248 lines
20 KiB
XML
248 lines
20 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.6.1.7" ID-ISSN="2624-2834" ID-Zenodo-Dep="7754940" ID-ZooBank="039F76C9-00B9-47A2-8589-1FE9E4705899" approvalRequired="6" approvalRequired_for_taxonomicNames="1" approvalRequired_for_treatments="5" checkinTime="1679381240438" checkinUser="plazi" docAuthor="KIM, JUNGGON, ROCA-CUSACHS, MARCOS, PHAM, THAI HONG PHAM & JUNG, SUNGHOON" docDate="2023" docId="03B9878AFFA20C1CFCC2FEC6DDB4FB9B" docLanguage="en" docName="palaeoentomology.6.1.7.pdf" docOrigin="Palaeoentomology 6 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.1.7" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Cretanabis KIM & ROCA-CUSACHS & PHAM & JUNG 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="FF80FFF2FFA30C1FFFE8FF94DF10FFFC" masterDocTitle="Cretanabis kerzhneri gen. et sp. nov., the oldest nabine genus and species (Hemiptera: Heteroptera: Nabidae) from the mid-Cretaceous Burmese amber" masterLastPageNumber="48" masterPageNumber="41" pageNumber="42" updateTime="1679381663387" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cretanabis kerzhneri gen. et sp. nov., the oldest nabine genus and species (Hemiptera: Heteroptera: Nabidae) from the mid-Cretaceous Burmese amber</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>KIM, JUNGGON</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ROCA-CUSACHS, MARCOS</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>PHAM, THAI HONG PHAM</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>JUNG, SUNGHOON</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>48</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.6.1.7</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.6.1.7</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7754940</mods:identifier>
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<mods:identifier type="ZooBank">039F76C9-00B9-47A2-8589-1FE9E4705899</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7754934" ID-Zenodo-Dep="7754934" LSID="urn:lsid:plazi:treatment:03B9878AFFA20C1CFCC2FEC6DDB4FB9B" httpUri="http://treatment.plazi.org/id/03B9878AFFA20C1CFCC2FEC6DDB4FB9B" lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42">
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<subSubSection box="[810,1287,338,365]" pageId="1" pageNumber="42" type="nomenclature">
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<paragraph blockId="1.[810,1287,338,365]" box="[810,1287,338,365]" pageId="1" pageNumber="42">
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<heading box="[810,1287,338,365]" fontSize="10" level="2" pageId="1" pageNumber="42" reason="2">
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<emphasis bold="true" box="[810,1287,338,365]" pageId="1" pageNumber="42">
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Genus
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<taxonomicName authority="Kim & Jung" authorityName="KIM & ROCA-CUSACHS & PHAM & JUNG" authorityYear="2023" box="[893,1175,338,365]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[893,1019,338,365]" italics="true" pageId="1" pageNumber="42">Cretanabis</emphasis>
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Kim & Jung
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</taxonomicName>
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<taxonomicNameLabel box="[1182,1287,339,365]" pageId="1" pageNumber="42" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="type_taxon">
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<paragraph blockId="1.[810,1452,411,1382]" pageId="1" pageNumber="42">
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<emphasis bold="true" box="[810,967,411,437]" pageId="1" pageNumber="42">
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<typeStatus box="[810,869,411,437]" pageId="1" pageNumber="42">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName authority="KIM & ROCA-CUSACHS & PHAM & JUNG, 2023" authorityName="KIM & ROCA-CUSACHS & PHAM & JUNG" authorityYear="2023" box="[977,1215,411,437]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
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<emphasis box="[977,1215,411,437]" italics="true" pageId="1" pageNumber="42">Cretanabis kerzhneri</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1225,1317,411,437]" pageId="1" pageNumber="42">
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<taxonomicNameLabel box="[1225,1317,411,437]" pageId="1" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="etymology">
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<paragraph blockId="1.[810,1452,411,1382]" pageId="1" pageNumber="42">
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<emphasis bold="true" box="[863,998,484,510]" pageId="1" pageNumber="42">Etymology.</emphasis>
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Name derives from a combination of Cretaceous and ‘
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<taxonomicName authorityName="Latreille" authorityYear="1802" box="[1010,1075,520,546]" class="Insecta" family="Nabidae" genus="Nabis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[1010,1075,520,546]" italics="true" pageId="1" pageNumber="42">Nabis</emphasis>
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</taxonomicName>
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’, referring to the nabine group from the Cretaceous; gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="diagnosis">
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<paragraph blockId="1.[810,1452,411,1382]" pageId="1" pageNumber="42">
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<emphasis bold="true" box="[863,985,593,619]" pageId="1" pageNumber="42">Diagnosis.</emphasis>
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Differs from the nymphs (III instars) of other nabine genera(
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<emphasis box="[1034,1046,630,655]" italics="true" pageId="1" pageNumber="42">e</emphasis>
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.
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<emphasis box="[1054,1068,630,655]" italics="true" pageId="1" pageNumber="42">g</emphasis>
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.,
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<taxonomicName authority="Scott, 1881" authorityName="Scott" authorityYear="1881" box="[1084,1366,629,655]" class="Insecta" family="Nabidae" genus="Arachnocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[1084,1236,629,655]" italics="true" pageId="1" pageNumber="42">Arachnocoris</emphasis>
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Scott,1881
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</taxonomicName>
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;
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<taxonomicName authority="Stal, 1859" authorityName="Stal" authorityYear="1859" class="Insecta" family="Nabidae" genus="Gorpis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[1374,1452,630,656]" italics="true" pageId="1" pageNumber="42">Gorpis</emphasis>
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Stål, 1859
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</taxonomicName>
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;
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<taxonomicName authorityName="J.P.Wolff" authorityYear="1811" box="[941,1064,666,691]" class="Insecta" family="Nabidae" genus="Himacerus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis box="[941,1064,666,691]" italics="true" pageId="1" pageNumber="42">Himacerus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1072,1134,666,692]" pageId="1" pageNumber="42" sensu="stricto">
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<emphasis box="[1072,1083,666,691]" italics="true" pageId="1" pageNumber="42">s</emphasis>
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.
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<emphasis box="[1098,1128,666,691]" italics="true" pageId="1" pageNumber="42">str</emphasis>
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.
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</taxonomicNameLabel>
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Wolff, 1811;
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<taxonomicName authority="Reuter, 1890" authorityName="Reuter" authorityYear="1890" class="Insecta" family="Nabidae" genus="Hoplistoscelis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[1293,1452,665,691]" italics="true" pageId="1" pageNumber="42">Hoplistoscelis</emphasis>
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Reuter, 1890
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</taxonomicName>
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;
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<taxonomicName authority="Reuter, 1890" authorityName="Reuter" authorityYear="1890" box="[983,1283,702,728]" class="Insecta" family="Nabidae" genus="Lasiomerus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[983,1114,703,728]" italics="true" pageId="1" pageNumber="42">Lasiomerus</emphasis>
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Reuter, 1890
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</taxonomicName>
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;
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<taxonomicName authorityName="Latreille" authorityYear="1802" box="[1302,1367,702,728]" class="Insecta" family="Nabidae" genus="Nabis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus" sensu="lato">
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<emphasis box="[1302,1367,702,728]" italics="true" pageId="1" pageNumber="42">Nabis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1382,1451,702,728]" pageId="1" pageNumber="42" sensu="lato">
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<emphasis box="[1382,1393,703,728]" italics="true" pageId="1" pageNumber="42">s</emphasis>
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.
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<emphasis box="[1415,1445,702,728]" italics="true" pageId="1" pageNumber="42">lat</emphasis>
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.
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</taxonomicNameLabel>
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Latreille, 1802;
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<taxonomicName authority="Reuter, 1872" authorityName="Reuter" authorityYear="1872" box="[996,1222,738,765]" class="Insecta" family="Tettigoniidae" genus="Stalia" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[996,1061,738,764]" italics="true" pageId="1" pageNumber="42">Stalia</emphasis>
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Reuter, 1872
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</taxonomicName>
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;
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<taxonomicName authority="Reuter, 1890" authorityName="Reuter" authorityYear="1890" class="Insecta" family="Nabidae" genus="Stenonabis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis box="[1237,1360,738,764]" italics="true" pageId="1" pageNumber="42">Stenonabis</emphasis>
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Reuter, 1890
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</taxonomicName>
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) by the following combination of characters: body ovoid, armed with long sparse spines; head length less than width, width more than first antennomere length; antennae shorter than body length; first segment significantly thick; tapered to apex; third and fourth antennomeres covered with long setae; pronotum trapezoid, basal width less then basal width of mesothorax; femora and tibiae with long spines; tarsus less than 1/2 length of tibia in midleg; tarsus less than 1/3 length of tibia in hindleg; second tarsomere longest in mid and hind legs.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42" type="description">
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<paragraph blockId="1.[810,1452,411,1382]" lastBlockId="3.[136,778,849,1127]" lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42">
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<emphasis bold="true" box="[863,1009,1138,1164]" pageId="1" pageNumber="42">Description.</emphasis>
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Body ovoid, unicolorous.
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<emphasis italics="true" pageId="1" pageNumber="42">Surface and vestiture</emphasis>
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. Body glabrous, armed with long spines; head with spines (
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<figureCitation box="[948,1067,1210,1237]" captionStart="FIGURE 2" captionStartId="2.[136,235,1819,1843]" captionTargetBox="[159,1426,204,1789]" captionTargetId="figure-91@2.[151,1436,198,1795]" captionTargetPageId="2" captionText="FIGURE 2. Cretanabis kerzhneri sp. nov., holotype (CNUHHNF001). A, Dorsal habitus. B, Head. C, Antennae. D, Mid-leg. E, Hind-leg. F, Ventral habitus. Scale bars = 1 mm (A, F), 0.5 mm (B–E)." figureDoi="http://doi.org/10.5281/zenodo.7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="1" pageNumber="42">Fig. 2A, B</figureCitation>
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); antennae covered with long setae (
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<figureCitation box="[818,904,1247,1273]" captionStart="FIGURE 2" captionStartId="2.[136,235,1819,1843]" captionTargetBox="[159,1426,204,1789]" captionTargetId="figure-91@2.[151,1436,198,1795]" captionTargetPageId="2" captionText="FIGURE 2. Cretanabis kerzhneri sp. nov., holotype (CNUHHNF001). A, Dorsal habitus. B, Head. C, Antennae. D, Mid-leg. E, Hind-leg. F, Ventral habitus. Scale bars = 1 mm (A, F), 0.5 mm (B–E)." figureDoi="http://doi.org/10.5281/zenodo.7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="1" pageNumber="42">Fig. 2C</figureCitation>
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); pronotum with spines laterally; legs with spines, tibia with dense spines.
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<emphasis box="[1090,1193,1284,1310]" italics="true" pageId="1" pageNumber="42">Structure</emphasis>
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. Head: hypognathous, wider than long; vertex narrow, narrower than a single compound eye width; antennae cylindrical, linear, shorter than body length; first antennomere shortest, shorter than head width. Thorax: pronotum trapezoid, anterior width shorter than head width; pronotal midline length as long as 1/2 of basal width, longer than mesothorax midline length; mesothorax basal width more than pronotal basal width; legs generally slender; fore femur slightly incrassate; fore femur as long as fore tibia; femora and tibiae with spines (
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<figureCitation box="[818,941,2066,2092]" captionStart="FIGURE 2" captionStartId="2.[136,235,1819,1843]" captionTargetBox="[159,1426,204,1789]" captionTargetId="figure-91@2.[151,1436,198,1795]" captionTargetPageId="2" captionText="FIGURE 2. Cretanabis kerzhneri sp. nov., holotype (CNUHHNF001). A, Dorsal habitus. B, Head. C, Antennae. D, Mid-leg. E, Hind-leg. F, Ventral habitus. Scale bars = 1 mm (A, F), 0.5 mm (B–E)." figureDoi="http://doi.org/10.5281/zenodo.7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="2" pageNumber="43">Fig. 2D, E</figureCitation>
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); tarsus three segmented, tarsus less than 1/2 length of tibia in midleg, tarsus less than 1/3 length of tibia in hindleg; second tarsomere longest. Abdomen: ovoid, abdominal segment VII widest. Genitalia: not visible.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7754942" ID-Zenodo-Dep="7754942" httpUri="https://zenodo.org/record/7754942/files/figure.png" pageId="1" pageNumber="42" startId="1.[136,235,2014,2038]" targetBox="[215,1364,1412,1978]" targetPageId="1">
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<paragraph blockId="1.[136,1452,2014,2075]" pageId="1" pageNumber="42">
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<emphasis bold="true" box="[136,262,2014,2038]" pageId="1" pageNumber="42">FIGURE 1.</emphasis>
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Amber and fossil insect (curatorial museum code: CNUHHNF001).
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<emphasis bold="true" box="[977,995,2014,2038]" pageId="1" pageNumber="42">A</emphasis>
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, Amber with scale.
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<emphasis bold="true" box="[1205,1222,2015,2039]" pageId="1" pageNumber="42">B</emphasis>
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, New fossil specimen
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<taxonomicName authority="KIM & ROCA-CUSACHS & PHAM & JUNG, 2023" authorityName="KIM & ROCA-CUSACHS & PHAM & JUNG" authorityYear="2023" box="[136,350,2051,2075]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
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<emphasis box="[136,350,2051,2075]" italics="true" pageId="1" pageNumber="42">Cretanabis kerzhneri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[356,437,2051,2075]" pageId="1" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7754944" ID-Zenodo-Dep="7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="2" pageNumber="43" startId="2.[136,235,1819,1843]" targetBox="[159,1426,204,1789]" targetPageId="2">
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||
<paragraph blockId="2.[136,1451,1819,1879]" pageId="2" pageNumber="43">
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||
<emphasis bold="true" box="[136,261,1819,1843]" pageId="2" pageNumber="43">FIGURE 2.</emphasis>
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||
<taxonomicName authority="KIM & ROCA-CUSACHS & PHAM & JUNG, 2023" authorityName="KIM & ROCA-CUSACHS & PHAM & JUNG" authorityYear="2023" box="[268,482,1819,1843]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
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||
<emphasis box="[268,482,1819,1843]" italics="true" pageId="2" pageNumber="43">Cretanabis kerzhneri</emphasis>
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||
</taxonomicName>
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||
<emphasis bold="true" box="[489,570,1819,1843]" pageId="2" pageNumber="43">
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||
<taxonomicNameLabel box="[489,570,1819,1843]" pageId="2" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
, holotype (CNUHHNF001).
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<emphasis bold="true" box="[871,889,1819,1843]" pageId="2" pageNumber="43">A</emphasis>
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||
, Dorsal habitus.
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||
<emphasis bold="true" box="[1064,1081,1819,1843]" pageId="2" pageNumber="43">B</emphasis>
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||
, Head.
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||
<emphasis bold="true" box="[1161,1179,1819,1843]" pageId="2" pageNumber="43">C</emphasis>
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||
, Antennae.
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||
<emphasis bold="true" box="[1301,1319,1819,1843]" pageId="2" pageNumber="43">D</emphasis>
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||
, Mid-leg.
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||
<emphasis bold="true" box="[1428,1445,1819,1843]" pageId="2" pageNumber="43">E</emphasis>
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, Hind-leg.
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||
<emphasis bold="true" box="[239,254,1855,1879]" pageId="2" pageNumber="43">F</emphasis>
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, Ventral habitus. Scale bars = 1 mm (
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<emphasis bold="true" box="[635,653,1855,1879]" pageId="2" pageNumber="43">A</emphasis>
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,
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||
<emphasis bold="true" box="[666,681,1855,1879]" pageId="2" pageNumber="43">F</emphasis>
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), 0.5 mm (
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<emphasis bold="true" box="[795,842,1855,1879]" pageId="2" pageNumber="43">B–E</emphasis>
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).
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</paragraph>
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</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7754946" ID-Zenodo-Dep="7754946" box="[136,1357,769,794]" httpUri="https://zenodo.org/record/7754946/files/figure.png" pageId="3" pageNumber="44" startId="3.[136,235,769,793]" targetBox="[306,1277,210,728]" targetPageId="3">
|
||
<paragraph blockId="3.[136,1357,769,794]" box="[136,1357,769,794]" pageId="3" pageNumber="44">
|
||
<emphasis bold="true" box="[136,260,769,794]" pageId="3" pageNumber="44">FIGURE 3.</emphasis>
|
||
Habitus drawings of
|
||
<taxonomicName authority="KIM & ROCA-CUSACHS & PHAM & JUNG, 2023" authorityName="KIM & ROCA-CUSACHS & PHAM & JUNG" authorityYear="2023" box="[478,692,770,794]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
|
||
<emphasis box="[478,692,770,794]" italics="true" pageId="3" pageNumber="44">Cretanabis kerzhneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[698,803,769,794]" pageId="3" pageNumber="44">
|
||
<taxonomicNameLabel box="[698,779,770,794]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
, Dorsal habitus.
|
||
<emphasis bold="true" box="[976,993,770,794]" pageId="3" pageNumber="44">B</emphasis>
|
||
, Ventral habitus. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="3" pageNumber="44" type="discussion">
|
||
<paragraph blockId="3.[136,778,849,1127]" pageId="3" pageNumber="44">
|
||
<emphasis bold="true" box="[189,305,957,983]" pageId="3" pageNumber="44">Remarks.</emphasis>
|
||
A structure that morphologically resembles a fossula spongiosa is thinly developed on the apex of fore and middle tibiae. Further detailed examination of this structure is needed using suitable methodology (
|
||
<emphasis box="[730,742,1066,1091]" italics="true" pageId="3" pageNumber="44">e</emphasis>
|
||
.
|
||
<emphasis box="[750,764,1066,1091]" italics="true" pageId="3" pageNumber="44">g</emphasis>
|
||
., micro-CT, laser scanning confocal microscopy).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |