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<document id="C39B967736045464B8212338BEAD4E99" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56680C0652A08276FD024AFC" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola mazandaranica Marusik, Zamani &amp; Mirshamsi 2014" docType="treatment" docVersion="11" lastPageNumber="34" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="32" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="6A65ED4CBEFE5278135024EB85DECE94">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="AA2DA98A41D7A6BE90479B785E1B070D">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="1FD00A8F0F36D2341C048EB49DAE34F3">Kunt, Kadir B.</mods:namePart>
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<mods:namePart id="778EF77509DE1EE310FF7634B832EAEB">Yağmur, Ersen A.</mods:namePart>
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<mods:title id="A9494FAFE5B6C39E91B249F6ADF704BB">European Journal of Taxonomy</mods:title>
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<mods:date id="B481069EB3948CDC7C5F3990FEDD859F">2018</mods:date>
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<treatment id="03B9B44C56680C0652A08276FD024AFC" ID-DOI="http://doi.org/10.5281/zenodo.5980304" ID-GBIF-Taxon="140561344" ID-Zenodo-Dep="5980304" LSID="urn:lsid:plazi:treatment:03B9B44C56680C0652A08276FD024AFC" httpUri="http://treatment.plazi.org/id/03B9B44C56680C0652A08276FD024AFC" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<subSubSection id="C30A56D156680C3852A08276FB4D4F7A" box="[412,1175,1893,1920]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BAF055A56680C3852A08276FB4D4F7A" blockId="31.[412,1175,1893,1958]" box="[412,1175,1893,1920]" pageId="31" pageNumber="32">
<taxonomicName id="4C107ED956680C3852A08276FB4D4F7A" ID-CoL="4RMTZ" authority="Marusik, Zamani &amp; Mirshamsi, 2014" authorityName="Marusik, Zamani &amp; Mirshamsi" authorityYear="2014" box="[412,1175,1893,1920]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mazandaranica">
<emphasis id="B964D94856680C3852A08276FD034F85" bold="true" box="[412,729,1893,1920]" italics="true" pageId="31" pageNumber="32">Raveniola mazandaranica</emphasis>
<bibRefCitation id="EF8178AB56680C3851DD8276FB4D4F7A" author="Marusik Y. M. &amp; Zamani A. &amp; Mirshamsi O." box="[737,1175,1893,1919]" journalOrPublisher="ZooKeys" pageId="31" pageNumber="32" pagination="1 - 10" part="463" publicationUrl="https://doi.org/10.3897/zookeys.463.8692" refString="Marusik Y. M., Zamani A. &amp; Mirshamsi O. 2014. Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae). ZooKeys 463: 1 - 10. https: // doi. org / 10.3897 / zookeys. 463.8692" title="Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae)" type="journal article" year="2014">Marusik, Zamani &amp; Mirshamsi, 2014</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156680C38511D829FFBC84FA3" box="[545,1042,1932,1958]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BAF055A56680C38511D829FFBC84FA3" blockId="31.[412,1175,1893,1958]" box="[545,1042,1932,1958]" pageId="31" pageNumber="32">
<figureCitation id="132B19DF56680C38511D829FFDA14FA3" box="[545,635,1932,1958]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="31" pageNumber="32">Figs 79</figureCitation>
,
<figureCitation id="132B19DF56680C3851BB829FFD6F4FA3" box="[647,693,1932,1958]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="31" pageNumber="32">112</figureCitation>
,
<figureCitation id="132B19DF56680C3851FD829FFD2A4FA3" box="[705,752,1932,1958]" captionStart="Figs 137–140" captionStartId="70.[189,241,1831,1857]" captionTargetBox="[211,1305,274,1777]" captionTargetId="figure@70.[197,1331,272,1777]" captionText="Figs 137–140. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 137. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 138. R. turcica sp. nov. (holotype). 139. R. micropa (Ausserer, 1871) (Uludağ).140. R. mazandaranica Marusik, Zamani &amp;Mirshamsi, 2014 (holotype). Scalebars: 1.0mm." httpUri="https://zenodo.org/record/1163464/files/figure.png" pageId="31" pageNumber="32">140</figureCitation>
,
<figureCitation id="132B19DF56680C3851C7829FFCF04FA3" box="[763,810,1932,1958]" captionStart="Figs 157–160" captionStartId="75.[189,241,1894,1920]" captionTargetBox="[189,1395,266,1870]" captionTargetId="figure@75.[189,1398,265,1875]" captionText="Figs 157–160. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 157. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 158. R. turcica sp. nov. (holotype). 159. R. micropa (Ausserer, 1871) (Uludağ). 160. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163474/files/figure.png" pageId="31" pageNumber="32">160</figureCitation>
,
<figureCitation id="132B19DF56680C38500A829FFC7A4FA3" box="[822,928,1932,1958]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="31" pageNumber="32">189, 190</figureCitation>
,
<figureCitation id="132B19DF56680C385097829FFC004FA3" box="[939,986,1932,1958]" captionStart="Figure 222" captionStartId="87.[189,272,1855,1881]" captionTargetBox="[189,1398,1034,1819]" captionTargetId="figure@87.[189,1398,1033,1819]" captionText="Figure 222. Localities of Raveniola spp. in Iran: R. dunini sp. nov. (), R. marusiki sp. nov. (). R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (), R. niedermeyeri (Brignoli, 1972) () andR.vonwickiZonstein,2000().Abbreviations:AJ=Azerbaijan;AM =Armenia;UZ =Uzbekistan." httpUri="https://zenodo.org/record/1163500/files/figure.png" pageId="31" pageNumber="32">222</figureCitation>
,
<figureCitation id="132B19DF56680C3850DA829FFBC84FA3" box="[998,1042,1932,1958]" captionStart="Figs 242–247" captionStartId="91.[189,241,1692,1718]" captionTargetBox="[189,1399,265,1643]" captionTargetId="figure@91.[189,1399,265,1643]" captionText="Figs 242–247. Raveniola spp., microhabitats of R. micropa (Ausserer, 1871) in Uludağ Mts (242– 243) and live spiders (244–247). — 244–246. ♀♀ of R. micropa (Ausserer, 1871) near their retreats. 247. Holotype, ♂of R. mazanderanicaMarusik, Zamani&amp;Mirshamsi, 2014(imagecourtesy ofAlireza Zamani)." httpUri="https://zenodo.org/record/1163508/files/figure.png" pageId="31" pageNumber="32">247</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156680C38538182C2FB554FEE" box="[189,1167,2000,2027]" pageId="31" pageNumber="32" type="reference_group">
<paragraph id="8BAF055A56680C38538182C2FB554FEE" blockId="31.[189,1167,2000,2027]" box="[189,1167,2000,2027]" pageId="31" pageNumber="32">
<taxonomicName id="4C107ED956680C38538182C2FC154FEE" ID-CoL="4RMTZ" authority="Marusik, Zamani &amp; Mirshamsi, 2014: 5" authorityName="Marusik, Zamani &amp; Mirshamsi" authorityPageNumber="5" authorityYear="2014" box="[189,975,2000,2027]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mazandaranica">
<emphasis id="B964D94856680C38538182C2FE284FEE" box="[189,498,2001,2027]" italics="true" pageId="31" pageNumber="32">Raveniola mazandaranica</emphasis>
<bibRefCitation id="EF8178AB56680C3852C582C2FC6E4FEE" author="Marusik Y. M. &amp; Zamani A. &amp; Mirshamsi O." box="[505,948,2001,2027]" journalOrPublisher="ZooKeys" pageId="31" pageNumber="32" pagination="1 - 10" part="463" publicationUrl="https://doi.org/10.3897/zookeys.463.8692" refString="Marusik Y. M., Zamani A. &amp; Mirshamsi O. 2014. Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae). ZooKeys 463: 1 - 10. https: // doi. org / 10.3897 / zookeys. 463.8692" title="Three new species of mygalomorph and filistatid spiders from Iran (Araneae, Cyrtaucheniidae, Nemesiidae and Filistatidae)" type="journal article" year="2014">Marusik, Zamani &amp; Mirshamsi, 2014</bibRefCitation>
: 5
</taxonomicName>
, figs 12–18 (♂).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156570C0753818418FC0B49C0" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BAF055A56570C0753818418FEEC4920" blockId="32.[189,310,267,293]" box="[189,310,267,293]" pageId="32" pageNumber="33">
<heading id="D0E7B23656570C0753818418FEEC4920" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B964D94856570C0753818418FEEC4920" bold="true" box="[189,310,267,293]" pageId="32" pageNumber="33">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56570C075381842FFC0B49C0" blockId="32.[189,1399,315,454]" pageId="32" pageNumber="33">
In the shape of the subapically spiraled embolus males of
<taxonomicName id="4C107ED956570C075076842FFBC54950" authorityName="Marusik, Zamani &amp; Mirshamsi" authorityYear="2014" box="[842,1055,315,341]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mazandaranica">
<emphasis id="B964D94856570C075076842FFBC54950" box="[842,1055,315,341]" italics="true" pageId="32" pageNumber="33">R. mazandaranica</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C107ED956570C0757CE842FFAAD4950" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[1266,1399,315,341]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856570C0757CE842FFAAD4950" box="[1266,1399,315,341]" italics="true" pageId="32" pageNumber="33">R. marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356570C0753818472FEC6497E" box="[189,284,353,379]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C107ED956570C0752168472FE6E497F" authorityName="Zonstein" authorityYear="2000" box="[298,436,352,378]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856570C0752168472FE6E497F" box="[298,436,352,378]" italics="true" pageId="32" pageNumber="33">R. vonwicki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C107ED956570C0752CE8472FDB6497F" baseAuthorityName="Charitonov" baseAuthorityYear="1948" box="[498,620,353,378]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856570C0752CE8472FDB6497F" box="[498,620,353,378]" italics="true" pageId="32" pageNumber="33">R. zaitzevi</emphasis>
</taxonomicName>
, but differ from them in having the distal corkscrew-shaped part of the embolus longer and more gently spiraled than in all the mentioned members of the same group (
<figureCitation id="132B19DF56570C0753F984B8FEB149C0" box="[197,363,427,453]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="32" pageNumber="33">Figs 189–190</figureCitation>
; cf.
<figureCitation id="132B19DF56570C0752A084B8FD9B49C3" box="[412,577,427,454]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="32" pageNumber="33">Figs 191–196</figureCitation>
). Female characters are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156570C07538184FDFD874A86" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BAF055A56570C07538184FDFE704A0D" blockId="32.[189,426,494,520]" box="[189,426,494,520]" pageId="32" pageNumber="33">
<heading id="D0E7B23656570C07538184FDFE704A0D" bold="true" box="[189,426,494,520]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B964D94856570C07538184FDFE704A0D" bold="true" box="[189,426,494,520]" pageId="32" pageNumber="33">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F0756570C0753D6870CFD874A86" ID-GBIF-Occurrence="1806244356" collectingDate="2014-04" collectionCode="SMF" collectorName="P. Beyhaghi" country="Iran" county="Māzandarān Province" latitude="36.65" location="Abbās Abad, Salmān Shahr, around Dāniāl Cave" longLatPrecision="1186" longitude="51.166668" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BAF055A56570C0753D6870CFE864A3C" blockId="32.[189,1399,543,644]" box="[234,348,543,569]" pageId="32" pageNumber="33">
<typeStatus id="54ABBBF856570C0753D6870CFE864A3C" box="[234,348,543,569]" pageId="32" pageNumber="33" type="holotype">
<emphasis id="B964D94856570C0753D6870CFE864A3C" bold="true" box="[234,348,543,569]" pageId="32" pageNumber="33">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BAF055A56570C0753818757FD874A86" blockId="32.[189,1399,543,644]" pageId="32" pageNumber="33">
<collectingCountry id="F30745CA56570C0753818757FED34A5B" box="[189,265,580,606]" name="Iran" pageId="32" pageNumber="33">IRAN</collectingCountry>
:
<specimenCount id="9D16CED356570C0752288757FEF64A5B" box="[276,300,580,606]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
,
<collectingCounty id="62CE7DD656570C0752058757FDE64A5B" box="[313,572,580,606]" pageId="32" pageNumber="33">Māzandarān Province</collectingCounty>
,
<location id="8ECF538156570C07517B8757FB5F4A5B" LSID="urn:lsid:plazi:treatment:03B9B44C56680C0652A08276FD024AFC:8ECF538156570C07517B8757FB5F4A5B" box="[583,1157,580,606]" country="Iran" county="Māzandarān Province" latitude="36.65" longLatPrecision="1186" longitude="51.166668" name="Abbās Abad, Salmān Shahr, around Dāniāl Cave" pageId="32" pageNumber="33">Abbās Abad, Salmān Shahr, around Dāniāl Cave</location>
,
<geoCoordinate id="EE24639D56570C0757AD8751FB274A5B" box="[1169,1277,578,606]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="36.65">36°39′ N</geoCoordinate>
,
<geoCoordinate id="EE24639D56570C0756358751FAAB4A5B" box="[1289,1393,578,606]" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="51.166668">51°10′ E</geoCoordinate>
,
<collectingDate id="EFEADA7256570C075381877AFEEC4A86" box="[189,310,617,643]" pageId="32" pageNumber="33" value="2014-04">
<date id="FFAE239A56570C075381877AFEEC4A86" box="[189,310,617,643]" pageId="32" pageNumber="33" value="2014-04">Apr. 2014</date>
</collectingDate>
,
<collectorName id="26E5608C56570C07527D8779FE174A86" box="[321,461,617,644]" pageId="32" pageNumber="33">P. Beyhaghi</collectorName>
leg. (
<collectionCode id="ED019D9F56570C07512C877AFD8B4A86" box="[528,593,617,643]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" pageId="32" pageNumber="33">SMF</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C30A56D156570C06538187BFFEE34A13" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8BAF055A56570C07538187BFFEB14AC3" blockId="32.[189,363,684,710]" box="[189,363,684,710]" pageId="32" pageNumber="33">
<heading id="D0E7B23656570C07538187BFFEB14AC3" bold="true" box="[189,363,684,710]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B964D94856570C07538187BFFEB14AC3" bold="true" box="[189,363,684,710]" pageId="32" pageNumber="33">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56570C0753D687CEFE714AFD" blockId="32.[189,454,733,797]" box="[234,427,733,760]" pageId="32" pageNumber="33">
<emphasis id="B964D94856570C0753D687CEFEF04AF2" bold="true" box="[234,298,733,759]" pageId="32" pageNumber="33">Male</emphasis>
(holotype)
</paragraph>
<paragraph id="8BAF055A56570C0753818610FE1C4B19" blockId="32.[189,454,733,797]" box="[189,454,770,797]" pageId="32" pageNumber="33">
HABITUS. See
<figureCitation id="132B19DF56570C0752618611FE1C4B19" box="[349,454,770,796]" captionStart="Figs 242–247" captionStartId="91.[189,241,1692,1718]" captionTargetBox="[189,1399,265,1643]" captionTargetId="figure@91.[189,1399,265,1643]" captionText="Figs 242–247. Raveniola spp., microhabitats of R. micropa (Ausserer, 1871) in Uludağ Mts (242– 243) and live spiders (244–247). — 244–246. ♀♀ of R. micropa (Ausserer, 1871) near their retreats. 247. Holotype, ♂of R. mazanderanicaMarusik, Zamani&amp;Mirshamsi, 2014(imagecourtesy ofAlireza Zamani)." httpUri="https://zenodo.org/record/1163508/files/figure.png" pageId="32" pageNumber="33">Fig. 247.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56570C0753818655FB7D4B5A" blockId="32.[189,1191,837,864]" box="[189,1191,837,864]" pageId="32" pageNumber="33">MEASUREMENTS. TBL 14.50, CL 5.07, CW 4.68, LL 0.38, LW 0.77, SL 2.57, SW 2.06.</paragraph>
<paragraph id="8BAF055A56570C075381869AFE714C17" blockId="32.[189,1399,905,1042]" pageId="32" pageNumber="33">COLOUR. Carapace, chelicerae, palps and legs dorsally brownish rufous; eye tubercle brownish black; sternum, labium, maxillae, palps, and legs ventrally light brownish rufous; abdomen dorsally brown with darker pattern consisting of partially fused irregular dark brown spots; ventral surface of abdomen light greyish brown.</paragraph>
<paragraph id="8BAF055A56570C075381812FFAB94CC0" blockId="32.[189,1399,1084,1221]" pageId="32" pageNumber="33">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF56570C07503E812FFC864C53" box="[770,860,1084,1110]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="32" pageNumber="33">Fig. 79</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF56570C0757EF812FFAE04C53" box="[1235,1338,1084,1110]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="32" pageNumber="33">Fig. 112</figureCitation>
. Eye diameters and interdistances: AME 0.15(0.21), ALE 0.33, PLE 0.22, PME 0.17, AME–AME 0.12(0.06), ALE–AME 0.07(0.04), ALE–PLE 0.05, PLE–PME 0.04, PME–PME 0.33. Each cheliceral furrow with 9 promarginal teeth and 1–2 mesobasal denticles. Maxillae with 10–11 relatively large cuspules each.
</paragraph>
<paragraph id="8BAF055A56570C07538181FCFECD4D7D" blockId="32.[189,1399,1262,1400]" pageId="32" pageNumber="33">
LEGS. Tibia and metatarsus I as shown in
<figureCitation id="132B19DF56570C07518281FDFCFC4D0C" box="[702,806,1262,1289]" captionStart="Figs 137–140" captionStartId="70.[189,241,1831,1857]" captionTargetBox="[211,1305,274,1777]" captionTargetId="figure@70.[197,1331,272,1777]" captionText="Figs 137–140. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 137. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 138. R. turcica sp. nov. (holotype). 139. R. micropa (Ausserer, 1871) (Uludağ).140. R. mazandaranica Marusik, Zamani &amp;Mirshamsi, 2014 (holotype). Scalebars: 1.0mm." httpUri="https://zenodo.org/record/1163464/files/figure.png" pageId="32" pageNumber="33">Fig. 140</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I–II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–12 each on tibiae, 14–19 on metatarsi, 11–14 on tarsi, 8–9 on cymbium. Paired claws on tarsi I–II with 8–11, on tarsi III–IV with 7–8 teeth on each margin.
</paragraph>
<paragraph id="8BAF055A56570C07538180B1FE7A4DB9" blockId="32.[189,416,1442,1468]" box="[189,416,1442,1468]" pageId="32" pageNumber="33">LEG MEASUREMENTS.</paragraph>
<paragraph id="8BAF055A56570C07525B80CDFAE34EC2" pageId="32" pageNumber="33">
<table id="F910F7FA5657F3D853F480CEFAE34ECD" box="[200,1337,1501,1736]" colsContinueFrom="27.[200,1384,1640,1875]" gridcols="7" gridrows="6" pageId="32" pageNumber="33" rowsContinueFrom="30.[200,1383,326,562]">
<tr id="352007185657F3D853F480CEFAE34DFD" box="[200,1337,1501,1528]" gridrow="0" pageId="32" pageNumber="33" rowspan-0="1">
<th id="76F16E645657F3D8525B80CEFE6E4DFD" box="[359,436,1501,1528]" gridcol="1" gridrow="0" pageId="32" pageNumber="33">Femur</th>
<th id="76F16E645657F3D8512680CEFDB04DFD" box="[538,618,1501,1528]" gridcol="2" gridrow="0" pageId="32" pageNumber="33">Patella</th>
<th id="76F16E645657F3D851E580CEFCCC4DFD" box="[729,790,1501,1528]" gridcol="3" gridrow="0" pageId="32" pageNumber="33">Tibia</th>
<th id="76F16E645657F3D8505180CEFC344DFD" box="[877,1006,1501,1528]" gridcol="4" gridrow="0" pageId="32" pageNumber="33">Metatarsus</th>
<th id="76F16E645657F3D8570080CEFB524DFD" box="[1084,1160,1501,1528]" gridcol="5" gridrow="0" pageId="32" pageNumber="33">Tarsus</th>
<th id="76F16E645657F3D857CB80CEFAE34DFD" box="[1271,1337,1501,1528]" gridcol="6" gridrow="0" pageId="32" pageNumber="33">Total</th>
</tr>
<tr id="352007185657F3D853F48314FAE34E24" box="[200,1337,1543,1569]" gridrow="1" pageId="32" pageNumber="33">
<th id="76F16E645657F3D853F48314FEC64E24" box="[200,284,1543,1569]" gridcol="0" gridrow="1" pageId="32" pageNumber="33">Palp</th>
<td id="76F16E645657F3D8525B8314FE6E4E24" box="[359,436,1543,1569]" gridcol="1" gridrow="1" pageId="32" pageNumber="33">2.71</td>
<td id="76F16E645657F3D851268314FDB04E24" box="[538,618,1543,1569]" gridcol="2" gridrow="1" pageId="32" pageNumber="33">1.48</td>
<td id="76F16E645657F3D851E58314FCCC4E24" box="[729,790,1543,1569]" gridcol="3" gridrow="1" pageId="32" pageNumber="33">2.24</td>
<td id="76F16E645657F3D850518314FC344E24" box="[877,1006,1543,1569]" gridcol="4" gridrow="1" pageId="32" pageNumber="33">–</td>
<td id="76F16E645657F3D857008314FB524E24" box="[1084,1160,1543,1569]" gridcol="5" gridrow="1" pageId="32" pageNumber="33">0.89</td>
<td id="76F16E645657F3D857CB8314FAE34E24" box="[1271,1337,1543,1569]" gridcol="6" gridrow="1" pageId="32" pageNumber="33">7.32</td>
</tr>
<tr id="352007185657F3D853F48323FAE34E4E" box="[200,1337,1584,1611]" gridrow="2" pageId="32" pageNumber="33">
<th id="76F16E645657F3D853F48323FEC64E4E" box="[200,284,1584,1611]" gridcol="0" gridrow="2" pageId="32" pageNumber="33">Leg I</th>
<td id="76F16E645657F3D8525B8323FE6E4E4E" box="[359,436,1584,1611]" gridcol="1" gridrow="2" pageId="32" pageNumber="33">4.02</td>
<td id="76F16E645657F3D851268323FDB04E4E" box="[538,618,1584,1611]" gridcol="2" gridrow="2" pageId="32" pageNumber="33">2.27</td>
<td id="76F16E645657F3D851E58323FCCC4E4E" box="[729,790,1584,1611]" gridcol="3" gridrow="2" pageId="32" pageNumber="33">3.36</td>
<td id="76F16E645657F3D850518323FC344E4E" box="[877,1006,1584,1611]" gridcol="4" gridrow="2" pageId="32" pageNumber="33">3.24</td>
<td id="76F16E645657F3D857008323FB524E4E" box="[1084,1160,1584,1611]" gridcol="5" gridrow="2" pageId="32" pageNumber="33">1.81</td>
<td id="76F16E645657F3D857CB8323FAE34E4E" box="[1271,1337,1584,1611]" gridcol="6" gridrow="2" pageId="32" pageNumber="33">14.70</td>
</tr>
<tr id="352007185657F3D853F48349FAE34E70" box="[200,1337,1626,1653]" gridrow="3" pageId="32" pageNumber="33">
<th id="76F16E645657F3D853F48349FEC64E70" box="[200,284,1626,1653]" gridcol="0" gridrow="3" pageId="32" pageNumber="33">Leg II</th>
<td id="76F16E645657F3D8525B8349FE6E4E70" box="[359,436,1626,1653]" gridcol="1" gridrow="3" pageId="32" pageNumber="33">3.85</td>
<td id="76F16E645657F3D851268349FDB04E70" box="[538,618,1626,1653]" gridcol="2" gridrow="3" pageId="32" pageNumber="33">2.02</td>
<td id="76F16E645657F3D851E58349FCCC4E70" box="[729,790,1626,1653]" gridcol="3" gridrow="3" pageId="32" pageNumber="33">2.73</td>
<td id="76F16E645657F3D850518349FC344E70" box="[877,1006,1626,1653]" gridcol="4" gridrow="3" pageId="32" pageNumber="33">2.76</td>
<td id="76F16E645657F3D857008349FB524E70" box="[1084,1160,1626,1653]" gridcol="5" gridrow="3" pageId="32" pageNumber="33">1.82</td>
<td id="76F16E645657F3D857CB8349FAE34E70" box="[1271,1337,1626,1653]" gridcol="6" gridrow="3" pageId="32" pageNumber="33">13.18</td>
</tr>
<tr id="352007185657F3D853F48397FAE34E9B" box="[200,1337,1668,1694]" gridrow="4" pageId="32" pageNumber="33">
<th id="76F16E645657F3D853F48397FEC64E9B" box="[200,284,1668,1694]" gridcol="0" gridrow="4" pageId="32" pageNumber="33">Leg III</th>
<td id="76F16E645657F3D8525B8397FE6E4E9B" box="[359,436,1668,1694]" gridcol="1" gridrow="4" pageId="32" pageNumber="33">3.37</td>
<td id="76F16E645657F3D851268397FDB04E9B" box="[538,618,1668,1694]" gridcol="2" gridrow="4" pageId="32" pageNumber="33">1.69</td>
<td id="76F16E645657F3D851E58397FCCC4E9B" box="[729,790,1668,1694]" gridcol="3" gridrow="4" pageId="32" pageNumber="33">3.27</td>
<td id="76F16E645657F3D850518397FC344E9B" box="[877,1006,1668,1694]" gridcol="4" gridrow="4" pageId="32" pageNumber="33">3.28</td>
<td id="76F16E645657F3D857008397FB524E9B" box="[1084,1160,1668,1694]" gridcol="5" gridrow="4" pageId="32" pageNumber="33">1.76</td>
<td id="76F16E645657F3D857CB8397FAE34E9B" box="[1271,1337,1668,1694]" gridcol="6" gridrow="4" pageId="32" pageNumber="33">13.37</td>
</tr>
<tr id="352007185657F3D853F483BEFAE34ECD" box="[200,1337,1709,1736]" gridrow="5" pageId="32" pageNumber="33">
<th id="76F16E645657F3D853F483BEFEC64ECD" box="[200,284,1709,1736]" gridcol="0" gridrow="5" pageId="32" pageNumber="33">Leg IV</th>
<td id="76F16E645657F3D8525B83BEFE6E4ECD" box="[359,436,1709,1736]" gridcol="1" gridrow="5" pageId="32" pageNumber="33">4.22</td>
<td id="76F16E645657F3D8512683BEFDB04ECD" box="[538,618,1709,1736]" gridcol="2" gridrow="5" pageId="32" pageNumber="33">1.99</td>
<td id="76F16E645657F3D851E583BEFCCC4ECD" box="[729,790,1709,1736]" gridcol="3" gridrow="5" pageId="32" pageNumber="33">3.36</td>
<td id="76F16E645657F3D8505183BEFC344ECD" box="[877,1006,1709,1736]" gridcol="4" gridrow="5" pageId="32" pageNumber="33">4.78</td>
<td id="76F16E645657F3D8570083BEFB524ECD" box="[1084,1160,1709,1736]" gridcol="5" gridrow="5" pageId="32" pageNumber="33">2.02</td>
<td id="76F16E645657F3D857CB83BEFAE34ECD" box="[1271,1337,1709,1736]" gridcol="6" gridrow="5" pageId="32" pageNumber="33">16.37</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A56570C07538183E1FE2E4FEE" blockId="32.[189,1399,1778,2027]" pageId="32" pageNumber="33">SPINATION. Patella I and metatarsus I aspinose. Palp: femur d1–1–1, pd1; patella p1; tibia d2–1–0, pd0– 1–1, r0–0–1, pv1–1–2, rv1–1–0; cymbium d7. Leg I: femur d1–1–1–1, pd0–1–1; tibia p1–1–0, pv v1– 1–0, rv1–1–m–m. Leg II: femur d1–1–1–1, pd1(0)–1–1; patella p1; tibia p1–1–1, v2–2–3; metatarsus p0–1–0, v2–2–2. Leg III: femur d1–1–1–1(0), pd1–1–1, rd0–1–1; patella p1, r1; tibia d1–1–0, p1–1–1, r1–1–1, v2–2–3(2); metatarsus d1–1–0, p1–1–1, r1–1–1, v3–2–3(2). Leg IV: femur d1–1–1–1, pd1–1–1, rd0–1–1; patella p1–1, r1; tibia d1(0)–1, p1–1–1–1(0), r1–1–1–1, v2–2–3; metatarsus d2–1–1–1, p1–1– 1–1, r1–1–1–1, v3–2–2–3.</paragraph>
<paragraph id="8BAF055A56560C065381841FFC18494C" blockId="33.[189,1398,267,329]" pageId="33" pageNumber="34">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF56560C0650768418FC6E4920" box="[842,948,267,293]" captionStart="Figs 157–160" captionStartId="75.[189,241,1894,1920]" captionTargetBox="[189,1395,266,1870]" captionTargetId="figure@75.[189,1398,265,1875]" captionText="Figs 157–160. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 157. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 158. R. turcica sp. nov. (holotype). 159. R. micropa (Ausserer, 1871) (Uludağ). 160. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163474/files/figure.png" pageId="33" pageNumber="34">Fig. 160</figureCitation>
. Embolic keel lost; embolus slightly corkscrew-shaped with bend located submedially (
<figureCitation id="132B19DF56560C065033843CFC6C494C" box="[783,950,303,329]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="33" pageNumber="34">Figs 189–190</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56560C0653818461FB7949B5" blockId="33.[189,1398,369,432]" pageId="33" pageNumber="34">SPINNERETS. PMS: length 0.31; diameter 0.16. PLS: maximal diameter 0.34; length of basal, medial and apical segments 0.46, 0.36, 0.37; total length 1.19; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A56560C0653D684CBFE9F49F7" blockId="33.[189,325,472,534]" box="[234,325,472,498]" pageId="33" pageNumber="34">
<emphasis id="B964D94856560C0653D684CBFE9F49F7" bold="true" box="[234,325,472,498]" pageId="33" pageNumber="34">Female</emphasis>
</paragraph>
<paragraph id="8BAF055A56560C06538184EFFEE34A13" blockId="33.[189,325,472,534]" box="[189,313,508,534]" pageId="33" pageNumber="34">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156560C065381872DFB954A82" box="[189,1103,574,648]" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="8BAF055A56560C065381872DFEFA4A5D" blockId="33.[189,288,574,600]" box="[189,288,574,600]" pageId="33" pageNumber="34">
<heading id="D0E7B23656560C065381872DFEFA4A5D" bold="true" box="[189,288,574,600]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B964D94856560C065381872DFEFA4A5D" bold="true" box="[189,288,574,600]" pageId="33" pageNumber="34">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56560C065381877EFB954A82" blockId="33.[189,1103,621,648]" box="[189,1103,621,648]" pageId="33" pageNumber="34">The male holotype was collected in a deciduous broad-leaved montane forest.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156560C06538187A3FD024AFC" box="[189,728,688,762]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BAF055A56560C06538187A3FE8C4ACF" blockId="33.[189,342,688,714]" box="[189,342,688,714]" pageId="33" pageNumber="34">
<heading id="D0E7B23656560C06538187A3FE8C4ACF" bold="true" box="[189,342,688,714]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B964D94856560C06538187A3FE8C4ACF" bold="true" box="[189,342,688,714]" pageId="33" pageNumber="34">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56560C06538187F3FD024AFC" blockId="33.[189,728,735,762]" box="[189,728,735,762]" pageId="33" pageNumber="34">
Known only from the type locality (
<figureCitation id="132B19DF56560C06515887CCFD114AFC" box="[612,715,735,761]" captionStart="Figure 222" captionStartId="87.[189,272,1855,1881]" captionTargetBox="[189,1398,1034,1819]" captionTargetId="figure@87.[189,1398,1033,1819]" captionText="Figure 222. Localities of Raveniola spp. in Iran: R. dunini sp. nov. (), R. marusiki sp. nov. (). R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (), R. niedermeyeri (Brignoli, 1972) () andR.vonwickiZonstein,2000().Abbreviations:AJ=Azerbaijan;AM =Armenia;UZ =Uzbekistan." httpUri="https://zenodo.org/record/1163500/files/figure.png" pageId="33" pageNumber="34">Fig. 222</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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