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<document id="F6F344FBD7D5244F19D260EE08A28943" ID-CLB-Dataset="2885" ID-DOI="10.1206/684.1" ID-GBIF-Dataset="9e73be91-fc4a-4e48-84ae-09453e2785a2" ID-ISSN="0003-0082" ID-Zenodo-Dep="4735531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1620069831530" checkinUser="felipe" docAuthor="Ochoa, José A. &amp; Prendini, Lorenzo" docDate="2010" docId="03B987CCFF99F13AFD1A000677B7FEC2" docLanguage="en" docName="AmMusNovit.2010.3687.1-56.pdf" docOrigin="American Museum Novitates 2010 (3687)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2010/issue-3687/684.1/The-Genus-Hadruroides-Pocock-1893-Scorpiones--Iuridae-in-Peru/10.1206/684.1.full" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Hadruroides juanchaparroi Ochoa &amp; Prendini 2010, n. sp." docType="treatment" docVersion="11" lastPageNumber="39" masterDocId="FF80FFB4FFBAF11CFFC50340737BFFBC" masterDocTitle="The Genus Hadruroides Pocock, 1893 (Scorpiones: Iuridae), in Peru: New Records and Descriptions of Six New Species" masterLastPageNumber="56" masterPageNumber="1" pageNumber="36" updateTime="1698938218626" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C774A02A7D0FE4277BE2D4C03190948B">The Genus Hadruroides Pocock, 1893 (Scorpiones: Iuridae), in Peru: New Records and Descriptions of Six New Species</mods:title>
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<mods:namePart id="5D13FE3F124C66AE2EDBD7CCE7E0BD55">Ochoa, José A.</mods:namePart>
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<mods:namePart id="39FE1B555C4850DC43C51E86001B81E2">Prendini, Lorenzo</mods:namePart>
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<treatment id="03B987CCFF99F13AFD1A000677B7FEC2" ID-DOI="http://doi.org/10.5281/zenodo.5795692" ID-GBIF-Taxon="191692367" ID-Zenodo-Dep="5795692" LSID="urn:lsid:plazi:treatment:03B987CCFF99F13AFD1A000677B7FEC2" httpUri="http://treatment.plazi.org/id/03B987CCFF99F13AFD1A000677B7FEC2" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36">
<subSubSection id="C30A6551FF99F13FFD1A00067721FCE2" box="[735,1114,838,862]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BAF36DAFF99F13FFD1A00067721FCE2" blockId="35.[735,1114,838,862]" box="[735,1114,838,862]" pageId="35" pageNumber="36">
<heading id="D0E781B6FF99F13FFD1A00067721FCE2" box="[735,1114,838,862]" centered="true" fontSize="36" level="2" pageId="35" pageNumber="36" reason="8">
<taxonomicName id="4C104D59FF99F13FFD1A00067770FCE2" ID-CoL="3J7DD" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[735,1035,838,862]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="juanchaparroi" status="sp. nov.">
<emphasis id="B964EAC8FF99F13FFD1A00067770FCE2" bold="true" box="[735,1035,838,862]" italics="true" pageId="35" pageNumber="36">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FF99F13FFBDE00067721FCE2" box="[1051,1114,838,862]" pageId="35" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C30A6551FF99F13FFD5A003770B8FC10" pageId="35" pageNumber="36" type="description">
<paragraph id="8BAF36DAFF99F13FFD5A003770B8FC10" blockId="35.[671,1178,887,940]" pageId="35" pageNumber="36">
<figureCitation id="132B2A5FFF99F13FFD5A00377076FC33" box="[671,781,887,911]" captionStart="Fig" captionStartId="3.[112,150,1574,1596]" captionTargetBox="[175,1100,210,1540]" captionTargetId="figure-1@3.[175,1100,210,1541]" captionTargetPageId="3" captionText="Fig. 2. Map of Peru, plotting known locality records of Hadruroides Pocock, 1893. Hadruroides carinatus Pocock, 1900, open circles. Hadruroides charcasus (Karsch, 1879), solid squares. Hadruroides chinchaysuyu, n. sp., diamonds. Hadruroides juanchaparroi, n. sp., solid circles. Hadruroides lunatus (L. Koch, 1867), open squares. Hadruroides mauryi Francke and Soleglad, 1980, open triangles. Undetermined Hadruroides species, stars." figureDoi="http://doi.org/10.5281/zenodo.4735535" httpUri="https://zenodo.org/record/4735535/files/figure.png" pageId="35" pageNumber="36">Figures 2</figureCitation>
,
<figureCitation id="132B2A5FFF99F13FFCDF00377046FC33" box="[794,829,887,911]" captionStart="Fig" captionStartId="6.[151,189,1553,1575]" captionTargetBox="[215,1140,214,1519]" captionTargetId="figure-0@6.[215,1140,210,1519]" captionTargetPageId="6" captionText="Fig. 3. Hadruroides Pocock, 1893, habitats in Peru. A. Puchaca Alto (Lambayeque Department), habitat of Hadruroides leopardus Pocock, 1900. B. Balsas (Cajamarca Department), habitat of Hadruroides geckoi, n. sp. C. Cerro Campana (La Libertad Department), habitat of Hadruroides juanchaparroi, n. sp. D. Wari (Ayacucho Department), habitat of Hadruroides bustamantei Ochoa and Chaparro, 2008. E. Isla Santa (Ancash Department), habitat of Hadruroides tishqu, n. sp. F. Playa Vichayitos (Piura Department), habitat of Hadruroides vichayitos, n. sp. G. Lomas de Lachay (Lima Department), habitat of Hadruroides lunatus (L. Koch, 1867)." figureDoi="http://doi.org/10.5281/zenodo.4735537" httpUri="https://zenodo.org/record/4735537/files/figure.png" pageId="35" pageNumber="36">3C</figureCitation>
,
<figureCitation id="132B2A5FFF99F13FFC8E00377014FC33" box="[843,879,887,911]" captionStart="Fig" captionStartId="8.[151,189,1615,1637]" captionTargetBox="[136,1218,210,1581]" captionTargetId="figure-0@8.[136,1218,210,1581]" captionText="Fig. 5. Hadruroides Pocock, 1893, habitus in life. A. Hadruroides juanchaparroi, n. sp., ♂. B. Hadruroides leopardus Pocock, 1900, ♂. C. Hadruroides lunatus (L. Koch, 1867), ♀. D. Hadruroides vichayitos, n. sp., ♂. E. Hadruroides tishqu, n. sp., ♂." figureDoi="http://doi.org/10.5281/zenodo.5794287" httpUri="https://zenodo.org/record/5794287/files/figure.png" pageId="35" pageNumber="36">5A</figureCitation>
, 18, 19, 20A, B, DF, 21;
<tableCitation id="C6920361FF99F13FFCB300D470B8FC10" box="[886,963,916,940]" captionStart="TABLE 2" captionStartId="27.[817,890,445,464]" captionTargetBox="[88,1623,611,1515]" captionTargetPageId="27" captionText="TABLE 2 Selected measurements (mm) of type specimens of six new species of Hadruroides Pocock, 1893 Hadruroides chinchaysuyu, n. sp., Hadruroides geckoi, n. sp., Hadruroides graceae, n. sp., Hadruroides juanchaparroi, n. sp., Hadruroides tishqu, n. sp., and Hadruroides vichayitos, n. sp. Abbreviations: AMNH 5 American Museum of Natural History, New York; MHNC 5 Museo de Historia Natural, Universidad Nacional de San Antonio Abad del Cusco, Peru." httpUri="http://table.plazi.org/id/DF6F6652FFA1F107FCF402FD7616FD87" pageId="35" pageNumber="36" tableUuid="DF6F6652FFA1F107FCF402FD7616FD87">table 2</tableCitation>
</paragraph>
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TYPE MATERIAL:
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:
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:
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<collectingMunicipality id="6BCBACA0FF99F13FFCF500A5708FFC41" box="[816,1012,997,1021]" pageId="35" pageNumber="36">Trujillo Province</collectingMunicipality>
</emphasis>
:
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♂,
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♂ (
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), 2
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(
<collectionCode id="ED01AE1FFF99F13FFD5B07607186FB84" box="[670,765,1056,1080]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="35" pageNumber="36">AMNH</collectionCode>
),
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,
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07
<emphasis id="B964EAC8FF99F13FFB8107607735FB84" bold="true" box="[1092,1102,1056,1080]" italics="true" pageId="35" pageNumber="36">°</emphasis>
58
<emphasis id="B964EAC8FF99F13FFBAD0761770AFB8B" bold="true" box="[1128,1137,1057,1079]" italics="true" pageId="35" pageNumber="36">'</emphasis>
08
<emphasis id="B964EAC8FF99F13FFB49076177EEFB8B" bold="true" box="[1164,1173,1057,1079]" italics="true" pageId="35" pageNumber="36">&quot;</emphasis>
S 79
<emphasis id="B964EAC8FF99F13FFD75077D71C1FBE9" bold="true" box="[688,698,1085,1109]" italics="true" pageId="35" pageNumber="36">°</emphasis>
05
<emphasis id="B964EAC8FF99F13FFD10077E71A5FBE8" bold="true" box="[725,734,1086,1108]" italics="true" pageId="35" pageNumber="36">'</emphasis>
59
<emphasis id="B964EAC8FF99F13FFD3D077E707AFBE8" bold="true" box="[760,769,1086,1108]" italics="true" pageId="35" pageNumber="36">&quot;</emphasis>
W
</geoCoordinate>
,
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<elevation id="003DD1E9FF99F13FFC8D077D70D5FBE9" box="[840,942,1085,1109]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="36" unit="m" value="450.0">450 m</elevation>
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,
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<collectingDate id="EFEAE9F2FF99F13FFC12077D7737FBE9" box="[983,1100,1085,1109]" pageId="35" pageNumber="36" value="2004-11-15">15.xi.2004</collectingDate>
</date>
,
<collectorName id="26E5530CFF99F13FFBB3077D707CFBCE" pageId="35" pageNumber="36">J.C. Chaparro</collectorName>
,
<collectorName id="26E5530CFF99F13FFCD0071A7005FBCE" box="[789,894,1114,1138]" pageId="35" pageNumber="36">J. Nuñez</collectorName>
and
<collectorName id="26E5530CFF99F13FFC78071A7732FBCE" box="[957,1097,1114,1138]" pageId="35" pageNumber="36">J.A. Ochoa.</collectorName>
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<emphasis id="B964EAC8FF99F13FFB96071A705BFB33" bold="true" pageId="35" pageNumber="36">
<collectingRegion id="49D4F838FF99F13FFB96071A77DFFBCE" box="[1107,1188,1114,1138]" country="Peru" name="Ancash" pageId="35" pageNumber="36">Ancash</collectingRegion>
Department:
</emphasis>
<emphasis id="B964EAC8FF99F13FFCEA073870A4FB2C" box="[815,991,1144,1168]" italics="true" pageId="35" pageNumber="36">
<location id="8ECF6001FF99F13FFCEA073870A4FB2C" LSID="urn:lsid:plazi:treatment:03B987CCFF99F13AFD1A000677B7FEC2:8ECF6001FF99F13FFCEA073870A4FB2C" box="[815,991,1144,1168]" country="Peru" county="Department" latitude="-78.652504" municipality="Trujillo Province" name="Santa Province" pageId="35" pageNumber="36" stateProvince="La Libertad">Santa Province</location>
</emphasis>
: 2
<specimenCount id="9D16FD53FF99F13FFBCA073277D8FB33" box="[1039,1187,1138,1169]" pageId="35" pageNumber="36" type="generic" typeStatus="paratypes">♂ paratypes</specimenCount>
(
<collectionCode id="ED01AE1FFF99F13FFD5B07D47186FB10" box="[670,765,1172,1196]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="35" pageNumber="36">AMNH</collectionCode>
), 1 ♂,
<specimenCount id="9D16FD53FF99F13FFCAE07D470E4FB12" box="[875,927,1170,1198]" pageId="35" pageNumber="36" type="female">1 ♀</specimenCount>
<typeStatus id="54AB8878FF99F13FFC7407D47764FB10" box="[945,1055,1172,1196]" pageId="35" pageNumber="36" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED01AE1FFF99F13FFBFC07D477EDFB10" box="[1081,1174,1172,1196]" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="35" pageNumber="36">MHNC</collectionCode>
),
<collectorName id="26E5530CFF99F13FFD5307F27071FB76" box="[662,778,1202,1226]" pageId="35" pageNumber="36">Chimbote</collectorName>
,
<collectorName id="26E5530CFF99F13FFCF007F27092FB76" box="[821,1001,1202,1226]" pageId="35" pageNumber="36">Caleta Santa</collectorName>
,
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08
<emphasis id="B964EAC8FF99F13FFBEA07F27742FB76" bold="true" box="[1071,1081,1202,1226]" italics="true" pageId="35" pageNumber="36">°</emphasis>
59
<emphasis id="B964EAC8FF99F13FFB9107F37726FB75" bold="true" box="[1108,1117,1203,1225]" italics="true" pageId="35" pageNumber="36">'</emphasis>
25.8
<emphasis id="B964EAC8FF99F13FFB4907F377EEFB75" bold="true" box="[1164,1173,1203,1225]" italics="true" pageId="35" pageNumber="36">&quot;</emphasis>
S 78
<emphasis id="B964EAC8FF99F13FFD75078F71C1FB5B" bold="true" box="[688,698,1231,1255]" italics="true" pageId="35" pageNumber="36">°</emphasis>
39
<emphasis id="B964EAC8FF99F13FFD10079071A5FB5A" bold="true" box="[725,734,1232,1254]" italics="true" pageId="35" pageNumber="36">'</emphasis>
12.9
<emphasis id="B964EAC8FF99F13FFCC80790706DFB5A" bold="true" box="[781,790,1232,1254]" italics="true" pageId="35" pageNumber="36">&quot;</emphasis>
W
</geoCoordinate>
,
<quantity id="4CE89B3FFF99F13FFC84078F70E8FB5B" box="[833,915,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.35" pageId="35" pageNumber="36" unit="m" value="13.5">
<elevation id="003DD1E9FF99F13FFC84078F70E8FB5B" box="[833,915,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.35" pageId="35" pageNumber="36" unit="m" value="13.5">13.5 m</elevation>
</quantity>
,
<date id="FFAE101AFF99F13FFC64078F7087FB5B" box="[929,1020,1231,1255]" pageId="35" pageNumber="36" value="2008-01-04">
<collectingDate id="EFEAE9F2FF99F13FFC64078F7087FB5B" box="[929,1020,1231,1255]" pageId="35" pageNumber="36" value="2008-01-04">4.i.2008</collectingDate>
</date>
,
<collectionCode id="ED01AE1FFF99F13FFBCF078F7759FB5B" box="[1034,1058,1231,1255]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="35" pageNumber="36">R</collectionCode>
.
<collectorName id="26E5530CFF99F13FFBF5078F71D4FAB8" pageId="35" pageNumber="36">Gutiérrez, D</collectorName>
. Apaza and
<collectorName id="26E5530CFF99F13FFC8407AC70B0FAB8" box="[833,971,1260,1284]" pageId="35" pageNumber="36">J.A. Ochoa.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FF99F13FFD75064A71A9FA09" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8BAF36DAFF99F13FFD75064A71A9FA09" blockId="35.[662,1188,968,1754]" pageId="35" pageNumber="36">
ETYMOLOGY: The specific name is a patronym honoring Peruvian herpetologist, Juan Carlos Chaparro (Universidad Nacional San Antonio Abad,
<collectingRegion id="49D4F838FF99F13FFC97062270E7FAC6" box="[850,924,1378,1402]" country="Peru" name="Cusco" pageId="35" pageNumber="36">Cusco</collectingRegion>
), in recognition of his help and participation in several fieldtrips in
<collectingCountry id="F307764AFF99F13FFD5306DD71B5FA09" box="[662,718,1437,1461]" name="Peru" pageId="35" pageNumber="36">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FF99F138FD7506FB7723FA81" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BAF36DAFF99F138FD7506FB7723FA81" blockId="35.[662,1188,968,1754]" lastBlockId="36.[701,1228,1141,1754]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
DIAGNOSIS:
<taxonomicName id="4C104D59FF99F13FFC8306FB770FFA6F" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[838,1140,1467,1491]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF99F13FFC8306FB770FFA6F" box="[838,1140,1467,1491]" italics="true" pageId="35" pageNumber="36">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
appears most closely similar to
<taxonomicName id="4C104D59FF99F13FFC2106987721FA4C" authorityName=": Kraepelin" authorityYear="1894" box="[996,1114,1496,1520]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF99F13FFC2106987721FA4C" box="[996,1114,1496,1520]" italics="true" pageId="35" pageNumber="36">H. lunatus</emphasis>
</taxonomicName>
in the following respects: the VSM carinae are absent and the VL carinae obsolete on metasomal segments IIV; sternite VII is acarinate; the EM carinae of the leg patella are absent; the fixed finger of the pedipalp chela of the adult male is curved, creating a distinct proximal gap with the movable finger when the fingers are closed; the granulation of metasomal segment V and telson are similar; and the relative dimensions of the pedipalp chela, metasomal segment V, and telson are similar. The two species may be distinguished as follows:
<taxonomicName id="4C104D59FF9EF138FEC707AB72BAFABF" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[258,449,1259,1283]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9EF138FEC707AB72BAFABF" box="[258,449,1259,1283]" italics="true" pageId="36" pageNumber="37">H. juanchaparroi</emphasis>
</taxonomicName>
is smaller in size, reaching ca.
<quantity id="4CE89B3FFF9EF138FEE3064872FBFA9C" box="[294,384,1288,1312]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="36" pageNumber="37" unit="mm" value="32.0">32 mm</quantity>
in total length (♂), compared with
<taxonomicName id="4C104D59FF9EF138FE8C066672B3FA82" authorityName=": Kraepelin" authorityYear="1894" box="[329,456,1318,1342]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FE8C066672B3FA82" box="[329,456,1318,1342]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
, which reaches
<quantity id="4CE89B3FFF9EF138FF45060373B4FAE7" box="[128,207,1347,1371]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="36" pageNumber="37" unit="mm" value="45.0">45 mm</quantity>
; the proximal gap between the pedipalp chela fingers of the male is less pronounced in
<taxonomicName id="4C104D59FF9EF138FF45063E7246FA2A" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[128,317,1406,1430]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9EF138FF45063E7246FA2A" box="[128,317,1406,1430]" italics="true" pageId="36" pageNumber="37">H. juanchaparroi</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C104D59FF9EF138FE5B063E716AFA2A" authorityName=": Kraepelin" authorityYear="1894" box="[414,529,1406,1430]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FE5B063E716AFA2A" box="[414,529,1406,1430]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
. There are also some differences in pigmentation between the two species: the dorsosubmedian spots on the tergites are rectangular in
<taxonomicName id="4C104D59FF9EF138FDCB069771F5FA53" authorityName=": Kraepelin" authorityYear="1894" box="[526,654,1495,1519]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FDCB069771F5FA53" box="[526,654,1495,1519]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A5FFF9EF138FF4206B4739FF9B0" box="[135,228,1524,1548]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="36" pageNumber="37">fig. 20C</figureCitation>
) and irregular in
<taxonomicName id="4C104D59FF9EF138FE0C06B471F5F9B0" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[457,654,1524,1548]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9EF138FE0C06B471F5F9B0" box="[457,654,1524,1548]" italics="true" pageId="36" pageNumber="37">H. juanchaparroi</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A5FFF9EF138FF4205517392F995" box="[135,233,1553,1577]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="36" pageNumber="37">fig. 20D</figureCitation>
);
<taxonomicName id="4C104D59FF9EF138FEC605517200F995" authorityName=": Kraepelin" authorityYear="1894" box="[259,379,1553,1577]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FEC605517200F995" box="[259,379,1553,1577]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
lacks pigmentation on sternite VII and metasomal segment I whereas several spots surround the insertion of setae on these segments in
<taxonomicName id="4C104D59FF9EF138FE78052A71F5F93E" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[445,654,1642,1666]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9EF138FE78052A71F5F93E" box="[445,654,1642,1666]" italics="true" pageId="36" pageNumber="37">H. juanchaparroi</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A5FFF9EF138FF4205C7739EF923" box="[135,229,1671,1695]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="36" pageNumber="37">fig. 20E</figureCitation>
); the pigmentation pattern on the ventral surfaces of metasomal segments IIIV is similar in the two species, but the stripes and spots are less evident and may be vestigial in some adult specimens of
<taxonomicName id="4C104D59FF9EF138FC1907D2772BFB16" authorityName=": Kraepelin" authorityYear="1894" box="[988,1104,1170,1194]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FC1907D2772BFB16" box="[988,1104,1170,1194]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
. The most important difference between the two species is the shape of hemispermatophore, which is slender with a small crest in
<taxonomicName id="4C104D59FF9EF138FBC907AA77B0FABE" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[1036,1227,1258,1282]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9EF138FBC907AA77B0FABE" box="[1036,1227,1258,1282]" italics="true" pageId="36" pageNumber="37">H. juanchaparroi</emphasis>
</taxonomicName>
(
<figureCitation id="132B2A5FFF9EF138FD0006487029FA9C" box="[709,850,1288,1312]" captionStart="Fig" captionStartId="36.[151,189,1044,1066]" captionTargetBox="[184,1170,210,1010]" captionTargetId="figure-297@36.[184,1171,210,1010]" captionText="Fig. 21. Hadruroides juanchaparroi, n. sp., paratype ³ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5713964" httpUri="https://zenodo.org/record/5713964/files/figure.png" pageId="36" pageNumber="37">fig. 21A, C</figureCitation>
), but broader basally with a relatively longer crest in
<taxonomicName id="4C104D59FF9EF138FC1E0665772BFA81" authorityName=": Kraepelin" authorityYear="1894" box="[987,1104,1317,1341]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B964EAC8FF9EF138FC1E0665772BFA81" box="[987,1104,1317,1341]" italics="true" pageId="36" pageNumber="37">H. lunatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF6F6652FF9EF138FF52075470B7FBF8" ID-DOI="http://doi.org/10.5281/zenodo.5713964" ID-Zenodo-Dep="5713964" httpUri="https://zenodo.org/record/5713964/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,189,1044,1066]" targetBox="[184,1170,210,1010]" targetPageId="36">
<paragraph id="8BAF36DAFF9EF138FF52075470B7FBF8" blockId="36.[128,1227,1039,1092]" pageId="36" pageNumber="37">
Fig. 21.
<taxonomicName id="4C104D59FF9EF138FF3807547172FB95" authority="Ochoa &amp; Prendini, 2010" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[253,521,1044,1065]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="juanchaparroi" status="sp. nov.">
<emphasis id="B964EAC8FF9EF138FF3807547172FB95" box="[253,521,1044,1065]" italics="true" pageId="36" pageNumber="37">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25757B3FF9EF138FDD20754712BFB96" box="[535,592,1044,1066]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
,
<typeStatus id="54AB8878FF9EF138FD9E075471CEFB96" box="[603,693,1044,1066]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
♂ (
<collectionCode id="ED01AE1FFF9EF138FD1A07547048FB96" box="[735,819,1044,1066]" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="36" pageNumber="37">MHNC</collectionCode>
), sinistral hemispermatophore.
<collectorName id="26E5530CFF9EF138FBB0075477B0FB96" box="[1141,1227,1044,1066]" pageId="36" pageNumber="37">
<emphasis id="B964EAC8FF9EF138FBB0075477F6FB96" bold="true" box="[1141,1165,1044,1066]" pageId="36" pageNumber="37">A.</emphasis>
Ental
</collectorName>
aspect.
<collectorName id="26E5530CFF9EF138FF08076E7249FBF8" box="[205,306,1070,1092]" pageId="36" pageNumber="37">
<emphasis id="B964EAC8FF9EF138FF08076E739FFBF8" bold="true" box="[205,228,1070,1092]" pageId="36" pageNumber="37">B.</emphasis>
Dorsal
</collectorName>
aspect.
<collectorName id="26E5530CFF9EF138FE42076E72A7FBF8" box="[391,476,1070,1092]" pageId="36" pageNumber="37">
<emphasis id="B964EAC8FF9EF138FE42076E72E4FBF8" bold="true" box="[391,415,1070,1092]" pageId="36" pageNumber="37">C.</emphasis>
Ectal
</collectorName>
aspect.
<collectorName id="26E5530CFF9EF138FDF4076E71E5FBF8" box="[561,670,1070,1092]" pageId="36" pageNumber="37">
<emphasis id="B964EAC8FF9EF138FDF4076E7131FBF8" bold="true" box="[561,586,1070,1092]" pageId="36" pageNumber="37">D.</emphasis>
Ventral
</collectorName>
aspect.
<collectorName id="26E5530CFF9EF138FD31076E7052FBF8" box="[756,809,1070,1092]" pageId="36" pageNumber="37">Scale</collectorName>
bar
<emphasis id="B964EAC8FF9EF138FC9E076E7014FBF8" bold="true" box="[859,879,1070,1092]" italics="true" pageId="36" pageNumber="37">=</emphasis>
<quantity id="4CE89B3FFF9EF138FCB2076E70B3FBF8" box="[887,968,1070,1092]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="37" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<subSubSection id="C30A6551FF9EF13AFD1D06037103FA13" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
<paragraph id="8BAF36DAFF9EF138FD1D060377E5FA2A" blockId="36.[701,1228,1141,1754]" pageId="36" pageNumber="37">
DESCRIPTION: Based on the
<typeStatus id="54AB8878FF9EF138FBF5060377EFFAE7" box="[1072,1172,1347,1371]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
and
<typeStatus id="54AB8878FF9EF138FD7806217054FAC5" box="[701,815,1377,1401]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
. Measurements of the
<typeStatus id="54AB8878FF9EF138FB81062177D3FAC5" box="[1092,1192,1377,1401]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
♂ and a
<typeStatus id="54AB8878FF9EF138FCC2063E7010FA2A" box="[775,875,1406,1430]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
<specimenCount id="9D16FD53FF9EF138FCB1063B70F2FA2B" box="[884,905,1403,1431]" pageId="36" pageNumber="37" type="female"></specimenCount>
are recorded in
<tableCitation id="C6920361FF9EF138FB8C063E77E2FA2A" box="[1097,1177,1406,1430]" captionStart="TABLE 2" captionStartId="27.[817,890,445,464]" captionTargetBox="[88,1623,611,1515]" captionTargetPageId="27" captionText="TABLE 2 Selected measurements (mm) of type specimens of six new species of Hadruroides Pocock, 1893 Hadruroides chinchaysuyu, n. sp., Hadruroides geckoi, n. sp., Hadruroides graceae, n. sp., Hadruroides juanchaparroi, n. sp., Hadruroides tishqu, n. sp., and Hadruroides vichayitos, n. sp. Abbreviations: AMNH 5 American Museum of Natural History, New York; MHNC 5 Museo de Historia Natural, Universidad Nacional de San Antonio Abad del Cusco, Peru." httpUri="http://table.plazi.org/id/DF6F6652FFA1F107FCF402FD7616FD87" pageId="36" pageNumber="37" tableUuid="DF6F6652FFA1F107FCF402FD7616FD87">table 2</tableCitation>
.
</paragraph>
<paragraph id="8BAF36DAFF9EF139FD1D06DC7248FB7D" blockId="36.[701,1228,1141,1754]" lastBlockId="37.[88,614,211,1754]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="B964EAC8FF9EF138FD1D06DC706CFA08" box="[728,791,1436,1460]" italics="true" pageId="36" pageNumber="37">Color</emphasis>
:
<collectorName id="26E5530CFF9EF138FCF706DC701CFA08" box="[818,871,1436,1460]" pageId="36" pageNumber="37">Base</collectorName>
color yellowish, except for pedipalp chela, metasomal segment V, and telson, which are brownish yellow.
<collectorName id="26E5530CFF9EF138FBA5069777B0FA53" box="[1120,1227,1495,1519]" pageId="36" pageNumber="37">Carapace</collectorName>
markedly pigmented, especially laterally, where two oblique stripes extend from median to anterolateral margins; ocular tubercle darker; anteromedian longitudinal sulcus markedly pigmented; four irregular spots situated near posterior margin; anterior margin with band of pigmentation connecting lateral ocelli.
<collectorName id="26E5530CFF9EF138FCA2058270A0F966" box="[871,987,1730,1754]" pageId="36" pageNumber="37">Tergites I</collectorName>
VI each with four irregular spots, two submedian and two sublateral, occasionally joining weakly at anterior margin; pretergites with two lateral spots (
<figureCitation id="132B2A5FFF9FF139FF63026C727CFEF8" box="[166,263,300,324]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="37" pageNumber="38">fig. 20D</figureCitation>
); VII faintly pigmented laterally, depigmented medially.
<collectorName id="26E5530CFF9FF139FE69020A7175FEDE" box="[428,526,330,354]" pageId="37" pageNumber="38">Sternites</collectorName>
IIIVI depigmented; VII with eight small spots surrounding insertion of setae (
<figureCitation id="132B2A5FFF9FF139FE3E02C57121FE21" box="[507,602,389,413]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="37" pageNumber="38">fig. 20E</figureCitation>
).
<collectorName id="26E5530CFF9FF139FF9D02E373ACFE07" box="[88,215,419,443]" pageId="37" pageNumber="38">Metasomal</collectorName>
segments IIV, dorsal surfaces faintly pigmented along DL carinae and granulation, occasionally with two submedian spots medially (as in
<typeStatus id="54AB8878FF9FF139FE8702BC72D1FDA8" box="[322,426,508,532]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
); lateral surfaces pigmented in posterior half; ML and LIM carinae pigmented; ventral surfaces with two stripes along VL carinae, joining posteriorly to lateral pigmentation on segments IIIV, often also with spots surrounding insertion of setae: usually with 2+2 spots on segment I, 3+3 on segments II and III, and 4+4 or 5+5 spots on segment IV (
<figureCitation id="132B2A5FFF9FF139FF2E01AA7231FCBE" box="[235,330,746,770]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="37" pageNumber="38">fig. 20E</figureCitation>
). Metasomal segment V, dorsal and lateral surfaces markedly pigmented in posterior half; ventral surface with two stripes along VL carinae, connected to lateral pigmentation in posterior half, additionally with 1215 spots; VM carina discontinuously pigmented medially (
<figureCitation id="132B2A5FFF9FF139FEA800DD72B6FC09" box="[365,461,925,949]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="37" pageNumber="38">fig. 20E</figureCitation>
). Cheliceral manus and fingers usually markedly pigmented, less so in specimens from Caleta Santa. Pedipalp femur pigmented along DI and DE carinae, and near articulations, densely so in some specimens; internal surfaces spotted; external surface with stripe along EM carinae. Patella markedly spotted on dorsal, internal, and external surfaces. Legs pigmented on prolateral surfaces.
</paragraph>
<paragraph id="8BAF36DAFF9FF139FFB7078772DFFAA7" blockId="37.[88,614,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FFB70787739CFB63" box="[114,231,1223,1247]" italics="true" pageId="37" pageNumber="38">Chelicerae</emphasis>
: Typical of genus; surfaces smooth; dorsal surface with two macrosetae situated near base of fingers.
</paragraph>
<paragraph id="8BAF36DAFF9FF139FFB7066172C7F99B" blockId="37.[88,614,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FFB7066173A7FA85" box="[114,220,1313,1337]" italics="true" pageId="37" pageNumber="38">Carapace</emphasis>
: Anterior margin with weak median projection and 89 macrosetae; surfaces coarsely granular, especially where pigmented, except for anterior third which is smooth; anteromedian longitudinal sulcus obsolete, bordered by few granules posteriorly; posteromedian longitudinal and posterolateral sulci well developed; median ocular sulcus obsolete; ocular tubercle well developed.
</paragraph>
<paragraph id="8BAF36DAFF9FF139FFB7056D7008FDD1" blockId="37.[88,614,211,1754]" lastBlockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FFB7056D73A6F9F9" box="[114,221,1581,1605]" italics="true" pageId="37" pageNumber="38">Pedipalps</emphasis>
: Femur with VI, DI, and DE carinae complete, granular; VM present in proximal half of segment; VE vestigial (
<figureCitation id="132B2A5FFF9FF139FFA505C673C6F922" box="[96,189,1670,1694]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="37" pageNumber="38">fig. 19E</figureCitation>
); dorsal surface with few scattered granules; internal surface with prominent granules medially; ventral surfaces smooth. Patella with DI and VI carinae complete, granular; DPP and VPP with prominent spiniform granules proximally (
<figureCitation id="132B2A5FFF9FF139FBFC024E77EDFE9A" box="[1081,1174,270,294]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="37" pageNumber="38">fig. 19F</figureCitation>
). Chela robust, with relatively elongated fingers; surfaces smooth and acarinate (
<figureCitation id="132B2A5FFF9FF139FBD0020A77EDFEDE" box="[1045,1174,330,354]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="37" pageNumber="38">fig. 19BD</figureCitation>
); fixed finger of adult ♂ strongly curved and lobed, creating distinct proximal gap with movable finger when fingers are closed, which is weakly developed in
<specimenCount id="9D16FD53FF9FF139FC7E02FE70ABFE66" box="[955,976,446,474]" pageId="37" pageNumber="38" type="female"></specimenCount>
. Movable finger, median denticle row comprising six subrows; one or two internal and external accessory denticles flanking subrows II and III; distal three subrows without external accessory denticles (
<figureCitation id="132B2A5FFF9FF139FCCD0115701CFDD1" box="[776,871,597,621]" captionStart="Fig" captionStartId="24.[151,189,1506,1528]" captionTargetBox="[231,1124,210,1472]" captionTargetId="figure-1@24.[231,1124,210,1472]" captionText="Fig. 20. Hadruroides Pocock, 1893, diagnostic characters. A, B. Hadruroides juanchaparroi, n. sp., paratype ♀ (MHNC). A. Telson, lateral aspect. B. Metasomal segment V, lateral aspect. C. Hadruroides lunatus (L. Koch, 1867), ♀ (AMNH), tergite V, dorsal aspect showing pigmentation pattern. DF. H. juanchaparroi, paratype ♂ (MHNC). D. Tergite IV, dorsal aspect showing pigmentation pattern. E. Sternite VII and metasomal segments IV, ventral aspect showing pigmentation pattern. F. Dextral pedipalp chela, movable finger, dorsal aspect showing dentition. G. Hadruroides geckoi, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. H. Hadruroides vichayitos, n. sp., paratype ♂ (MHNC), dextral pedipalp chela, movable finger, dorsal aspect showing dentition. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713962" httpUri="https://zenodo.org/record/5713962/files/figure.png" pageId="37" pageNumber="38">fig. 20F</figureCitation>
).
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD7501347745FC9C" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD75013470CAFD30" box="[688,945,628,652]" italics="true" pageId="37" pageNumber="38">Trichobothrial pattern</emphasis>
: Typical of genus; femur with three trichobothria, patella with 20, chela with 26; chelal trichobothrium
<emphasis id="B964EAC8FF9FF139FB4E01EF77DFFD7B" box="[1163,1188,687,711]" italics="true" pageId="37" pageNumber="38">eb</emphasis>
situated in line with proximal gap between fixed and movable fingers (♂); trichobothrium
<emphasis id="B964EAC8FF9FF139FD53004871CAFC9C" box="[662,689,776,800]" italics="true" pageId="37" pageNumber="38">Et</emphasis>
<subScript id="1794349FFF9FF139FD75005271C1FC9E" attach="left" box="[688,698,786,802]" fontSize="7" pageId="37" pageNumber="38">5</subScript>
situated level with
<emphasis id="B964EAC8FF9FF139FC59004870CCFC9C" box="[924,951,776,800]" italics="true" pageId="37" pageNumber="38">Et</emphasis>
<subScript id="1794349FFF9FF139FC72005270BAFC9E" attach="left" box="[951,961,786,802]" fontSize="7" pageId="37" pageNumber="38">4</subScript>
(
<figureCitation id="132B2A5FFF9FF139FC140048774AFC9C" box="[977,1073,776,800]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="37" pageNumber="38">fig. 19C</figureCitation>
).
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD7500667043FBDB" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD750066719EFC82" box="[688,741,806,830]" italics="true" pageId="37" pageNumber="38">Legs</emphasis>
: Prolateral surfaces granular; retrolateral surfaces smooth. Leg III, femur tetracarinate, DI and DE carinae obsolete, VI and EM well developed; patella, DE carinae comprising few granules in proximal third of segment, DM comprising few granules proximally, DI present in distal half, IM comprising four or five granules medially, VI comprising few granules distally, EM absent. Telotarsus with 813 ventromedian spinule clusters (setaceous tufts).
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD75072E70F9FB40" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD75072E7070FB3A" box="[688,779,1134,1158]" italics="true" pageId="37" pageNumber="38">Tergites</emphasis>
:
<materialsCitation id="3B783C87FF9FF139FCD8072D7005FB40" ID-GBIF-Occurrence="3421081351" collectionCode="VI, VII" collectorName="Post-tergites I" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1">
Pretergites finely granular (♂) or smooth (
<specimenCount id="9D16FD53FF9FF139FD3907C8706AFB18" box="[764,785,1160,1188]" pageId="37" pageNumber="38" type="female"></specimenCount>
).
<collectorName id="26E5530CFF9FF139FCED07CB70BDFB1F" box="[808,966,1163,1187]" pageId="37" pageNumber="38">Post-tergites I</collectorName>
<collectionCode id="ED01AE1FFF9FF139FC1307CB708EFB1F" box="[982,1013,1163,1187]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="37" pageNumber="38">VI</collectionCode>
, surfaces finely granular, slightly more so laterally;
<collectionCode id="ED01AE1FFF9FF139FBB907E977DFFB7D" box="[1148,1188,1193,1217]" pageId="37" pageNumber="38">VII</collectionCode>
coarsely granular, with four well-developed longitudinal carinae
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD75064377DFFAEA" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD7506437070FAA7" box="[688,779,1283,1307]" italics="true" pageId="37" pageNumber="38">Sternum</emphasis>
: Subpentagonal; surface granular medially, with 6 macrosetae; posterolateral surfaces granular; median sulcus well developed.
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD75061D7074FA2E" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD75061D7077FAC9" box="[688,780,1373,1397]" italics="true" pageId="37" pageNumber="38">Pectines</emphasis>
: Pectinal tooth count: 1618 (♂), 1111 (
<specimenCount id="9D16FD53FF9FF139FD2F06377184FA2F" box="[746,767,1399,1427]" pageId="37" pageNumber="38" type="female"></specimenCount>
).
</paragraph>
<paragraph id="8BAF36DAFF9FF139FD7506D97194F9F9" blockId="37.[662,1188,211,1754]" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD7506D97069FA0D" box="[688,786,1433,1457]" italics="true" pageId="37" pageNumber="38">Sternites</emphasis>
: Sternites IIIVI, surfaces smooth (
<specimenCount id="9D16FD53FF9FF139FD5B06F371C8FA73" box="[670,691,1459,1487]" pageId="37" pageNumber="38" type="female"></specimenCount>
) or becoming slightly matte laterally (♂); spiracles narrow, situated in posterior half of segment; VII, surface smooth, acarinate, only a few small granules evident in position of VL carinae.
</paragraph>
<paragraph id="8BAF36DAFF9FF13AFD75050B7136FDD1" blockId="37.[662,1188,211,1754]" lastBlockId="38.[128,654,211,1754]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="B964EAC8FF9FF139FD75050B705EF9DF" box="[688,805,1611,1635]" italics="true" pageId="37" pageNumber="38">Metasoma</emphasis>
: Segments IIV, dorsal surfaces coarsely granular, less so on IV; DL and ML carinae complete; LIM carinae complete on segment I, restricted to posterior half of II, and posterior third of III, absent on IV; surfaces between DL and ML carinae granular on segments IIII; VL carinae obsolete, smooth on segments IIII, comprising few granules on IV (
<figureCitation id="132B2A5FFF9CF13AFEF0026C72EDFEF8" box="[309,406,300,324]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="38" pageNumber="39">fig. 19A</figureCitation>
); VSM carinae absent on all segments. Segment V short (
<figureCitation id="132B2A5FFF9CF13AFDDB020A73E3FEC3" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="38" pageNumber="39">figs. 19A, G</figureCitation>
, 20B); DL carinae complete; VL carinae complete but well developed in posterior three-quarters of segment; VM carinae complete but obscured by granulation of ventral surface; dorsal and lateral surfaces smooth. Segment I with two pairs of ventral setae; II and III each with three pairs; IV with five pairs; V with 1115 ventral setae and 47 additional setae along posterior margin.
</paragraph>
<paragraph id="8BAF36DAFF9CF13AFF5F013471F2FD7B" blockId="38.[128,654,211,1754]" pageId="38" pageNumber="39">
<emphasis id="B964EAC8FF9CF13AFF5F01347398FD30" box="[154,227,628,652]" italics="true" pageId="38" pageNumber="39">Telson</emphasis>
: Vesicle, surfaces sparsely setose, ventral surface sparsely granular anteriorly (♂) or densely granular (
<specimenCount id="9D16FD53FF9CF13AFE6601EC72C3FD74" box="[419,440,684,712]" pageId="38" pageNumber="39" type="female"></specimenCount>
) (
<figureCitation id="132B2A5FFF9CF13AFE1401EF7140FD7B" box="[465,571,687,711]" captionStart="Fig" captionStartId="23.[112,150,1616,1638]" captionTargetBox="[152,1124,210,1582]" captionTargetId="figure-1@23.[152,1124,210,1582]" captionText="Fig. 19. Hadruroides juanchaparroi, n. sp., holotype ♂ (MHNC). A. Metasomal segments IV, V and telson, lateral aspect. B. Dextral pedipalp chela, ventral aspect. C. Dextral pedipalp chela, external aspect. D. Dextral pedipalp chela, ventrointernal aspect. E. Dextral pedipalp femur, dorsal aspect. F. Dextral pedipalp patella, dorsal aspect. G. Metasomal segment V, ventral aspect. Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5713960" httpUri="https://zenodo.org/record/5713960/files/figure.png" pageId="38" pageNumber="39">figs. 19A</figureCitation>
, 20A).
</paragraph>
<paragraph id="8BAF36DAFF9CF13AFF5F018E72E3FC82" blockId="38.[128,654,211,1754]" pageId="38" pageNumber="39">
<emphasis id="B964EAC8FF9CF13AFF5F018E7201FD5A" box="[154,378,718,742]" italics="true" pageId="38" pageNumber="39">Hemispermatophore</emphasis>
: Distal lamina slender, slightly curved, with apex rounded; crest equal to or less than half lamina length; basal portion slender (
<figureCitation id="132B2A5FFF9CF13AFEFA006672F0FC82" box="[319,395,806,830]" captionStart="Fig" captionStartId="36.[151,189,1044,1066]" captionTargetBox="[184,1170,210,1010]" captionTargetId="figure-297@36.[184,1171,210,1010]" captionText="Fig. 21. Hadruroides juanchaparroi, n. sp., paratype ³ (MHNC), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. D. Ventral aspect. Scale bar 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5713964" httpUri="https://zenodo.org/record/5713964/files/figure.png" pageId="38" pageNumber="39">fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BAF36DAFF9CF13AFF5F00057103FA13" blockId="38.[128,654,211,1754]" pageId="38" pageNumber="39">
<emphasis id="B964EAC8FF9CF13AFF5F00057278FCE1" box="[154,259,837,861]" italics="true" pageId="38" pageNumber="39">Variation</emphasis>
: Total length: ♂, 31.935.8 (mean
<emphasis id="B964EAC8FF9CF13AFF45002473EDFCC6" bold="true" box="[128,150,868,890]" italics="true" pageId="38" pageNumber="39">=</emphasis>
33.8,
<emphasis id="B964EAC8FF9CF13AFF2A00237258FCC6" box="[239,291,867,891]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFEC800247258FCC6" bold="true" box="[269,291,868,890]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7);
<specimenCount id="9D16FD53FF9CF13AFEA100207202FCC0" box="[356,377,864,892]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 33.6. Pedipalp chela, length:width ratio: ♂, 3.293.70 (mean
<emphasis id="B964EAC8FF9CF13AFDB200C271F6FC24" bold="true" box="[631,653,898,920]" italics="true" pageId="38" pageNumber="39">=</emphasis>
3.48,
<emphasis id="B964EAC8FF9CF13AFF0500DE7394FC0A" box="[192,239,926,950]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFF1C00E07394FC0A" bold="true" box="[217,239,928,950]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7);
<specimenCount id="9D16FD53FF9CF13AFEE700DB724CFC0B" box="[290,311,923,951]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 3.49; length:height ratio: ♂, 3.003.34 (mean
<emphasis id="B964EAC8FF9CF13AFE9500FD721DFC6F" bold="true" box="[336,358,957,979]" italics="true" pageId="38" pageNumber="39">=</emphasis>
3.18,
<emphasis id="B964EAC8FF9CF13AFE7900FC7294FC6F" box="[444,495,956,980]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE1C00FD7294FC6F" bold="true" box="[473,495,957,979]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7);
<specimenCount id="9D16FD53FF9CF13AFDE900F9713AFC69" box="[556,577,953,981]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 3.32. Pedipalp femur, length:width ratio: ♂, 3.00 3.33 (mean
<emphasis id="B964EAC8FF9CF13AFECF00B9725BFBB3" bold="true" box="[266,288,1017,1039]" italics="true" pageId="38" pageNumber="39">=</emphasis>
3.17,
<emphasis id="B964EAC8FF9CF13AFEAE00B872E1FBB3" box="[363,410,1016,1040]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE4100B972E1FBB3" bold="true" box="[388,410,1017,1039]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7);
<specimenCount id="9D16FD53FF9CF13AFE0900B4729AFBAC" box="[460,481,1012,1040]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 3.00. Pectinal tooth count: ♂ (
<emphasis id="B964EAC8FF9CF13AFEFD07557219FB91" box="[312,354,1045,1069]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE8907577219FB91" bold="true" box="[332,354,1047,1069]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
14), 16 (
<emphasis id="B964EAC8FF9CF13AFE0107557295FB91" box="[452,494,1045,1069]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE1D07577295FB91" bold="true" box="[472,494,1047,1069]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
3), 17 (10), 18 (1);
<materialsCitation id="3B783C87FF9CF13AFF7A07707269FB3A" ID-GBIF-Occurrence="3421081366" collectionCode="V" latitude="5.116667" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D16FD53FF9CF13AFF7A077073AFFBF0" box="[191,212,1072,1100]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 11 (2). Metasomal segment
<collectionCode id="ED01AE1FFF9CF13AFDB6077371F2FBF7" box="[627,649,1075,1099]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="39" type="Museum">V</collectionCode>
, length:width ratio: ♂, 2.042.33 (mean
<emphasis id="B964EAC8FF9CF13AFDB2071271F6FBD4" bold="true" box="[631,653,1106,1128]" italics="true" pageId="38" pageNumber="39">=</emphasis>
2.14,
<emphasis id="B964EAC8FF9CF13AFF05072E7394FB3A" box="[192,239,1134,1158]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFF1C07307394FB3A" bold="true" box="[217,239,1136,1158]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7)
</materialsCitation>
;
<specimenCount id="9D16FD53FF9CF13AFEE7072B724CFB3B" box="[290,311,1131,1159]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 1.97; length:height ratio: ♂, 2.092.29 (mean
<emphasis id="B964EAC8FF9CF13AFE9507CE721DFB18" bold="true" box="[336,358,1166,1188]" italics="true" pageId="38" pageNumber="39">=</emphasis>
2.19,
<emphasis id="B964EAC8FF9CF13AFE7907CC7294FB18" box="[444,495,1164,1188]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE1C07CE7294FB18" bold="true" box="[473,495,1166,1188]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7);
<specimenCount id="9D16FD53FF9CF13AFDE907C9713AFB19" box="[556,577,1161,1189]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 2.02; number of setae: dorsolateral (
<emphasis id="B964EAC8FF9CF13AFE1807EA7172FB7D" box="[477,521,1194,1218]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE3607EB7172FB7D" bold="true" box="[499,521,1195,1217]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
16): 5 (
<emphasis id="B964EAC8FF9CF13AFDA707EA71F6FB7D" box="[610,653,1194,1218]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFDB207EB71F6FB7D" bold="true" box="[631,653,1195,1217]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
1), 6 (5), 8 (9), 9 (1); lateral (
<emphasis id="B964EAC8FF9CF13AFE73078872A4FB63" box="[438,479,1224,1248]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE0C078972A4FB63" bold="true" box="[457,479,1225,1247]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
16): 5 (5), 6 (8), 7 (3); ventrolateral (
<emphasis id="B964EAC8FF9CF13AFEA507A572F0FB41" box="[352,395,1253,1277]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFEB007A772F0FB41" bold="true" box="[373,395,1255,1277]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
16): 8 (9), 9 (6), 10 (1); ventral (
<emphasis id="B964EAC8FF9CF13AFF1B06437272FAA7" box="[222,265,1283,1307]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFF3606457272FAA7" bold="true" box="[243,265,1285,1307]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
7): 11 (3), 12 (1), 13 (1), 14 (1), 15 (1). Telson, length:height ratio: ♂, 3.153.40 (mean
<emphasis id="B964EAC8FF9CF13AFF1606007392FAEA" bold="true" box="[211,233,1344,1366]" italics="true" pageId="38" pageNumber="39">=</emphasis>
3.26,
<emphasis id="B964EAC8FF9CF13AFEFF067F7210FAEA" box="[314,363,1343,1367]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE9006007210FAEA" bold="true" box="[341,363,1344,1366]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
6);
<materialsCitation id="3B783C87FF9CF13AFE61067B72D5FA2E" ID-GBIF-Occurrence="3421081337" collectionCode="III" latitude="5.266667" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D16FD53FF9CF13AFE61067B72C2FAEB" box="[420,441,1339,1367]" pageId="38" pageNumber="39" type="female"></specimenCount>
, 3.06. Telotarsus, number of ventromedian spinule clusters (setaceous tufts):
<collectionCode id="ED01AE1FFF9CF13AFEE3063A723EFA2E" box="[294,325,1402,1426]" pageId="38" pageNumber="39">III</collectionCode>
(
<emphasis id="B964EAC8FF9CF13AFE90063A72FBFA2D" box="[341,384,1402,1426]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFEAF063B72FBFA2D" bold="true" box="[362,384,1403,1425]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
16)
</materialsCitation>
, 8 (
<emphasis id="B964EAC8FF9CF13AFE1D063A7178FA2D" box="[472,515,1402,1426]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE28063B7178FA2D" bold="true" box="[493,515,1403,1425]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
1),
<materialsCitation id="3B783C87FF9CF13AFDF5063A72FFFA13" ID-GBIF-Occurrence="3421081347" collectionCode="IV" latitude="5.25" pageId="38" pageNumber="39" specimenCount="1">
9 (3), 10 (9), 11 (3);
<collectionCode id="ED01AE1FFF9CF13AFF3806D77261FA13" box="[253,282,1431,1455]" pageId="38" pageNumber="39">IV</collectionCode>
(
<emphasis id="B964EAC8FF9CF13AFEEE06D8722DFA13" box="[299,342,1432,1456]" italics="true" pageId="38" pageNumber="39">
n
<emphasis id="B964EAC8FF9CF13AFE8506D9722DFA13" bold="true" box="[320,342,1433,1455]" italics="true" pageId="38" pageNumber="39">=</emphasis>
</emphasis>
15)
</materialsCitation>
, 11 (7), 12 (3), 13 (5).
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FF9CF13AFF5F06F6726BF9DE" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BAF36DAFF9CF13AFF5F06F6726BF9DE" blockId="38.[128,654,211,1754]" pageId="38" pageNumber="39">
DISTRIBUTION: This species is presently known from only two localities in northern
<collectingCountry id="F307764AFF9CF13AFF4506B173C3F9B5" box="[128,184,1521,1545]" name="Peru" pageId="38" pageNumber="39">Peru</collectingCountry>
, Cerro Campana (
<quantity id="4CE89B3FFF9CF13AFE6706B17289F9B5" box="[418,498,1521,1545]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="38" pageNumber="39" unit="m" value="450.0">450 m</quantity>
) and Caleta Salta (
<quantity id="4CE89B3FFF9CF13AFF0C054F727FF99B" box="[201,260,1551,1575]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="38" pageNumber="39" unit="m" value="13.0">13 m</quantity>
), close to the coast. Both localities occur in the Pacific desert ecoregion (
<bibRefCitation id="EF814B2BFF9CF13AFD87056D73C3F9DE" author="Brack, A." pageId="38" pageNumber="39" pagination="175 - 319" refId="ref28821" refString="Brack, A. 1986. Ecologia de un pais complejo. In Gran Geografia del Peru: Naturaleza y Hombre. Barcelona: Manfer-Juan Mejia Baca, 2: 175 - 319." type="journal article" year="1986">Brack, 1986</bibRefCitation>
; fig. 2).
</paragraph>
</subSubSection>
<subSubSection id="C30A6551FF9CF13AFF5F052977B7FEC2" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="8BAF36DAFF9CF13AFF5F052977B7FEC2" blockId="38.[128,654,211,1754]" lastBlockId="38.[701,1228,211,382]" pageId="38" pageNumber="39">
ECOLOGY:
<taxonomicName id="4C104D59FF9CF13AFEE70529712FF93D" authorityName="Ochoa &amp; Prendini" authorityYear="2010" box="[290,596,1641,1665]" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="juanchaparroi">
<emphasis id="B964EAC8FF9CF13AFEE70529712FF93D" box="[290,596,1641,1665]" italics="true" pageId="38" pageNumber="39">Hadruroides juanchaparroi</emphasis>
</taxonomicName>
was collected at night with UV light detection. At the
<typeStatus id="54AB8878FF9CF13AFF6B05E473A5F900" box="[174,222,1700,1724]" pageId="38" pageNumber="39">type</typeStatus>
locality (Cerro Campana), a typical Lomas biotope (
<bibRefCitation id="EF814B2BFF9CF13AFE9405827296F966" author="Pefaur, J. E." box="[337,493,1730,1754]" pageId="38" pageNumber="39" pagination="31 - 42" refId="ref30935" refString="Pefaur, J. E. 1981. Composition and phenology of epigeic animal communities in the Lomas of southern Peru. Journal of Arid Environments 4: 31 - 42." type="journal article" year="1981">Péfaur, 1981</bibRefCitation>
) with shrub vegetation and columnar cacti (fig. 3C), specimens were observed climbing on terrestrial bromeliads (
<taxonomicName id="4C104D59FF9CF13AFC96024E708AFE9A" box="[851,1009,270,294]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
), a behavior previously described for
<taxonomicName id="4C104D59FF9CF13AFC74026B7009FEDC" authority="(Francke and Soleglad, 1980)" baseAuthorityName="Francke and Soleglad" baseAuthorityYear="1980" class="Arachnida" family="Caraboctonidae" genus="Hadruroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="aguilari">
<emphasis id="B964EAC8FF9CF13AFC74026B7750FEFF" box="[945,1067,299,323]" italics="true" pageId="38" pageNumber="39">H. aguilari</emphasis>
(
<bibRefCitation id="EF814B2BFF9CF13AFBFF026B7011FEDC" author="Francke, O. F. &amp; M. E. Soleglad" pageId="38" pageNumber="39" pagination="1 - 13" refId="ref29871" refString="Francke, O. F., and M. E. Soleglad. 1980. Two new Hadruroides Pocock from Peru (Scorpiones, Vaejovidae). Occasional Papers of the Museum, Texas Tech University 69: 1 - 13." type="journal article" year="1980">Francke and Soleglad, 1980</bibRefCitation>
)
</taxonomicName>
. At Caleta Santa, the species was collected in an area with shrub vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>