1991-09-22">22.ix.1991</collectingDate>
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</date>
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(
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<collectorName id="26E65367FDAFFDDDFEC6BFD9A794FBD7" box="[292,388,1021,1042]" pageId="634" pageNumber="633">P. Hanson</collectorName>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B7B3CECFDAFFDDDFE76BFD9A533FBEB" collectingDate="1995-02" collectorName="P. Hanson" country="Costa Rica" elevation="2500" location="Volcan Poas" pageId="634" pageNumber="633" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
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<typeStatus id="54A88813FDAFFDDDFE76BFD9A7E0FBD7" box="[404,496,1021,1042]" pageId="634" pageNumber="633">Paratypes</typeStatus>
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:
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<emphasis id="B967EAA3FDAFFDDDFE1ABFD9A494FBD6" bold="true" box="[504,644,1021,1043]" pageId="634" pageNumber="633">
|
|
<collectingCountry id="F3047621FDAFFDDDFE1ABFD9A494FBD6" box="[504,644,1021,1043]" name="Costa Rica" pageId="634" pageNumber="633">COSTA RICA</collectingCountry>
|
|
</emphasis>
|
|
, 1E, same data as holotype; 3G,
|
|
<emphasis id="B967EAA3FDAFFDDDFC5BBFD9A21BFBD6" bold="true" box="[953,1035,1021,1043]" pageId="634" pageNumber="633">
|
|
<collectingRegion id="49D7F853FDAFFDDDFC5BBFD9A21BFBD6" box="[953,1035,1021,1043]" country="Costa Rica" name="Alajuela" pageId="634" pageNumber="633">Alajuela</collectingRegion>
|
|
</emphasis>
|
|
,
|
|
<location id="8ECC606AFDAFFDDDFBF5BFD9A299FBD7" LSID="urn:lsid:plazi:treatment:03BA87A7FDACFDDDFE31BA35A26DFBEB:8ECC606AFDAFFDDDFBF5BFD9A299FBD7" box="[1047,1161,1021,1043]" country="Costa Rica" name="Volcan Poas" pageId="634" pageNumber="633" stateProvince="Alajuela">Volcan Poas</location>
|
|
, ex triozid fruit gall
|
|
<collectedFrom id="380F754CFDAFFDDDFE84B83DA44AFBE8" box="[358,602,1048,1070]" pageId="634" pageNumber="633">
|
|
on
|
|
<taxonomicName id="4C134D32FDAFFDDDFE60B83CA44AFBE8" box="[386,602,1048,1069]" class="Insecta" family="Cerambycidae" genus="Phoebe" kingdom="Animalia" order="Coleoptera" pageId="634" pageNumber="633" phylum="Arthropoda" rank="species" species="cinnamomifolia">
|
|
<emphasis id="B967EAA3FDAFFDDDFE60B83CA44AFBE8" box="[386,602,1048,1069]" italics="true" pageId="634" pageNumber="633">Phoebe cinnamomifolia</emphasis>
|
|
</taxonomicName>
|
|
</collectedFrom>
|
|
, i/
|
|
<date id="FFAD1071FDAFFDDDFD91B83CA4A2FBEB" box="[627,690,1048,1070]" pageId="634" pageNumber="633" value="1995-02">
|
|
<collectingDate id="EFE9E999FDAFFDDDFD91B83CA4A2FBEB" box="[627,690,1048,1070]" pageId="634" pageNumber="633" value="1995-02">ii.1995</collectingDate>
|
|
</date>
|
|
(
|
|
<collectorName id="26E65367FDAFFDDDFD5CB83CA50DFBEB" box="[702,797,1048,1070]" pageId="634" pageNumber="633">P. Hanson</collectorName>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3B7B3CECFDAFFDDDFCCEB83CA269FBEB" box="[812,1145,1048,1070]" collectionCode="NHMUK" country="Costa Rica" pageId="634" pageNumber="633" specimenCount="2" stateProvince="Alajuela" typeStatus="holotype">
|
|
<typeStatus id="54A88813FDAFFDDDFCCEB83CA590FBEB" box="[812,896,1048,1070]" pageId="634" pageNumber="633">Holotype</typeStatus>
|
|
and
|
|
<typeStatus id="54A88813FDAFFDDDFC4FB83DA214FBEB" box="[941,1028,1049,1070]" pageId="634" pageNumber="633">paratypes</typeStatus>
|
|
in
|
|
<collectionCode id="ED02AE74FDAFFDDDFBC2B83DA269FBEB" box="[1056,1145,1049,1070]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="634" pageNumber="633" type="Museum">NHMUK</collectionCode>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |