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<document id="C56BA6C335EAEDD7E495A28899FF556B" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FD9BFDE8FE30B9DDA56EFB73" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Psyllaephagus creusa Noyes &amp; Hanson 1996" docType="treatment" docVersion="2" lastPageNumber="590" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="589" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="AB32C0DF5BF68C3DE22AFBE9B7642D5C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5E4AFA5D5747E6F08354CB4819107585">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="468240732C3E61593F4521A2DA04E139">Noyes, John Stuart</mods:namePart>
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<mods:typeOfResource id="841456F488824AC50DB7289FAAB5540A">text</mods:typeOfResource>
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<mods:title id="63FD5EE30102BF4BC24B4B8FF491A491">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
</mods:titleInfo>
<mods:part id="75423292186FA40151FA02898B99DDBD">
<mods:date id="B6724294FA1EE63B65E7379314F804E3">2023</mods:date>
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<mods:title id="79945DF9A04CAF618E33A0DF35BDB287">Oxford, England</mods:title>
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<mods:number id="E2049179BC9A7F958C82A76FD73976DD">2023-06-30</mods:number>
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<mods:number id="796B14E092A6055085402CC3F930BF0C">2</mods:number>
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<mods:url id="F3024391E2E288743A647BBF00E4E7B1">http://dx.doi.org/10.5281/zenodo.8074943</mods:url>
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<mods:classification id="B36362D82F1B7BFF8357E533B0DD8D22">journal article</mods:classification>
<mods:identifier id="C41FB2A666FDBBF370024976D7010047" type="DOI">10.5281/zenodo.8074943</mods:identifier>
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</mods:mods>
<treatment id="03BA87A7FD9BFDE8FE30B9DDA56EFB73" LSID="urn:lsid:plazi:treatment:03BA87A7FD9BFDE8FE30B9DDA56EFB73" httpUri="http://treatment.plazi.org/id/03BA87A7FD9BFDE8FE30B9DDA56EFB73" lastPageId="591" lastPageNumber="590" pageId="590" pageNumber="589">
<subSubSection id="C309653AFD9BFDE9FE30B9DDA57BF9E9" pageId="590" pageNumber="589" type="nomenclature">
<paragraph id="8BAC36B1FD9BFDE9FE30B9DDA560F9D4" blockId="590.[466,880,1529,1580]" box="[466,880,1529,1553]" pageId="590" pageNumber="589">
<heading id="D0E481DDFD9BFDE9FE30B9DDA560F9D4" bold="true" box="[466,880,1529,1553]" centered="true" fontSize="10" level="2" pageId="590" pageNumber="589" reason="2">
<taxonomicName id="4C134D32FD9BFDE9FE30B9DDA560F9D4" ID-CoL="4PS25" authority="Noyes &amp; Hanson" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[466,880,1529,1553]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="590" pageNumber="589" phylum="Arthropoda" rank="species" species="creusa">
<emphasis id="B967EAA3FD9BFDE9FE30B9DDA560F9D4" bold="true" box="[466,880,1529,1553]" pageId="590" pageNumber="589">
<emphasis id="B967EAA3FD9BFDE9FE30B9DDA4A4F9D5" bold="true" box="[466,692,1529,1553]" italics="true" pageId="590" pageNumber="589">Psyllaephagus creusa</emphasis>
Noyes &amp; Hanson
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAC36B1FD9BFDE9FE35BA30A57BF9E9" blockId="590.[466,880,1529,1580]" box="[471,875,1556,1580]" pageId="590" pageNumber="589">(Figs 6,1547-1554; Hab. E 229, G 230)</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9BFDE8FF57BA6DA6F3FF5A" lastPageId="591" lastPageNumber="590" pageId="590" pageNumber="589" type="reference_group">
<paragraph id="8BAC36B1FD9BFDE9FF57BA6DA7C4F9B9" blockId="590.[181,1164,1609,1660]" pageId="590" pageNumber="589">
<taxonomicName id="4C134D32FD9BFDE9FF57BA6DA4A1F9A4" authority="Noyes &amp; Hanson, 1996" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[181,689,1609,1634]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="590" pageNumber="589" phylum="Arthropoda" rank="species" species="creusa">
<emphasis id="B967EAA3FD9BFDE9FF57BA6DA78BF9A4" box="[181,411,1609,1633]" italics="true" pageId="590" pageNumber="589">Psyllaephagus creusa</emphasis>
<materialsCitation id="3B7B3CECFD9BFDE9FE4BBA6EA4A1F9A4" box="[425,689,1609,1634]" collectionCode="NHMUK" country="Costa Rica" location="Costa Rica" pageId="590" pageNumber="589" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EF824B40FD9BFDE9FE4BBA6EA4A1F9A4" author="Noyes, J. S. &amp; Hanson, P." box="[425,689,1609,1634]" pageId="590" pageNumber="589" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. &amp; Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes &amp; Hanson, 1996</bibRefCitation>
</materialsCitation>
</taxonomicName>
:126,127,130-131.
<typeStatus id="54A88813FD9BFDE9FC9BBA6DA5C7F9A4" box="[889,983,1609,1633]" pageId="590" pageNumber="589">Holotype</typeStatus>
E,
<collectingCountry id="F3047621FD9BFDE9FBEDBA6DA298F9A4" box="[1039,1160,1609,1633]" name="Costa Rica" pageId="590" pageNumber="589">Costa Rica</collectingCountry>
, NHMUK, examined.
</paragraph>
<paragraph id="8BAC36B1FD9AFDE8FF57BCA6A6F3FF5A" blockId="591.[181,227,130,159]" box="[181,227,130,159]" pageId="591" pageNumber="590">
<pageNumber id="BBB83D67FD9AFDE8FF57BCA6A6F3FF5A" ambiguity="2" box="[181,227,130,159]" fuzzyness="0" pageId="591" pageNumber="590" score="12" value="590">
<pageTitle id="CB8CEED6FD9AFDE8FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="591" pageNumber="590">590</pageTitle>
</pageNumber>
</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9AFDE8FEB5BCA4A43FFD34" pageId="591" pageNumber="590" type="description">
<paragraph id="8BAC36B1FD9AFDE8FEB5BCA4A5FBFF5D" blockId="591.[343,1003,128,152]" box="[343,1003,128,152]" pageId="591" pageNumber="590">
<pageTitle id="CB8CEED6FD9AFDE8FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="591" pageNumber="590">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FD9AFDE8FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FD9AFDE8FF57BCFAA43FFD34" blockId="591.[181,1165,222,753]" pageId="591" pageNumber="590">
<emphasis id="B967EAA3FD9AFDE8FF57BCFAA711FF33" box="[181,257,222,246]" italics="true" pageId="591" pageNumber="590">Female</emphasis>
(length about
<quantity id="4CEB9B54FD9AFDE8FE7BBCFAA41DFF33" box="[409,525,222,246]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="591" pageNumber="590" unit="mm" value="1.9" valueMax="2.3" valueMin="1.5">1.5-2.3mm</quantity>
): head and thorax generally dull metallic green or blue-green; antenna (Fig. 1554) with scape brown; flagellum brown; tegula dark brown; femora and tibiae dark brown; head (Fig. 1547) about 2.8-2.9X as wide as mouth opening; head about 2X frontovertex width and about 2.8-2.9X as wide as mouth; frontovertex with piliferous punctures deep and conspicuous, largest subequal in size to ocellus and all much larger than eye facet; ocelli forming an angle of about 105°, posterior ocellus slightly nearer occipital margin than eye; scrobes deep, but separated from anterior ocellus by at least about 2X length of torulus; antenna (Fig. 1554) with scape about 4X as long as broad, clearly shorter than width of frontovertex and with fairly conspicuous campaniform sensilla on outer surface near apex (Fig. 1554, inset); pedicel about as long as F1, all funicle segments longer than broad, F6 subquadrate, linear sensilla on all segments; clava with apical segment short, almost giving clava a 2-segmented appearance, apex rounded; clypeal margin roundly convex and clearly protuberant; mandible with two teeth and a truncation; mesoscutum and scutellum (Fig. 1553) with similar polygonally reticulate sculpture, perhaps slightly deeper and more elongate on scutellum; fore wing (Fig. 1551) about 2.3X as long as broad, costal cell with only a single line of setae dorsally, marginal vein (Fig. 1552) 1.0-1.5X as long as broad, linea calva partially closed; Gt3 without a distinct, strongly reticulate area anteromedially; hypopygium Fig. 1550) ovipositor shorter than mid tibia and hidden, second valvifer with about 5-7 subapical setae.
<emphasis id="B967EAA3FD9AFDE8FDABBE80A46DFD79" box="[585,637,676,700]" italics="true" pageId="591" pageNumber="590">Male</emphasis>
(length about
<quantity id="4CEB9B54FD9AFDE8FCEDBE80A594FD79" box="[783,900,676,700]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="591" pageNumber="590" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5-2.2mm</quantity>
): generally very similar to female but for slightly wider frontovertex, higher placement of antenna and structure of antenna (Fig. 1549) and genitalia (Fig. 1548).
</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9AFDE8FF57BF2BA4FEFCE2" box="[181,750,783,807]" pageId="591" pageNumber="590" type="distribution">
<paragraph id="8BAC36B1FD9AFDE8FF57BF2BA4FEFCE2" blockId="591.[181,750,783,807]" box="[181,750,783,807]" pageId="591" pageNumber="590">
DISTRIBUTION.
<collectingCountry id="F3047621FD9AFDE8FE91BF2BA7F1FCE2" box="[371,481,783,807]" name="Costa Rica" pageId="591" pageNumber="590">Costa Rica</collectingCountry>
(
<bibRefCitation id="EF824B40FD9AFDE8FE0DBF2BA4F3FCE2" author="Noyes, J. S. &amp; Hanson, P." box="[495,739,783,807]" pageId="591" pageNumber="590" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. &amp; Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes &amp; Hanson, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9AFDE8FF57BF60A742FC54" pageId="591" pageNumber="590" type="biology_ecology">
<paragraph id="8BAC36B1FD9AFDE8FF57BF60A742FC54" blockId="591.[181,1164,836,913]" pageId="591" pageNumber="590">
HOSTS. Reared from
<taxonomicName id="4C134D32FD9AFDE8FE75BF60A4DAFC99" authority="White &amp; Hodkinson" authorityName="White &amp; Hodkinson" box="[407,714,836,860]" class="Insecta" family="Triozidae" genus="Neolithus" kingdom="Animalia" order="Hemiptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B967EAA3FD9AFDE8FE75BF60A7E8FC99" box="[407,504,836,860]" italics="true" pageId="591" pageNumber="590">Neolithus</emphasis>
White &amp; Hodkinson
</taxonomicName>
(
<taxonomicName id="4C134D32FD9AFDE8FD35BF60A558FC99" box="[727,840,836,860]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
:
<taxonomicName id="4C134D32FD9AFDE8FCB2BF60A5A2FC99" box="[848,946,836,860]" class="Insecta" family="Triozidae" kingdom="Animalia" order="Hemiptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="family">Triozidae</taxonomicName>
) forming closed galls on leaves of
<taxonomicName id="4C134D32FD9AFDE8FED8BF7BA4CEFCB2" authority="K. Schum. &amp; Pittier" authorityName="K. Schum. &amp; Pittier" box="[314,734,863,887]" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="species" species="oligoneurum">
<emphasis id="B967EAA3FD9AFDE8FED8BF7BA41FFCB2" box="[314,527,863,887]" italics="true" pageId="591" pageNumber="590">Sapium oligoneurum</emphasis>
K.Schum. &amp; Pittier
</taxonomicName>
(
<taxonomicName id="4C134D32FD9AFDE8FD0EBF7BA566FCB2" box="[748,886,863,887]" class="Insecta" kingdom="Animalia" order="Malpighiales" pageId="591" pageNumber="590" phylum="Arthropoda" rank="order">Malpighiales</taxonomicName>
:
<taxonomicName id="4C134D32FD9AFDE8FC63BF7BA209FCB2" box="[897,1049,863,887]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) (
<bibRefCitation id="EF824B40FD9AFDE8FBCCBF7BA756FC54" author="Noyes, J. S. &amp; Hanson, P." pageId="591" pageNumber="590" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. &amp; Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes &amp; Hanson, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9AFDE8FF57BF94A7A2FB45" pageId="591" pageNumber="590" type="materials_examined">
<paragraph id="8BAC36B1FD9AFDE8FF57BF94A7BDFC00" blockId="591.[181,1165,944,1153]" box="[181,429,944,965]" pageId="591" pageNumber="590">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FD9AFDE8FF39BFEFA7A2FB45" blockId="591.[181,1165,944,1153]" pageId="591" pageNumber="590">
Type material.
<typeStatus id="54A88813FD9AFDE8FE85BFEFA7ABFC24" box="[359,443,971,993]" pageId="591" pageNumber="590">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FD9AFDE8FE00BFEFA47EFC24" bold="true" box="[482,622,971,993]" pageId="591" pageNumber="590">
<collectingCountry id="F3047621FD9AFDE8FE00BFEFA47EFC24" box="[482,622,971,993]" name="Costa Rica" pageId="591" pageNumber="590">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FD9AFDE8FD99BFEFA4C1FC25" bold="true" box="[635,721,970,993]" pageId="591" pageNumber="590">
<collectingRegion id="49D7F853FD9AFDE8FD99BFEFA4C1FC25" box="[635,721,970,993]" country="Costa Rica" name="San Jose" pageId="591" pageNumber="590">San José</collectingRegion>
</emphasis>
,
<quantity id="4CEB9B54FD9AFDE8FD3FBFEFA516FC24" box="[733,774,971,993]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="591" pageNumber="590" unit="km" value="4.0">4km</quantity>
E San Jeronimo, ex
<taxonomicName id="4C134D32FD9AFDE8FC2BBFEFA252FC25" box="[969,1090,971,992]" class="Insecta" family="Triozidae" genus="Neolithus" kingdom="Animalia" order="Hemiptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B967EAA3FD9AFDE8FC2BBFEFA230FC25" box="[969,1056,971,992]" italics="true" pageId="591" pageNumber="590">Neolithus</emphasis>
sp.
</taxonomicName>
in galls on
<taxonomicName id="4C134D32FD9AFDE8FF36BFC2A79EFC3E" box="[212,398,998,1019]" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="species" species="oligoneuron">
<emphasis id="B967EAA3FD9AFDE8FF36BFC2A79EFC3E" box="[212,398,998,1019]" italics="true" pageId="591" pageNumber="590">Sapium oligoneuron</emphasis>
</taxonomicName>
, collected
<date id="FFAD1071FD9AFDE8FE17BFC1A45CFC3E" box="[501,588,997,1019]" pageId="591" pageNumber="590" value="1993-01-26">26.i.1993</date>
, emerged i-ii.1993 (D. Hollis, P. Hanson).
<typeStatus id="54A88813FD9AFDE8FC3DBFC2A22BFC3E" box="[991,1083,998,1019]" pageId="591" pageNumber="590">Paratypes</typeStatus>
:
<collectionCode id="ED02AE74FD9AFDE8FBA7BFC2A6FDFBD0" pageId="591" pageNumber="590">COSTA RICA</collectionCode>
, 2E, 3G,
<collectingRegion id="49D7F853FD9AFDE8FEABB824A78AFBD3" box="[329,410,1024,1046]" inferredBySuffix="true" pageId="591" pageNumber="590">San José</collectingRegion>
,
<quantity id="4CEB9B54FD9AFDE8FE46B824A7DCFBD3" box="[420,460,1024,1046]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="591" pageNumber="590" unit="km" value="6.0">6km</quantity>
NE San Jeronimo de Moravia, Carr. Carillo,
<quantity id="4CEB9B54FD9AFDE8FC88B824A5BCFBD3" box="[874,940,1024,1046]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="591" pageNumber="590" unit="m" value="1500.0">1500m</quantity>
, ex gall
<taxonomicName id="4C134D32FD9AFDE8FC1BB824A260FBD3" box="[1017,1136,1024,1046]" class="Insecta" family="Triozidae" genus="Neolithus" kingdom="Animalia" order="Hemiptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B967EAA3FD9AFDE8FC1BB824A240FBD0" box="[1017,1104,1024,1045]" italics="true" pageId="591" pageNumber="590">Neolithus</emphasis>
sp.
</taxonomicName>
on
<taxonomicName id="4C134D32FD9AFDE8FF57B83FA77EFBF5" box="[181,366,1051,1072]" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="species" species="oligoneuron">
<emphasis id="B967EAA3FD9AFDE8FF57B83FA77EFBF5" box="[181,366,1051,1072]" italics="true" pageId="591" pageNumber="590">Sapium oligoneuron</emphasis>
</taxonomicName>
, ii-v.1988 (P. Hanson); 3E, 1G,
<collectingRegion id="49D7F853FD9AFDE8FD7DB83FA4E1FBF4" box="[671,753,1051,1073]" inferredBySuffix="true" pageId="591" pageNumber="590">San José</collectingRegion>
, Zurqui de Moravia, psyllid gall on
<taxonomicName id="4C134D32FD9AFDE8FBA0B83FA296FBF5" box="[1090,1158,1051,1072]" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="genus">
<emphasis id="B967EAA3FD9AFDE8FBA0B83FA296FBF5" box="[1090,1158,1051,1072]" italics="true" pageId="591" pageNumber="590">Sapium</emphasis>
</taxonomicName>
,
<date id="FFAD1071FD9AFDE8FF57B811A70DFB8E" box="[181,285,1077,1099]" pageId="591" pageNumber="590" value="1990-12-22">22.xii.1990</date>
(P. Hanson); 7E, 10G, same data as
<typeStatus id="54A88813FD9AFDE8FD8EB811A4AEFB8E" box="[620,702,1077,1099]" pageId="591" pageNumber="590">holotype</typeStatus>
; 1G,
<emphasis id="B967EAA3FD9AFDE8FD0DB812A54DFB8E" bold="true" box="[751,861,1078,1099]" pageId="591" pageNumber="590">
<collectingRegion id="49D7F853FD9AFDE8FD0DB812A54DFB8E" box="[751,861,1078,1099]" country="Costa Rica" name="Puntarenas" pageId="591" pageNumber="590">Puntarenas</collectingRegion>
</emphasis>
, Monteverde,
<geoCoordinate id="EE275076FD9AFDE8FC03B812A23FFB8E" box="[993,1071,1078,1099]" degrees="10" direction="north" minutes="18" orientation="latitude" pageId="591" pageNumber="590" precision="925" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="EE275076FD9AFDE8FBD7B812A298FB8E" box="[1077,1160,1078,1099]" degrees="84" direction="west" minutes="04" orientation="longitude" pageId="591" pageNumber="590" precision="925" value="-84.066666">84°04’W</geoCoordinate>
,
<quantity id="4CEB9B54FD9AFDE8FF57B874A6E7FBA0" box="[181,247,1104,1125]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="591" pageNumber="590" unit="m" value="1400.0">1400m</quantity>
, ex psyllid gall on
<taxonomicName id="4C134D32FD9AFDE8FE4BB875A472FBA0" box="[425,610,1104,1126]" class="Magnoliopsida" family="Euphorbiaceae" genus="Sapium" kingdom="Plantae" order="Malpighiales" pageId="591" pageNumber="590" phylum="Tracheophyta" rank="species" species="oligoneuron">
<emphasis id="B967EAA3FD9AFDE8FE4BB875A472FBA0" box="[425,610,1104,1126]" italics="true" pageId="591" pageNumber="590">Sapium oligoneuron</emphasis>
</taxonomicName>
,
<date id="FFAD1071FD9AFDE8FD8CB874A4BBFBA0" box="[622,683,1104,1125]" pageId="591" pageNumber="590" value="1989-05">v.1989</date>
(K. Brodshaw).
<typeStatus id="54A88813FD9AFDE8FCA5B874A58BFBA3" box="[839,923,1104,1126]" pageId="591" pageNumber="590">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FD9AFDE8FC58B875A202FBA3" box="[954,1042,1105,1126]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="591" pageNumber="590" type="Museum">NHMUK</collectionCode>
,
<typeStatus id="54A88813FD9AFDE8FBFFB875A264FBA3" box="[1053,1140,1105,1126]" pageId="591" pageNumber="590">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FD9AFDE8FF57B84FA6E0FB44" box="[181,240,1131,1153]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="591" pageNumber="590">INBio</collectionCode>
,
<collectionCode id="ED02AE74FD9AFDE8FF1BB84FA741FB45" box="[249,337,1131,1152]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="591" pageNumber="590" type="Museum">NHMUK</collectionCode>
,
<collectionCode id="ED02AE74FD9AFDE8FEB8B84FA7BEFB45" box="[346,430,1131,1152]" pageId="591" pageNumber="590">MZUCR</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFD9AFDE8FF57B8BBA56EFB73" box="[181,894,1182,1207]" pageId="591" pageNumber="590" type="discussion">
<paragraph id="8BAC36B1FD9AFDE8FF57B8BBA56EFB73" blockId="591.[181,894,1182,1207]" box="[181,894,1182,1207]" pageId="591" pageNumber="590">
COMMENTS. See comments under
<taxonomicName id="4C134D32FD9AFDE8FDC9B8BAA470FB73" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[555,608,1182,1206]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="species" species="ufens">
<emphasis id="B967EAA3FD9AFDE8FDC9B8BAA470FB73" box="[555,608,1182,1206]" italics="true" pageId="591" pageNumber="590">ufens</emphasis>
</taxonomicName>
(p. 583) and
<taxonomicName id="4C134D32FD9AFDE8FD0AB8BBA530FB73" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[744,800,1183,1206]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="591" pageNumber="590" phylum="Arthropoda" rank="species" species="gyces">
<emphasis id="B967EAA3FD9AFDE8FD0AB8BBA530FB73" box="[744,800,1183,1206]" italics="true" pageId="591" pageNumber="590">gyces</emphasis>
</taxonomicName>
(p. 608).
</paragraph>
</subSubSection>
</treatment>
</document>
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