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<document id="C3C877765E7F45155D37873B2296C413" ID-CLB-Dataset="56108" ID-DOI="10.6620/ZS.2019.58-14" ID-GBIF-Dataset="de0647eb-9b73-4bac-9ff7-f7bff7518dc0" ID-PMC="PMC6759935" ID-PubMed="31966315" ID-Zenodo-Dep="8065869" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="tatiana" checkinTime="1687182873363" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Nguyen, Manh-Ha, Nguyen, Thu-Anh T. &amp; Phung, Hong-Luong T." docDate="2019" docId="03B987C4FFE9AE5BFCB1FE58FD28AAD4" docLanguage="en" docName="ZoolStud.58.14.1-31.pdf" docOrigin="Zoological Studies 58 (14)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Hylomus cattienensis" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="FF80FFBCFFEAAE5FFFF0FFA5FF8AAB21" masterDocTitle="Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="4" updateTime="1699534267754" updateUser="ExternalLinkService">
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<mods:title id="927D62697716893CAB2050AED1890176">Review of Dragon Millipedes (Diplopoda, Polydesmida, Paradoxosomatidae) in the Fauna of Vietnam, with Descriptions of Three New Species</mods:title>
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<mods:namePart id="1291BFF6F0DD862AE13B1D5040ADD760">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="A452BB90DD49B94125B765EDF2BB981C">Nguyen, Manh-Ha</mods:namePart>
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<mods:namePart id="54DB403552581F44C2D3629C86583860">Nguyen, Thu-Anh T.</mods:namePart>
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<mods:namePart id="ABE7AC2046E2A7BD30C1319B9188077A">Phung, Hong-Luong T.</mods:namePart>
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<mods:title id="3B8B52591D2A61299F8F9886035C3419">Zoological Studies</mods:title>
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<mods:date id="9CACC55A7D819882D474EA4EC7584C2B">2019</mods:date>
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<treatment id="03B987C4FFE9AE5BFCB1FE58FD28AAD4" ID-DOI="http://doi.org/10.5281/zenodo.8065886" ID-GBIF-Taxon="210357016" ID-Zenodo-Dep="8065886" LSID="urn:lsid:plazi:treatment:03B987C4FFE9AE5BFCB1FE58FD28AAD4" httpUri="http://treatment.plazi.org/id/03B987C4FFE9AE5BFCB1FE58FD28AAD4" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="C30A6559FFE9AE5CFCB1FE58FB28A974" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BAF36D2FFE9AE5CFCB1FE58FB28A974" blockId="3.[833,1393,508,597]" pageId="3" pageNumber="4">
<heading id="D0E781BEFFE9AE5CFCB1FE58FB30A917" bold="true" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="4C104D51FFE9AE5CFCB1FE58FB30A917" ID-CoL="3NGCS" authority="(Nguyen, Golovatch &amp; Anichkin, 2005)" baseAuthorityName="Nguyen, Golovatch &amp; Anichkin" baseAuthorityYear="2005" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B964EAC0FFE9AE5CFCB1FE58FB30A917" bold="true" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFCB1FE58FBD9A937" bold="true" box="[833,1107,509,534]" italics="true" pageId="3" pageNumber="4">Hylomus cattienensis</emphasis>
(
<bibRefCitation id="EF814B23FFE9AE5CFB92FE58FAFBA937" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" box="[1122,1393,508,534]" pageId="3" pageNumber="4" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen, Golovatch &amp;</bibRefCitation>
Anichkin, 2005)
</emphasis>
</taxonomicName>
</heading>
(
<figureCitation id="132B2A57FFE9AE5CFBE8FD98FBE8A974" box="[1048,1122,573,597]" captionStart="Fig" captionStartId="2.[175,206,1813,1832]" captionTargetBox="[440,1185,515,1783]" captionTargetId="figure-96@2.[440,1185,486,1804]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Hylomus species in mainland Vietnam. 1: Hoang Lien National Park (Sa Pa, Lao Cai Province); 2: Nam Xay Commune (Van Ban District, Lao Cai Province); 3: Duc Xuan Commune (Bac Quang District, Ha Giang Province); 4: Xuan Son National Park (Phu Tho Province); 5: Tam Dao (Vinh Phuc Province); 6: Cuc Phuong National Park (Ninh Binh Province); 7: Phong Nha – Ke Bang National Park (Quang Binh Province); 8: Lien Chieu District (Da Nang Province); 9: Ba Na National Park (Da Nang Province); 10: Lo Xo Pass (Kon Tum Province); 11: Ngoc Linh Mt. (Kon Tum Province); 12: Kon Chu Rang Natural Reserve (Gia Lai Province); 13: Buon Ho District (Dak Lak Province); 14: Hon Ba Mt. (Khanh Hoa Province); 15: Phan Rang (Ninh Thuan Province); 16: Cat Tien National Park (Dong Nai Province)." figureDoi="http://doi.org/10.5281/zenodo.8065873" httpUri="https://zenodo.org/record/8065873/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="132B2A57FFE9AE5CFB9FFD98FB13A974" box="[1135,1177,573,597]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[174,205,1813,1832]" captionStartId-1="5.[136,167,954,973]" captionTargetBox-0="[174,1451,755,1785]" captionTargetBox-1="[283,1266,249,929]" captionTargetId-0="figure-318@4.[174,1451,755,1785]" captionTargetId-1="figure-5@5.[283,1266,249,929]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 3. Hylomus cattienensis (Nguyen, Golovatch &amp; Anichkin, 2005) from Cat Tien National Park. Anterior part, lateral view (A), dorsal view (B), ventral view (C); body rings 10−12, dorsal view (D), lateral view (E); posterior part, lateral view (F), ventral view (G); gonopods, ventral view (H); sternite 5, ventral view (I). Anterior part, lateral view (A), dorsal view (B), ventral view (C); body rings 10−12, dorsal view (D), lateral view (E); posterior part, lateral view (F), ventral view (G); gonopods, ventral view (H); sternite 5, ventral view (I). No scale bars were inserted while taking photographs for this figure. This figure emphasises colouration, shapes of paraterga (Pa), metaterga (Me), head (He), collum (Co), gonopods (Go), sternal tubercles (st), epiproct (Epi) and hypoproct (Hy)." captionText-1="Fig. 4. Hylomus cattienensis (Nguyen, Golovatch &amp; Anichkin, 2005) from Cat Tien National Park. Right gonopod, lateral view (A), mesal view (B). Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8065879" figureDoi-1="http://doi.org/10.5281/zenodo.8065881" httpUri-0="https://zenodo.org/record/8065879/files/figure.png" httpUri-1="https://zenodo.org/record/8065881/files/figure.png" pageId="3" pageNumber="4">3−4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE9AE5CFCDDFDDCFB5DA9E0" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BAF36D2FFE9AE5CFCDDFDDCFB87A987" blockId="3.[813,1414,632,705]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D51FFE9AE5CFCDDFDDCFC22A987" ID-CoL="6CNKJ" authority="Nguyen, Golovatch &amp; Anichkin, 2005: 252 - 254" authorityName="Nguyen, Golovatch &amp; Anichkin" authorityPageNumber="252 - 254" authorityYear="2005" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B964EAC0FFE9AE5CFCDDFDDCFB85A9AA" box="[813,1039,633,651]" italics="true" pageId="3" pageNumber="4">Desmoxytes cattienensis</emphasis>
<bibRefCitation id="EF814B23FFE9AE5CFBE8FDDCFC22A987" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" pageId="3" pageNumber="4" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen, Golovatch &amp; Anichkin, 2005: 252−254</bibRefCitation>
</taxonomicName>
, figs. 6−13.
</paragraph>
<paragraph id="8BAF36D2FFE9AE5CFCDDFD08FB5DA9E0" blockId="3.[813,1414,632,705]" box="[813,1239,685,705]" pageId="3" pageNumber="4">
<taxonomicName id="4C104D51FFE9AE5CFCDDFD08FB59A9E0" ID-CoL="3NGCS" authority=": Srisonchai et al. 2018 a: 10" authorityName="Srisonchai" authorityPageNumber="10" authorityYear="2018" box="[813,1235,685,705]" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="B964EAC0FFE9AE5CFCDDFD08FC69A9E1" box="[813,995,685,704]" italics="true" pageId="3" pageNumber="4">Hylomus cattienensis</emphasis>
:
<bibRefCitation id="EF814B23FFE9AE5CFC1EFD08FB59A9E0" author="Srisonchai R &amp; Enghoff H &amp; Likhitrakarn N &amp; Panha S." box="[1006,1235,685,705]" pageId="3" pageNumber="4" pagination="1 - 177" refId="ref12093" refString="Srisonchai R, Enghoff H, Likhitrakarn N, Panha S. 2018 a. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). Zookeys 761: 1 - 177. doi: 10.3897 / zookeys. 761.24214." type="journal article" year="2018">Srisonchai et al. 2018a: 10</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE9AE5CFC99FD4CFBA6A840" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BAF36D2FFE9AE5CFC99FD4CFBA6A840" blockId="3.[813,1414,745,993]" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFC99FD4CFBB2A820" box="[873,1080,745,769]" italics="true" pageId="3" pageNumber="4">Material examined</emphasis>
:
<specimenCount id="9D16FD5BFFE9AE5CFBB9FD4CFB19A820" box="[1097,1171,745,769]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
(IEBR-27)
<collectingRegion id="49D4F830FFE9AE5CFAEAFD4CFC05A800" country="Vietnam" name="Dong Nai" pageId="3" pageNumber="4">Dong Nai Province</collectingRegion>
, Cat Tien NP, forest,
<date id="FFAE1012FFE9AE5CFB87FCACFAC1A800" box="[1143,1355,777,801]" pageId="3" pageNumber="4" value="2004-09-23">23 September 2004</date>
, coll. Anh D. Nguyen;
<specimenCount id="9D16FD5BFFE9AE5CFC14FC8CFBA4A860" box="[996,1070,809,833]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
,
<specimenCount id="9D16FD5BFFE9AE5CFBC9FC8CFB1CA860" box="[1081,1174,809,833]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
(IEBR-40) same data, but
<date id="FFAE1012FFE9AE5CFCA5FCECFBA2A840" box="[853,1064,841,865]" pageId="3" pageNumber="4" value="2004-09-24">24 September 2004</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFE9AE5BFC99FCCFFE36AAB7" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BAF36D2FFE9AE5BFC99FCCFFE36AAB7" blockId="3.[813,1414,745,993]" lastBlockId="4.[174,774,253,501]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B964EAC0FFE9AE5CFC99FCCFFC6AA8A0" box="[873,992,874,897]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
: The species can be distinguished from its congeners by the combination of the following characters: colouration pink to reddish; paraterga antler-shaped; metaterga with a row of 1 + 1 short setae; male leg femora unmodified; epiproct modified with several setiferous knobs (
<figureCitation id="132B2A57FFEEAE5BFE83FEB8FE32AA17" box="[371,440,285,310]" captionStart="Fig" captionStartId="4.[174,205,1813,1832]" captionTargetBox="[174,1451,755,1785]" captionTargetId="figure-318@4.[174,1451,755,1785]" captionTargetPageId="4" captionText="Fig. 3. Hylomus cattienensis (Nguyen, Golovatch &amp; Anichkin, 2005) from Cat Tien National Park. Anterior part, lateral view (A), dorsal view (B), ventral view (C); body rings 10−12, dorsal view (D), lateral view (E); posterior part, lateral view (F), ventral view (G); gonopods, ventral view (H); sternite 5, ventral view (I). Anterior part, lateral view (A), dorsal view (B), ventral view (C); body rings 10−12, dorsal view (D), lateral view (E); posterior part, lateral view (F), ventral view (G); gonopods, ventral view (H); sternite 5, ventral view (I). No scale bars were inserted while taking photographs for this figure. This figure emphasises colouration, shapes of paraterga (Pa), metaterga (Me), head (He), collum (Co), gonopods (Go), sternal tubercles (st), epiproct (Epi) and hypoproct (Hy)." figureDoi="http://doi.org/10.5281/zenodo.8065879" httpUri="https://zenodo.org/record/8065879/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
); gonopod suberect; femorite, solenomere and solenophore subequal in length; postfemoral region not extremely short; tip of lamina medialis pointed (
<figureCitation id="132B2A57FFEEAE5BFE9EFED8FE25AAB7" box="[366,431,381,406]" captionStart="Fig" captionStartId="5.[136,167,954,973]" captionTargetBox="[283,1266,249,929]" captionTargetId="figure-5@5.[283,1266,249,929]" captionTargetPageId="5" captionText="Fig. 4. Hylomus cattienensis (Nguyen, Golovatch &amp; Anichkin, 2005) from Cat Tien National Park. Right gonopod, lateral view (A), mesal view (B). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8065881" httpUri="https://zenodo.org/record/8065881/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A6559FFEEAE5BFF1AFE3BFD28AAD4" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BAF36D2FFEEAE5BFF1AFE3BFDBFAAF4" blockId="4.[174,774,253,501]" pageId="4" pageNumber="5">
<emphasis id="B964EAC0FFEEAE5BFF1AFE3BFE2DAA94" box="[234,423,413,437]" italics="true" pageId="4" pageNumber="5">Previous records</emphasis>
:
<collectingRegion id="49D4F830FFEEAE5BFE49FE3BFD19AA94" box="[441,659,413,438]" country="Vietnam" name="Dong Nai" pageId="4" pageNumber="5">Dong Nai Province</collectingRegion>
(Cat Tien National Park) (
<bibRefCitation id="EF814B23FFEEAE5BFEA8FE1BFDA3AAF4" author="Nguyen AD &amp; Golovatch SI &amp; Anichkin AE" box="[344,553,445,470]" pageId="4" pageNumber="5" pagination="251 - 257" refId="ref12004" refString="Nguyen AD, Golovatch SI, Anichkin AE. 2005. The dragon millipedes in Vietnam (Polydesmida: Paradoxosomatidae, genus Desmoxytes Chamberlin, 1923). Arthropoda Selecta 14 (3): 251 - 257." type="journal article" year="2005">Nguyen et al. 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8BAF36D2FFEEAE5BFF1AFE78FD28AAD4" blockId="4.[174,774,253,501]" box="[234,674,477,501]" pageId="4" pageNumber="5">
<emphasis id="B964EAC0FFEEAE5BFF1AFE78FEE6AAD4" box="[234,364,477,501]" italics="true" pageId="4" pageNumber="5">Distribution</emphasis>
: Only known from
<collectingCountry id="F3077642FFEEAE5BFDB0FE78FD14AAD4" box="[576,670,477,501]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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