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<mods:title id="C30F1867A56CBD0B5D763E7BB45E5904">Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species</mods:title>
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<mods:namePart id="9B9EE063D8815B94D1C2AA86B4407E9F">Nguyen, Anh D.</mods:namePart>
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<treatment id="03B987C4FFEBAE5CFC99F979FDC9AA77" ID-DOI="http://doi.org/10.5281/zenodo.8065871" ID-GBIF-Taxon="210357023" ID-Zenodo-Dep="8065871" LSID="urn:lsid:plazi:treatment:03B987C4FFEBAE5CFC99F979FDC9AA77" httpUri="http://treatment.plazi.org/id/03B987C4FFEBAE5CFC99F979FDC9AA77" lastPageId="3" lastPageNumber="3" pageId="1" pageNumber="2">
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<paragraph id="8BAF36D2FFEBAE5EFC99F979FAC3ADD4" blockId="1.[861,1365,1660,1782]" box="[873,1353,1756,1782]" pageId="1" pageNumber="2">
<emphasis id="B964EAC0FFEBAE5EFC99F979FAC3ADD4" bold="true" box="[873,1353,1756,1782]" pageId="1" pageNumber="2">
Genus
<taxonomicName id="4C104D51FFEBAE5EFC33F979FAC3ADD4" ID-CoL="7PL2Z" ID-ENA="614353" authority="Cook &amp; Loomis, 1924" authorityName="Cook &amp; Loomis" authorityYear="1924" box="[963,1353,1756,1782]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC0FFEBAE5EFC33F979FBBEADD4" bold="true" box="[963,1076,1756,1781]" italics="true" pageId="1" pageNumber="2">Hylomus</emphasis>
<bibRefCitation id="EF814B23FFEBAE5EFBCBF979FAC3ADD4" author="Cook OF &amp; Loomis HF" box="[1083,1353,1756,1782]" pageId="1" pageNumber="2" pagination="103 - 108" refId="ref11518" refString="Cook OF, Loomis HF. 1924. A new family of spined millipeds from central China. Journal of the Washington Academy of Sciences 14: 103 - 108." type="journal article" year="1924">Cook &amp; Loomis, 1924</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BAF36D2FFEBAE5EFCDDF8B9FB15AC5E" blockId="1.[813,1415,1820,1920]" pageId="1" pageNumber="2">
<taxonomicName id="4C104D51FFEBAE5EFCDDF8B9FBE7AC0E" ID-CoL="7PL2Z" authority="Cook &amp; Loomis, 1924: 105" authorityName="Cook &amp; Loomis" authorityPageNumber="105" authorityYear="1924" box="[813,1133,1820,1839]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC0FFEBAE5EFCDDF8B9FCF3AC0E" box="[813,889,1820,1839]" italics="true" pageId="1" pageNumber="2">Hylomus</emphasis>
<bibRefCitation id="EF814B23FFEBAE5EFC8EF8B9FBE7AC0E" author="Cook OF &amp; Loomis HF" box="[894,1133,1820,1839]" pageId="1" pageNumber="2" pagination="103 - 108" refId="ref11518" refString="Cook OF, Loomis HF. 1924. A new family of spined millipeds from central China. Journal of the Washington Academy of Sciences 14: 103 - 108." type="journal article" year="1924">Cook &amp; Loomis, 1924: 105</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF814B23FFEBAE5EFB85F8B9FA9AAC0E" author="Attems C." box="[1141,1296,1820,1839]" pageId="1" pageNumber="2" pagination="1 - 300" refId="ref11356" refString="Attems C. 1937. Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae. Das Tierreich 68: 1 - 300." type="journal article" year="1937">Attems 1937: 126</bibRefCitation>
;
<bibRefCitation id="EF814B23FFEBAE5EFAEAF8B9FCFBAC6B" author="Jeekel CAW" pageId="1" pageNumber="2" refId="ref11849" refString="Jeekel CAW. 1968. On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida). Bronder-Offset Rotterdam." type="book" year="1968">Jeekel 1968: 51</bibRefCitation>
;
<bibRefCitation id="EF814B23FFEBAE5EFC8CF892FB98AC6B" author="Jeekel CAW" box="[892,1042,1847,1866]" pageId="1" pageNumber="2" pagination="651 - 670" refId="ref11873" refString="Jeekel CAW. 1980. The generic allocation of some little-known Paradoxosomatidae from South-ast Asia (Diplopoda, Polydesmida). Revue suisse de zoologie 87: 651 - 670." type="journal article" year="1980">Jeekel 1980: 655</bibRefCitation>
;
<bibRefCitation id="EF814B23FFEBAE5EFBECF892FA8CAC6B" author="Srisonchai R &amp; Enghoff H &amp; Likhitrakarn N &amp; Panha S." box="[1052,1286,1847,1866]" pageId="1" pageNumber="2" pagination="1 - 177" refId="ref12093" refString="Srisonchai R, Enghoff H, Likhitrakarn N, Panha S. 2018 a. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zookeys 761: 1 - 177. doi: 10.3897 / zookeys. 761.24214." type="journal article" year="2018">Srisonchai et al. 2018a: 15</bibRefCitation>
. Synonymized with
<taxonomicName id="4C104D51FFEBAE5EFC78F8F7FC7AAC45" ID-CoL="7P8ZX" authorityName="Chamberlin" authorityYear="1923" box="[904,1008,1874,1892]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC0FFEBAE5EFC78F8F7FC7AAC45" box="[904,1008,1874,1892]" italics="true" pageId="1" pageNumber="2">Desmoxytes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF814B23FFEBAE5EFBE5F8F4FAB5AC45" author="Golovatch SI &amp; Enghoff H." box="[1045,1343,1873,1893]" pageId="1" pageNumber="2" pagination="45 - 71" refId="ref11598" refString="Golovatch SI, Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article" year="1994">Golovatch &amp; Enghoff (1994: 46)</bibRefCitation>
, but re-validated by
<bibRefCitation id="EF814B23FFEBAE5EFC38F8C9FB11AC5E" author="Srisonchai R &amp; Enghoff H &amp; Likhitrakarn N &amp; Panha S." box="[968,1179,1900,1920]" pageId="1" pageNumber="2" pagination="1 - 177" refId="ref12093" refString="Srisonchai R, Enghoff H, Likhitrakarn N, Panha S. 2018 a. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zookeys 761: 1 - 177. doi: 10.3897 / zookeys. 761.24214." type="journal article" year="2018">Srisonchai et al. (2018a)</bibRefCitation>
.
</paragraph>
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<footnote id="E80B2ADCFFEBAE5EFF78F840FE12ACD6" box="[136,408,2020,2039]" pageId="1" pageNumber="2">
<paragraph id="8BAF36D2FFEBAE5EFF78F840FE12ACD6" blockId="1.[136,408,2020,2039]" box="[136,408,2020,2039]" pageId="1" pageNumber="2">© 2019 Academia Sinica, Taiwan</paragraph>
</footnote>
<subSubSection id="C30A6559FFE8AE5DFF1AFF5BFE64AA14" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="8BAF36D2FFE8AE5DFF1AFF5BFE64AA14" blockId="2.[174,774,253,469]" pageId="2" pageNumber="3">
<emphasis id="B964EAC0FFE8AE5DFF1AFF5BFEF0AA34" box="[234,378,254,277]" italics="true" pageId="2" pageNumber="3">
<typeStatus id="54AB8870FFE8AE5DFF1AFF5BFE94AA34" box="[234,286,254,277]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C104D51FFE8AE5DFE7CFF58FF6CAA17" ID-CoL="3NGCV" authority="Cook &amp; Loomis, 1924" authorityName="Cook &amp; Loomis" authorityYear="1924" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="draco">
<emphasis id="B964EAC0FFE8AE5DFE7CFF58FDB1AA34" box="[396,571,253,277]" italics="true" pageId="2" pageNumber="3">Hylomus draco</emphasis>
<bibRefCitation id="EF814B23FFE8AE5DFDB5FF58FF6CAA17" author="Cook OF &amp; Loomis HF" pageId="2" pageNumber="3" pagination="103 - 108" refId="ref11518" refString="Cook OF, Loomis HF. 1924. A new family of spined millipeds from central China. Journal of the Washington Academy of Sciences 14: 103 - 108." type="journal article" year="1924">Cook &amp; Loomis, 1924</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE8AE5DFF1AFE9BFC48AA94" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BAF36D2FFE8AE5DFF1AFE9BFC48AA94" blockId="2.[174,774,253,469]" lastBlockId="2.[850,1451,253,470]" pageId="2" pageNumber="3">
<emphasis id="B964EAC0FFE8AE5DFF1AFE9BFEE9AA74" box="[234,355,318,341]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
: The genus can be recognised by a combination of characters: small to large size (
<quantity id="4CE89B37FFE8AE5DFF46FED8FEB8AAB4" box="[182,306,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.5" metricValueMin="1.2" pageId="2" pageNumber="3" unit="mm" value="23.5" valueMax="35.0" valueMin="12.0">1235 mm</quantity>
long); often brightly coloured; paraterga strongly to exceedingly well developed, wing-, spine- and antler-shaped; metazonal surface often more or less roughly granulate, rarely smooth; epiproct of telson sometimes modified with hypertrophied setiferous tubercles near tip; male leg femora 5/6/7/9 often inflated or humped ventrally; gonopod suberect, seldom subfalcate; solenophore and solenomere usually shortened.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE8AE5CFC7FFE18FDC9AA77" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BAF36D2FFE8AE5CFC7FFE18FDC9AA77" blockId="2.[850,1451,253,470]" lastBlockId="3.[136,736,253,342]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B964EAC0FFE8AE5DFC7FFE18FC7FAAF4" box="[911,1013,445,469]" italics="true" pageId="2" pageNumber="3">Remarks</emphasis>
: The genus is widely distributed in southern
<collectingCountry id="F3077642FFE9AE5CFF03FF58FEBFAA34" box="[243,309,253,277]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
and
<collectingCountry id="F3077642FFE9AE5CFE80FF58FE58AA34" box="[368,466,253,277]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
. Few species are found in
<collectingCountry id="F3077642FFE9AE5CFF58FEBBFF54AA14" box="[168,222,286,309]" name="Laos" pageId="3" pageNumber="4">Laos</collectingCountry>
and
<collectingCountry id="F3077642FFE9AE5CFEEAFEB8FE08AA14" box="[282,386,285,309]" name="Thailand" pageId="3" pageNumber="4">Thailand</collectingCountry>
. The distribution of the genus
<taxonomicName id="4C104D51FFE9AE5CFF78FE98FF6DAA74" authorityName="Cook &amp; Loomis" authorityYear="1924" box="[136,231,317,341]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAC0FFE9AE5CFF78FE98FF6DAA74" box="[136,231,317,341]" italics="true" pageId="3" pageNumber="4">Hylomus</emphasis>
</taxonomicName>
in
<collectingCountry id="F3077642FFE9AE5CFEF8FE98FEE8AA74" box="[264,354,317,341]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
is shown in figure 1.
</paragraph>
</subSubSection>
</treatment>
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