treatments-xml/data/03/BA/87/03BA8787FFACFF9DFD5FFA9EFF7E2A48.xml
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<document id="999B756D10FFEF0D32C821BAC79134CD" ID-DOI="10.5281/zenodo.5390317" ID-ISSN="1638-9387" ID-Zenodo-Dep="5390317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630312953056" checkinUser="marcus" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2004" docId="03BA8787FFACFF9DFD5FFA9EFF7E2A48" docLanguage="en" docName="Zoosystema.2004.26.1.87-94.pdf" docOrigin="Zoosystema 26 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Halicarcinus bedfordi Montgomery 1931" docType="treatment" docVersion="4" lastPageNumber="89" masterDocId="FF83FFFFFFADFF9FFFAFFFB5FFA12F0D" masterDocTitle="The Hymenosomatidae (Crustacea, Decapoda, Brachyura) of Timika (Irian Jaya, Indonesia)" masterLastPageNumber="94" masterPageNumber="87" pageNumber="88" updateTime="1699222838032" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E755599336B7348D27658ACC58B44334">The Hymenosomatidae (Crustacea, Decapoda, Brachyura) of Timika (Irian Jaya, Indonesia)</mods:title>
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<mods:affiliation id="C405C37B273B4BA3DB8C612F26145519">Department of Biological Sciences, National University of Singapore, Kent Ridge, Singapore 119260 (Republic of Singapore) peterng @ nus. edu. sg ..</mods:affiliation>
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<mods:date id="87C1D145CD1DA3AEAE5FE6B2AF4F9FD7">2004</mods:date>
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<taxonomicName id="4C134D12FFACFF9EFD5FFA9EFB182A48" ID-CoL="3JBLV" authority="Montgomery, 1931" authorityName="Montgomery" authorityYear="1931" box="[752,1209,1322,1349]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="bedfordi">
<emphasis id="B967EA83FFACFF9EFD5FFA9EFC7C2A48" bold="true" box="[752,989,1323,1349]" italics="true" pageId="1" pageNumber="88">Halicarcinus bedfordi</emphasis>
<bibRefCitation id="EF824B60FFACFF9EFC4BFA9FFB182A48" author="MONTGOMERY S. K." box="[996,1209,1322,1349]" pageId="1" pageNumber="88" pagination="405 - 465" refId="ref4305" refString="MONTGOMERY S. K. 1931. - Report on the Crustacea Brachyura of the Percy Sladen Expedition to the Abrolhos Islands under the leadership of Professor W. J. Daikin in 1913; along with other crabs from Western Australia. Journal of the Linnean Society (Zoology) 37 (253): 405 - 465, pls 24 - 30." type="journal article" year="1931">Montgomery, 1931</bibRefCitation>
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<treatmentCitation id="0AB21080FFACFF9EFD65FAEBFB322A7B" author="MONTGOMERY S. K." box="[714,1171,1373,1398]" page="425" pageId="1" pageNumber="88" year="1931">
<taxonomicName id="4C134D12FFACFF9EFD65FAEBFB322A7B" ID-CoL="3JBLV" authority="Montgomery, 1931: 425" authorityName="Montgomery" authorityPageNumber="425" authorityYear="1931" box="[714,1171,1373,1398]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="bedfordi">
<emphasis id="B967EA83FFACFF9EFD65FAEBFC372A7B" box="[714,918,1374,1398]" italics="true" pageId="1" pageNumber="88">Halicarcinus bedfordi</emphasis>
<bibRefCitation id="EF824B60FFACFF9EFC32FAE8FB322A7B" author="MONTGOMERY S. K." box="[925,1171,1373,1398]" pageId="1" pageNumber="88" pagination="405 - 465" refId="ref4305" refString="MONTGOMERY S. K. 1931. - Report on the Crustacea Brachyura of the Percy Sladen Expedition to the Abrolhos Islands under the leadership of Professor W. J. Daikin in 1913; along with other crabs from Western Australia. Journal of the Linnean Society (Zoology) 37 (253): 405 - 465, pls 24 - 30." type="journal article" year="1931">Montgomery, 1931: 425</bibRefCitation>
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, pl. 27, figs 3, 3a, b. —
<treatmentCitation id="0AB21080FFACFF9EFCC9FAC2FBAB2A82" author="LUCAS J. S." box="[870,1034,1399,1423]" page="181" pageId="1" pageNumber="88" year="1980">
<bibRefCitation id="EF824B60FFACFF9EFCC9FAC2FBAB2A82" author="LUCAS J. S." box="[870,1034,1399,1423]" pageId="1" pageNumber="88" pagination="148 - 247" refId="ref4226" refString="LUCAS J. S. 1980. - Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33: 148 - 247, figs 1 - 10." type="journal article" year="1980">Lucas 1980: 181</bibRefCitation>
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, figs 3A, 5E, 6N, 7G, 9E,
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<treatmentCitation id="0AB21080FFACFF9EFC97FA24FB862AA4" author="CHUANG C. T. N. &amp; NG P. K. L." box="[824,1063,1424,1449]" page="87" pageId="1" pageNumber="88" year="1994">
<bibRefCitation id="EF824B60FFACFF9EFC97FA24FB862AA4" author="CHUANG C. T. N. &amp; NG P. K. L." box="[824,1063,1424,1449]" pageId="1" pageNumber="88" pagination="85 - 92" refId="ref4066" refString="CHUANG C. T. N. &amp; NG P. K. L. 1994. - The ecology and biology of Southeast Asian false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae). Hydrobiologia 285: 85 - 92." type="journal article" year="1994">Chuang &amp; Ng 1994: 87</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AB21080FFACFF9EFBFFFA25FCB92ACF" author="NG P. K. L. &amp; CHUANG C. T. N." page="6" pageId="1" pageNumber="88" year="1996">
<bibRefCitation id="EF824B60FFACFF9EFBFFFA25FCB92ACF" author="NG P. K. L. &amp; CHUANG C. T. N." pageId="1" pageNumber="88" pagination="1 - 82" refId="ref4370" refString="NG P. K. L. &amp; CHUANG C. T. N. 1996. - The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82." type="journal article" year="1996">Ng &amp; Chuang 1996: 6</bibRefCitation>
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.
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MATERIAL EXAMINED. —
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<collectingCountry id="F3047601FFACFF9EFC65FA66FB8D2AE6" box="[970,1068,1491,1515]" name="Indonesia" pageId="1" pageNumber="88">Indonesia</collectingCountry>
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. Otakwa,
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,
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, sandy mud,
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, estuary,
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, coll. A. Haris, 1 3.2 ×
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, 1 3.3 ×
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(
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2002.606), 3 3.1 ×
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, 2.9 ×
<quantity id="4CEB9B74FFACFF9EFBFFF9AAFB012938" box="[1104,1184,1566,1591]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="88" unit="mm" value="3.2">3.2 mm</quantity>
, 3.1 ×
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, 1 3.0 ×
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(
<collectionCode id="ED02AE54FFACFF9EFC46F98DFB82295D" box="[1001,1059,1592,1616]" pageId="1" pageNumber="88">RCO</collectionCode>
).
</paragraph>
<paragraph id="8BAC3691FFACFF9EFD65F9E4FB0F298E" blockId="1.[714,1248,1373,1667]" pageId="1" pageNumber="88">
Ajkwa, intertidal, mud substrate in mangrove area,
<date id="FFAD1051FFACFF9EFD65F9DEFC9F298E" box="[714,830,1643,1667]" pageId="1" pageNumber="88" value="2001-06-12">12.VI.2001</date>
, coll. A. Pratiwi, 1 2.3 ×
<quantity id="4CEB9B74FFACFF9EFBF8F9DEFB34298C" box="[1111,1173,1642,1667]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="88" unit="mm" value="3.0">3 mm</quantity>
, 3
</paragraph>
<paragraph id="8BAC3691FFAFFF9DFF38FF1FFD802FF8" blockId="2.[151,684,169,447]" pageId="2" pageNumber="89">
(2 ovigerous)
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3.3 ×
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, 2.2 ×
<quantity id="4CEB9B74FFAFFF9DFDBFFF1FFDC62FCD" box="[528,615,169,194]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="89" unit="mm" value="2.4">2.4 mm</quantity>
, 3.0 ×
<quantity id="4CEB9B74FFAFFF9DFF38FF76FF442FD7" box="[151,229,195,219]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="89" unit="mm" value="3.3">3.3 mm</quantity>
(
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), 1 3.3 ×
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(
<collectionCode id="ED02AE54FFAFFF9DFDC9FF76FD002FD6" box="[614,673,195,219]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="89" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7B3CCCFFAFFF9DFF38FF68FDBD2FF8" box="[151,540,220,245]" collectingDate="2001-06-15" collectionCode="MZB" pageId="2" pageNumber="89" specimenCount="1">
<date id="FFAD1051FFAFFF9DFF38FF68FEAA2FF8" box="[151,267,221,245]" pageId="2" pageNumber="89" value="2001-06-15">
<collectingDate id="EFE9E9B9FFAFFF9DFF38FF68FEAA2FF8" box="[151,267,221,245]" pageId="2" pageNumber="89" value="2001-06-15">15.VI.2001</collectingDate>
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, 1 2.0 ×
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(
<collectionCode id="ED02AE54FFAFFF9DFE75FF68FDB42FF8" box="[474,533,221,245]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="89" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
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<paragraph id="8BAC3691FFAFFF9DFF38FF40FD312EB2" blockId="2.[151,684,169,447]" pageId="2" pageNumber="89">
<materialsCitation id="3B7B3CCCFFAFFF9DFF38FF40FF442E7E" collectingDate="2001-07-11" collectionCode="RCO, MNHN, MZB" collectorName="Kamora &amp; A. Pratiwi" pageId="2" pageNumber="89" specimenCount="1">
<collectorName id="26E65347FFAFFF9DFF38FF40FF482E01" box="[151,233,245,268]" pageId="2" pageNumber="89">Kamora</collectorName>
, intertidal, mud substrate in mangrove area,
<date id="FFAD1051FFAFFF9DFF38FEBAFEB52E2A" box="[151,276,271,295]" pageId="2" pageNumber="89" value="2001-07-11">
<collectingDate id="EFE9E9B9FFAFFF9DFF38FEBAFEB52E2A" box="[151,276,271,295]" pageId="2" pageNumber="89" value="2001-07-11">11.VII.2001</collectingDate>
</date>
, coll.
<collectorName id="26E65347FFAFFF9DFEE0FEBAFE142E2A" box="[335,437,271,295]" pageId="2" pageNumber="89">A. Pratiwi</collectorName>
, 5 (3 ovigerous) 3.3 ×
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, 2.5 ×
<quantity id="4CEB9B74FFAFFF9DFE91FE9CFE2F2E32" box="[318,398,296,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="89" unit="mm" value="2.8">2.8 mm</quantity>
, 2.4 ×
<quantity id="4CEB9B74FFAFFF9DFE7DFE9CFD812E32" box="[466,544,296,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="89" unit="mm" value="2.8">2.8 mm</quantity>
(
<collectionCode id="ED02AE54FFAFFF9DFD82FE9CFDC62E4C" box="[557,615,297,321]" pageId="2" pageNumber="89">RCO</collectionCode>
), 3.0 ×
<quantity id="4CEB9B74FFAFFF9DFF38FEF7FF462E55" box="[151,231,321,346]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="89" unit="mm" value="3.3">3.3 mm</quantity>
, 3.3 ×
<quantity id="4CEB9B74FFAFFF9DFE82FEF7FEDD2E55" box="[301,380,321,346]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="89" unit="mm" value="3.5">3.5 mm</quantity>
(
<collectionCode id="ED02AE54FFAFFF9DFE25FEF7FE402E57" box="[394,481,322,346]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="89">MNHN</collectionCode>
), 1 2.9 ×
<quantity id="4CEB9B74FFAFFF9DFDF2FEF7FD0D2E55" box="[605,684,321,346]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="89" unit="mm" value="3.3">3.3 mm</quantity>
(
<collectionCode id="ED02AE54FFAFFF9DFF30FEEEFF7C2E7E" box="[159,221,347,371]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="89" type="Museum">MZB</collectionCode>
)
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;
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, 1 2.3 ×
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(
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;
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, 3 2.0 ×
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(
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), 3.3 ×
<quantity id="4CEB9B74FFAFFF9DFD23FEC0FF632EA9" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="89" unit="mm" value="3.8">3.8 mm</quantity>
, 2.2 ×
<quantity id="4CEB9B74FFAFFF9DFEA1FE3BFEC02EA9" box="[270,353,397,422]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="89" unit="mm" value="2.7">2.7 mm</quantity>
, 1 (ovigerous) 3.0 ×
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(
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)
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;
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<date id="FFAD1051FFAFFF9DFEA8FE12FEDB2EB2" box="[263,378,423,447]" pageId="2" pageNumber="89" value="2001-09-20">
<collectingDate id="EFE9E9B9FFAFFF9DFEA8FE12FEDB2EB2" box="[263,378,423,447]" pageId="2" pageNumber="89" value="2001-09-20">20.IX.2001</collectingDate>
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, 1 3.2 ×
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(
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFAFFF9DFF38FE5FFF7E2A48" pageId="2" pageNumber="89" type="discussion">
<paragraph id="8BAC3691FFAFFF9DFF38FE5FFF5A2D0F" blockId="2.[151,684,490,1349]" box="[151,251,490,516]" pageId="2" pageNumber="89">REMARKS</paragraph>
<paragraph id="8BAC3691FFAFFF9DFF38FDBEFDAF2B89" blockId="2.[151,684,490,1349]" pageId="2" pageNumber="89">
The genus
<taxonomicName id="4C134D12FFAFFF9DFEBDFDBEFE372D28" authorityName="White" authorityYear="1846" box="[274,406,523,549]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B967EA83FFAFFF9DFEBDFDBEFE372D28" box="[274,406,523,549]" italics="true" pageId="2" pageNumber="89">Halicarcinus</emphasis>
</taxonomicName>
(
<typeStatus id="54A88833FFAFFF9DFE08FDBEFE762D28" box="[423,471,523,549]" pageId="2" pageNumber="89">type</typeStatus>
species
<taxonomicName id="4C134D12FFAFFF9DFD9EFDBFFEDB2D48" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="planatus">
<emphasis id="B967EA83FFAFFF9DFD9EFDBFFF6E2D4E" italics="true" pageId="2" pageNumber="89">Cancer planatus</emphasis>
Fabricius, 1775
</taxonomicName>
) contains 20 species but only one (
<emphasis id="B967EA83FFAFFF9DFF63FDFEFE332D68" box="[204,402,587,613]" italics="true" pageId="2" pageNumber="89">Halicarcinusfilholi</emphasis>
De Man, 1887, from Java) has been reported previously from Indonesian waters.
<taxonomicName id="4C134D12FFAFFF9DFF42FD3EFE732DA8" authorityName="Montgomery" authorityYear="1931" box="[237,466,651,677]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="bedfordi">
<emphasis id="B967EA83FFAFFF9DFF42FD3EFE732DA8" box="[237,466,651,677]" italics="true" pageId="2" pageNumber="89">Halicarcinus bedfordi</emphasis>
</taxonomicName>
was originally described from
<collectingRegion id="49D7F873FFAFFF9DFEB0FD1FFE402DC8" box="[287,481,682,709]" country="Australia" name="Western Australia" pageId="2" pageNumber="89">Western Australia</collectingRegion>
but is also known from Northern
<collectingCountry id="F3047601FFAFFF9DFEEBFD7EFE022DE8" box="[324,419,715,741]" name="Australia" pageId="2" pageNumber="89">Australia</collectingCountry>
(
<bibRefCitation id="EF824B60FFAFFF9DFE1CFD7EFD962DE8" author="LUCAS J. S." box="[435,567,715,741]" pageId="2" pageNumber="89" pagination="148 - 247" refId="ref4226" refString="LUCAS J. S. 1980. - Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33: 148 - 247, figs 1 - 10." type="journal article" year="1980">Lucas 1980</bibRefCitation>
). The present specimens agree very well with the redescription and figures of this species by
<bibRefCitation id="EF824B60FFAFFF9DFDBBFCBFFD062C28" author="LUCAS J. S." box="[532,679,778,805]" pageId="2" pageNumber="89" pagination="148 - 247" refId="ref4226" refString="LUCAS J. S. 1980. - Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33: 148 - 247, figs 1 - 10." type="journal article" year="1980">Lucas (1980)</bibRefCitation>
, although there is some variation. The chelipeds of the largest males are substantially longer than those figured by
<bibRefCitation id="EF824B60FFAFFF9DFEA5FCDFFE482C88" author="MONTGOMERY S. K." box="[266,489,874,901]" pageId="2" pageNumber="89" pagination="405 - 465" refId="ref4305" refString="MONTGOMERY S. K. 1931. - Report on the Crustacea Brachyura of the Percy Sladen Expedition to the Abrolhos Islands under the leadership of Professor W. J. Daikin in 1913; along with other crabs from Western Australia. Journal of the Linnean Society (Zoology) 37 (253): 405 - 465, pls 24 - 30." type="journal article" year="1931">Montgomery (1931)</bibRefCitation>
and
<bibRefCitation id="EF824B60FFAFFF9DFDB1FCDFFD0D2C88" author="LUCAS J. S." box="[542,684,874,901]" pageId="2" pageNumber="89" pagination="148 - 247" refId="ref4226" refString="LUCAS J. S. 1980. - Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33: 148 - 247, figs 1 - 10." type="journal article" year="1980">Lucas (1980)</bibRefCitation>
for this species, but this is not unexpected, being previously observed in the related
<taxonomicName id="4C134D12FFAFFF9DFD8DFC1FFEEE2CE8" authority="(Rathbun, 1909)" baseAuthorityName="Rathbun" baseAuthorityYear="1909" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="coralicola">
<emphasis id="B967EA83FFAFFF9DFD8DFC1FFD0D2CC8" box="[546,684,938,965]" italics="true" pageId="2" pageNumber="89">H. coralicola</emphasis>
(Rathbun, 1909)
</taxonomicName>
(see
<bibRefCitation id="EF824B60FFAFFF9DFE27FC7FFDCD2CE8" author="NG P. K. L. &amp; CHUANG C. T. N." box="[392,620,970,997]" pageId="2" pageNumber="89" pagination="1 - 82" refId="ref4370" refString="NG P. K. L. &amp; CHUANG C. T. N. 1996. - The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82." type="journal article" year="1996">Ng &amp; Chuang 1996</bibRefCitation>
). The form of the anterolateral margin varies. In smaller specimens, the lateral angle is marked by a small spine, but in larger specimens, especially males, it expands to form a large dentiform structure, giving the carapace a broader appearance.
</paragraph>
<paragraph id="8BAC3691FFAFFF9DFF38FB3EFF7E2A48" blockId="2.[151,684,490,1349]" pageId="2" pageNumber="89">
The male abdomen for this species is as described by
<bibRefCitation id="EF824B60FFAFFF9DFF11FB1EFD062BC8" author="GUINOT D. &amp; RICHER DE FORGES B." box="[190,679,1194,1221]" pageId="2" pageNumber="89" pagination="453 - 502" refId="ref4151" refString="GUINOT D. &amp; RICHER DE FORGES B. 1997. - Affinites entre les Hymenosomatidae MacLeay, 1838 et les Inachoididae Dana, 1851 (Crustacea, Decapoda, Brachyura). Zoosystema 19 (2 - 3): 453 - 502." type="journal article" year="1997">Guinot &amp; Richer de Forges (1997: 466</bibRefCitation>
, fig. 4A) for the genus, in which there are six segments, the telson being fused with segment 6 and barely separated by a faint suture (see also
<bibRefCitation id="EF824B60FFAFFF9DFDC1FABFFF722A48" author="LUCAS J. S." pageId="2" pageNumber="89" pagination="148 - 247" refId="ref4226" refString="LUCAS J. S. 1980. - Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33: 148 - 247, figs 1 - 10." type="journal article" year="1980">Lucas 1980</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>