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<document id="ED08AA4D1D7EECCB59EB6D46B68DEF58" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFC3FFF2BF14FD440C1005A3" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus javanus Cameron 1936" docType="treatment" docVersion="2" lastPageNumber="29" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="26" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CA12019753743189E6414BF43A2E0737">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="F724717CB8B71651D851016B480030D0">Frisch, Johannes</mods:namePart>
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<mods:namePart id="6034574A12F752A90C8D67A3493072D7">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="904818BCFC7306218BF276E8B1C99AFA">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="75B6C2A32876E3118F6E3FDEDE11CE02">Soil Organisms</mods:title>
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<mods:date id="D9AE54B192BC98144B736702DBA3C7F4">2022</mods:date>
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<mods:number id="507A3D7D1033CA197B69F16FFD123957">2022-04-01</mods:number>
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<mods:number id="64926EED267F17CC0DA96942132267D5">95</mods:number>
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<mods:number id="92C8F23F9A3884E13D393AE7FE3A4CD0">1</mods:number>
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<mods:identifier id="5893994CE16BEB1D53B6A2BAD8A12A7D" type="DOI">10.25674/so95iss1id311</mods:identifier>
<mods:identifier id="0F31B9EB72ABE76F4E53ED72F21A193A" type="ISSN">2509-9523</mods:identifier>
<mods:identifier id="A84AAC8D1D8301A7AFBCC28AA221AF25" type="Zenodo-Dep">10724236</mods:identifier>
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<treatment id="03B7C77AFFC3FFF2BF14FD440C1005A3" LSID="urn:lsid:plazi:treatment:03B7C77AFFC3FFF2BF14FD440C1005A3" httpUri="http://treatment.plazi.org/id/03B7C77AFFC3FFF2BF14FD440C1005A3" lastPageId="6" lastPageNumber="29" pageId="3" pageNumber="26">
<subSubSection id="C30425E7FFC3FFF7BF14FD4409F106DB" box="[847,1260,700,726]" pageId="3" pageNumber="26" type="nomenclature">
<paragraph id="8BA1766CFFC3FFF7BF14FD4409F106DB" blockId="3.[847,1260,700,760]" box="[847,1260,700,726]" pageId="3" pageNumber="26">
<heading id="D0E9C100FFC3FFF7BF14FD4409F106DB" box="[847,1260,700,726]" fontSize="11" level="2" pageId="3" pageNumber="26" reason="5">
<taxonomicName id="4C1E0DEFFFC3FFF7BF14FD4409F106DB" ID-CoL="79XFL" authority="Cameron, 1936" authorityName="Cameron" authorityYear="1936" box="[847,1260,700,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC3FFF7BF14FD4409F106DB" bold="true" box="[847,1260,700,726]" pageId="3" pageNumber="26">
<emphasis id="B96AAA7EFFC3FFF7BF14FD44093A06DB" bold="true" box="[847,1063,700,726]" italics="true" pageId="3" pageNumber="26">Scopaeus javanus</emphasis>
<bibRefCitation id="EF8F0B9DFFC3FFF7B875FD4409F106DB" author="Cameron, M." box="[1070,1260,700,726]" pageId="3" pageNumber="26" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron, 1936</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC3FFF7BF14FD2509D506F5" box="[847,1224,733,760]" pageId="3" pageNumber="26" type="description">
<paragraph id="8BA1766CFFC3FFF7BF14FD2509D506F5" blockId="3.[847,1260,700,760]" box="[847,1224,733,760]" pageId="3" pageNumber="26">
(
<figureCitation id="13256AE9FFC3FFF7BF0CFD250EDC06FA" box="[855,961,733,759]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="3" pageNumber="26">Figs 1, 2</figureCitation>
,
<figureCitation id="13256AE9FFC3FFF7BF96FD250EF006FA" box="[973,1005,733,759]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="3" pageNumber="26">27</figureCitation>
,
<figureCitation id="13256AE9FFC3FFF7BFA2FD25095106F5" box="[1017,1100,733,760]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="3" pageNumber="26">50 –52</figureCitation>
,
<figureCitation id="13256AE9FFC3FFF7B803FD25099B06FA" box="[1112,1158,733,759]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="3" pageNumber="26">119</figureCitation>
,
<figureCitation id="13256AE9FFC3FFF7B8C9FD2509A206F5" box="[1170,1215,733,760]" captionStart="Figures 146–154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146–154. Sperm pump (146–154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150–152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="3" pageNumber="26">146</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC3FFF7BF14FCD8080D0737" box="[847,1296,799,826]" pageId="3" pageNumber="26" type="reference_group">
<paragraph id="8BA1766CFFC3FFF7BF14FCD8080D0737" blockId="3.[847,1296,799,826]" box="[847,1296,799,826]" pageId="3" pageNumber="26">
<treatmentCitationGroup id="AB0E5142FFC3FFF7BF14FCD8080D0737" box="[847,1296,799,826]" pageId="3" pageNumber="26">
<treatmentCitation id="0ABF507DFFC3FFF7BF14FCD808100737" author="Cameron, M." box="[847,1293,799,826]" page="52" pageId="3" pageNumber="26" year="1936">
<taxonomicName id="4C1E0DEFFFC3FFF7BF14FCD808100737" ID-CoL="79XFL" authority="Cameron, 1936: 52" authorityName="Cameron" authorityPageNumber="52" authorityYear="1936" box="[847,1293,799,826]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC3FFF7BF14FCD8093D0734" box="[847,1056,800,826]" italics="true" pageId="3" pageNumber="26">Scopaeus javanus</emphasis>
<bibRefCitation id="EF8F0B9DFFC3FFF7B87CFCE708100737" author="Cameron, M." box="[1063,1293,799,826]" pageId="3" pageNumber="26" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron, 1936: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC3FFF1BF14FC9A0E0B061C" lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26" type="materials_examined">
<paragraph id="8BA1766CFFC3FFF7BF14FC9A0EBD01C7" blockId="3.[824,1437,866,1976]" pageId="3" pageNumber="26">
<emphasis id="B96AAA7EFFC3FFF7BF14FC9A099A0777" bold="true" box="[847,1159,866,890]" pageId="3" pageNumber="26">Type specimens examined:</emphasis>
<typeStatus id="54A5C8CEFFC3FFF7B8C9FC9B09E00776" box="[1170,1277,867,891]" pageId="3" pageNumber="26" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D18BDE5FFC3FFF7B953FC9B08030777" box="[1288,1310,867,890]" count="1" pageId="3" pageNumber="26" type="male">♂</specimenCount>
,
<collectingCountry id="F30936FCFFC3FFF7B975FC9A08850777" box="[1326,1432,866,890]" name="Indonesia" pageId="3" pageNumber="26">Indonesia</collectingCountry>
,
<collectingRegion id="49DAB88EFFC3FFF7BF63FC7C0EDC0796" box="[824,961,900,923]" country="Indonesia" name="Jawa Barat" pageId="3" pageNumber="26">Jawa Barat</collectingRegion>
, N Bandung: Mt Tangkuban Perahu,
<date id="FFA050ACFFC3FFF7BF63FC5C0EAC07B1" box="[824,945,932,956]" pageId="3" pageNumber="26" value="1929-12-11">12.11.1929</date>
, leg. Drescher, by present designation; labeled ‘Type’ (red edged, round, printed label), ‘
<collectingRegion id="49DAB88EFFC3FFF7B952FC3D084D07D0" box="[1289,1360,965,989]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="26">JAVA</collectingRegion>
: / F.C. Drescher. / B.M. 1934.-264.’ (printed), ‘F.C. DRESCHER / G. Tangkoeban Prahoe / 4000-5000 Voet / Pranger,
<collectingRegion id="49DAB88EFFC3FFF7B936FBFF08810013" box="[1389,1436,1031,1054]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="26">Java</collectingRegion>
/
<date id="FFA050ACFFC3FFF7BF16FBD00ED4004D" box="[845,969,1064,1088]" pageId="3" pageNumber="26" value="1929-11-12">12.XI.1929</date>
’ (printed except date), ‘
<taxonomicName id="4C1E0DEFFFC3FFF7B884FBD00880004D" authority="Cam." authorityName="Cameron" authorityYear="1936" box="[1247,1437,1063,1088]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">S. javanus Cam.</taxonomicName>
TYPE’ (handwritten), ‘
<typeStatus id="54A5C8CEFFC3FFF7B86DFBB009BB006D" box="[1078,1190,1096,1120]" pageId="3" pageNumber="26" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFC3FFF7B89EFBB10894006D" authorityName="Cameron" authorityYear="1936" box="[1221,1417,1097,1120]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC3FFF7B89EFBB10894006D" box="[1221,1417,1097,1120]" italics="true" pageId="3" pageNumber="26">Scopaeus javanus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFC3FFF7BF63FB910EC4008C" author="Cameron, M." box="[824,985,1129,1153]" pageId="3" pageNumber="26" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron 1936</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFC3FFF7B829FB9109E3008C" author="Frisch, J. &amp; T. Mainda" box="[1138,1278,1129,1153]" pageId="3" pageNumber="26" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
’ (red, printed) (
<collectionCode id="ED0FEEA9FFC3FFF7BF64FB720E8900AF" box="[831,916,1162,1186]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="3" pageNumber="26">NHML</collectionCode>
).
<typeStatus id="54A5C8CEFFC3FFF7BFFCFB72092100AF" box="[935,1084,1162,1186]" pageId="3" pageNumber="26" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="9D18BDE5FFC3FFF7B813FB72096E00AF" box="[1096,1139,1162,1186]" count="1" pageId="3" pageNumber="26" type="male">1 ♂</specimenCount>
, labeled ‘Paratype’ (yellow edged, round, printed label), ‘cultus ex. / [illegible name] / Triatar-Kina’ (handwritten except for ‘cultus ex.’), ‘F.C. Drescher / G. Tangkoeban Prahoe / 4000 – 5000 Voet / Preanger,
<collectingRegion id="49DAB88EFFC3FFF7BFF3FAF60ECA0128" box="[936,983,1294,1317]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="26">Java</collectingRegion>
/
<date id="FFA050ACFFC3FFF7BFACFAF6096E012B" box="[1015,1139,1294,1318]" pageId="3" pageNumber="26" value="1929-11-12">12.XI.1929</date>
’ (printed except date), ‘
<taxonomicName id="4C1E0DEFFFC3FFF7B9DFFAF60ECF014A" authority="Cam." authorityName="Cameron" authorityYear="1936" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">S. javanus Cam.</taxonomicName>
COTYPE’ (handwritten), ‘M. Cameron / Bequest / B.M.1955-147.’ (printed), ‘
<typeStatus id="54A5C8CEFFC3FFF7B8AAFAB7089A016A" box="[1265,1415,1359,1383]" pageId="3" pageNumber="26" type="paralectotype">Paralectotype</typeStatus>
/
<taxonomicName id="4C1E0DEFFFC3FFF7BF63FA890EE70185" authorityName="Cameron" authorityYear="1936" box="[824,1018,1393,1416]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC3FFF7BF63FA890EE70185" box="[824,1018,1393,1416]" italics="true" pageId="3" pageNumber="26">Scopaeus javanus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF8F0B9DFFC3FFF7B84DFA8809AA0185" author="Cameron, M." box="[1046,1207,1392,1416]" pageId="3" pageNumber="26" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron 1936</bibRefCitation>
/ label by J.
<bibRefCitation id="EF8F0B9DFFC3FFF7B908FA880E6D01A4" author="Frisch, J. &amp; T. Mainda" pageId="3" pageNumber="26" pagination="139 - 147" refId="ref31869" refString="Frisch, J. &amp; T. Mainda (2022): The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species. - Soil Organisms 94 (3): 139 - 147." type="journal article" year="2022">Frisch, 2022</bibRefCitation>
’ (subsequent
<typeStatus id="54A5C8CEFFC3FFF7B840FA6909B601A4" box="[1051,1195,1425,1449]" pageId="3" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
label, red, printed) (
<collectionCode id="ED0FEEA9FFC3FFF7BF64FA4A0E8901C7" box="[831,916,1458,1482]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="3" pageNumber="26">NHML</collectionCode>
).
</paragraph>
<paragraph id="8BA1766CFFC3FFF1BF14FA2B0FFF0527" blockId="3.[824,1437,866,1976]" lastBlockId="5.[189,802,241,857]" lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26">
A
<typeStatus id="54A5C8CEFFC3FFF7BF34FA2B0ECD01E6" box="[879,976,1491,1515]" pageId="3" pageNumber="26" type="lectotype">lectotype</typeStatus>
is designated according to ICZN 1999, Article 74.1., because
<bibRefCitation id="EF8F0B9DFFC3FFF7B816FA0C08220201" author="Cameron, M." box="[1101,1343,1524,1548]" pageId="3" pageNumber="26" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron (1936: 52)</bibRefCitation>
neither distinguished a ‘type’ nor specified the total number of type specimens. The round type labels constitute subsequent labels attached by collection staff of NHML. The specimen selected as
<typeStatus id="54A5C8CEFFC3FFF7BFEAF98F090F0282" box="[945,1042,1655,1679]" pageId="3" pageNumber="26" type="lectotype">lectotype</typeStatus>
had already been labeled ‘
<taxonomicName id="4C1E0DEFFFC3FFF7B976F98F0E6C02BD" authority="Cam." authorityName="Cameron" authorityYear="1936" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">S. javanus Cam.</taxonomicName>
TYPE’, handwritten by Cameron, but this was not published. Another male of
<taxonomicName id="4C1E0DEFFFC3FFF7B8F7F941083D02DC" authorityName="Cameron" authorityYear="1936" box="[1196,1312,1721,1745]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC3FFF7B8F7F941083D02DC" box="[1196,1312,1721,1745]" italics="true" pageId="3" pageNumber="26">S. javanus</emphasis>
</taxonomicName>
at NHML doubtlessly belongs to the same sample as the
<typeStatus id="54A5C8CEFFC3FFF7B967F922088002FF" box="[1340,1437,1754,1778]" pageId="3" pageNumber="26" type="lectotype">lectotype</typeStatus>
and is therefore a
<typeStatus id="54A5C8CEFFC3FFF7B843F90309B7031E" box="[1048,1194,1787,1811]" pageId="3" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
. It was collected by Drescher at Mount Tangkuban Perahu on
<date id="FFA050ACFFC3FFF7B8A3F8E408720339" box="[1272,1391,1820,1844]" pageId="3" pageNumber="26" value="1929-12-11">12.11.1929</date>
and bears the same labels as the
<typeStatus id="54A5C8CEFFC3FFF7B835F8C509CF0358" box="[1134,1234,1853,1877]" pageId="3" pageNumber="26" type="lectotype">lectotype</typeStatus>
. It lacks, however, Cameron’s handwritten type and identification label. A female at MZB, which Drescher subsequently collected at the type locality on
<date id="FFA050ACFFC3FFF7B84CF867098D03BA" box="[1047,1168,1951,1975]" pageId="3" pageNumber="26" value="1929-12-23">23.12.1929</date>
, is not a type specimen, because
<bibRefCitation id="EF8F0B9DFFC5FFF1BD41FF0A0CEA0504" author="Cameron, M." box="[282,503,241,266]" pageId="5" pageNumber="28" pagination="25 - 54" refId="ref31076" refString="Cameron, M. (1936): Fauna Javanica. The Staphylinidae (Col.) Collected by Mr. F. C. Drescher. - Tijdschrift voor Entomologie 79: 25 - 54." type="journal article" year="1936">Cameron (1936: 52)</bibRefCitation>
based the description of
<taxonomicName id="4C1E0DEFFFC5FFF1BF56FF0A0C120527" authorityName="Cameron" authorityYear="1936" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC5FFF1BF56FF0A0C120527" italics="true" pageId="5" pageNumber="28">S. javanus</emphasis>
</taxonomicName>
on specimens collected on
<date id="FFA050ACFFC5FFF1BE6AFEEA0FB50527" box="[561,680,274,298]" pageId="5" pageNumber="28" value="1929-12-11">12.11.1929</date>
only.
</paragraph>
<caption id="DF6126E4FFC4FFF0BCCCF8A60C9A03B5" ID-DOI="http://doi.org/10.5281/zenodo.10724238" ID-Zenodo-Dep="10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="4" pageNumber="27" startId="4.[151,224,1886,1908]" targetBox="[224,1327,277,1777]" targetPageId="4" targetType="figure">
<paragraph id="8BA1766CFFC4FFF0BCCCF8A60C9A03B5" blockId="4.[151,1399,1886,1977]" pageId="4" pageNumber="27">
<emphasis id="B96AAA7EFFC4FFF0BCCCF8A60C10037E" bold="true" box="[151,269,1886,1908]" pageId="4" pageNumber="27">Figures 1–7.</emphasis>
Habitus of
<taxonomicName id="4C1E0DEFFFC4FFF0BD22F8A70F060379" authorityName="Cameron" authorityYear="1936" box="[377,539,1887,1908]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC4FFF0BD22F8A70F060379" box="[377,539,1887,1908]" italics="true" pageId="4" pageNumber="27">Scopaeus javanus</emphasis>
</taxonomicName>
, Jawa Barat, Sukabumi: Kiara Dua (
<emphasis id="B96AAA7EFFC4FFF0BF31F8A60E68037E" bold="true" box="[874,885,1886,1907]" pageId="4" pageNumber="27">1</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BFD2F8A70EFA0379" authorityName="Cameron" authorityYear="1936" box="[905,999,1887,1908]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC4FFF0BFD2F8A70EFA0379" box="[905,999,1887,1908]" italics="true" pageId="4" pageNumber="27">S. javanus</emphasis>
</taxonomicName>
, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (
<emphasis id="B96AAA7EFFC4FFF0BD15F88D0C440387" bold="true" box="[334,345,1909,1930]" pageId="4" pageNumber="27">2</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BD35F88E0F190387" authorityName="Frisch" authorityYear="2022" box="[366,516,1909,1931]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="batukaruensis">
<emphasis id="B96AAA7EFFC4FFF0BD35F88E0C640386" box="[366,377,1910,1931]" italics="true" pageId="4" pageNumber="27">S</emphasis>
.
<emphasis id="B96AAA7EFFC4FFF0BDDEF88D0F190387" box="[389,516,1909,1930]" italics="true" pageId="4" pageNumber="27">batukaruensis</emphasis>
</taxonomicName>
, paratype, Bali, Tabanan: Batukaru (
<emphasis id="B96AAA7EFFC4FFF0BF0DF88D0E7C0387" bold="true" box="[854,865,1909,1930]" pageId="4" pageNumber="27">3</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BF2EF88E0EDA0387" authorityName="Frisch" authorityYear="2022" box="[885,967,1909,1931]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="B96AAA7EFFC4FFF0BF2EF88E0EDA0387" box="[885,967,1909,1931]" italics="true" pageId="4" pageNumber="27">S. riedeli</emphasis>
</taxonomicName>
, holotype, Sumatra, South Lampung: Rajabasa Mts (
<emphasis id="B96AAA7EFFC4FFF0BC91F8740DC803AC" bold="true" box="[202,213,1932,1953]" pageId="4" pageNumber="27">4</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BCB7F8740C4303AC" authorityName="Cameron" authorityYear="1930" box="[236,350,1932,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="B96AAA7EFFC4FFF0BCB7F8740C4303AC" box="[236,350,1932,1953]" italics="true" pageId="4" pageNumber="27">S. jacobsoni</emphasis>
</taxonomicName>
, Jawa Barat: Mt Salak (
<emphasis id="B96AAA7EFFC4FFF0BE1CF8740F4F03AC" bold="true" box="[583,594,1932,1953]" pageId="4" pageNumber="27">5</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BE32F8740FA003AF" authorityName="Cameron" authorityYear="1941" box="[617,701,1932,1954]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="B96AAA7EFFC4FFF0BE32F8740FA003AF" box="[617,701,1932,1954]" italics="true" pageId="4" pageNumber="27">S. mixtus</emphasis>
</taxonomicName>
, Malaysia, Sabah: Lokan (
<emphasis id="B96AAA7EFFC4FFF0BFE6F8740ED503AC" bold="true" box="[957,968,1932,1953]" pageId="4" pageNumber="27">6</emphasis>
);
<taxonomicName id="4C1E0DEFFFC4FFF0BF84F874098203AC" authorityName="Frisch" authorityYear="2022" box="[991,1183,1932,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="halimunsalakensis">
<emphasis id="B96AAA7EFFC4FFF0BF84F8740EF703AC" box="[991,1002,1932,1953]" italics="true" pageId="4" pageNumber="27">S</emphasis>
.
<emphasis id="B96AAA7EFFC4FFF0BFA3F874098203AC" box="[1016,1183,1932,1953]" italics="true" pageId="4" pageNumber="27">halimunsalakensis</emphasis>
</taxonomicName>
, paratype, Jawa Barat, Sukabumi: Sirnaresmi (
<emphasis id="B96AAA7EFFC4FFF0BD35F85B0C6403B5" bold="true" box="[366,377,1955,1976]" pageId="4" pageNumber="27">7</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA1766CFFC5FFF1BC8FFECB0E0B061C" blockId="5.[189,802,241,857]" pageId="5" pageNumber="28">
<emphasis id="B96AAA7EFFC5FFF1BC8FFECB0F140546" bold="true" box="[212,521,307,331]" pageId="5" pageNumber="28">New Indonesian records:</emphasis>
<materialsCitation id="3B767C31FFC5FFF1BE46FECC0FB90561" collectingDate="1988-03-23" collectionCode="FMNH" collectorName="Wrensch &amp; Johnston" location="Bandung" pageId="5" pageNumber="28" specimenCount="1" stateProvince="Jawa Barat">
<collectingRegion id="49DAB88EFFC5FFF1BE46FECC0FB90546" box="[541,676,308,331]" country="Indonesia" name="Jawa Barat" pageId="5" pageNumber="28">Jawa Barat</collectingRegion>
:
<location id="8EC120B7FFC5FFF1BEE1FECB0E000546" LSID="urn:lsid:plazi:treatment:03B7C77AFFC3FFF2BF14FD440C1005A3:8EC120B7FFC5FFF1BEE1FECB0E000546" box="[698,797,307,331]" name="Bandung" pageId="5" pageNumber="28" stateProvince="Jawa Barat">Bandung</location>
,
<date id="FFA050ACFFC5FFF1BCE6FEAC0C370561" box="[189,298,340,364]" pageId="5" pageNumber="28" value="1988-03-23">
<collectingDate id="EFE4A944FFC5FFF1BCE6FEAC0C370561" box="[189,298,340,364]" pageId="5" pageNumber="28" value="1988-03-23">23.3.1988</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFC5FFF1BD3BFEAC0CA30561" box="[352,446,340,364]" pageId="5" pageNumber="28">Wrensch</collectorName>
&amp;
<collectorName id="26EB13BAFFC5FFF1BD85FEAC0F200561" box="[478,573,340,364]" pageId="5" pageNumber="28">Johnston</collectorName>
(
<collectionCode id="ED0FEEA9FFC5FFF1BE11FEAC0F800561" box="[586,669,340,364]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="28" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFC5FFF1BEF5FEAC0E0005A3" collectingDate="24.9. 2015, 27.9. 2015" collectionCode="MFNB, MZB" collectorName="Frisch" county="Sukabumi" elevation="710" latitude="-7.1408334" location="Ciletuh River" longLatPrecision="21" longitude="106.62945" municipality="Kiara Dua" pageId="5" pageNumber="28" specimenCount="1" stateProvince="Jawa Barat">
<collectingCounty id="62C00EE0FFC5FFF1BEF5FEAC0E000561" box="[686,797,340,364]" pageId="5" pageNumber="28">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFC5FFF1BCE6FE8D0C280580" box="[189,309,373,397]" pageId="5" pageNumber="28">Kiara Dua</collectingMunicipality>
:
<location id="8EC120B7FFC5FFF1BD18FE8D0CC50580" LSID="urn:lsid:plazi:treatment:03B7C77AFFC3FFF2BF14FD440C1005A3:8EC120B7FFC5FFF1BD18FE8D0CC50580" box="[323,472,373,397]" county="Sukabumi" latitude="-7.1408334" longLatPrecision="21" longitude="106.62945" municipality="Kiara Dua" name="Ciletuh River" pageId="5" pageNumber="28" stateProvince="Jawa Barat">Ciletuh River</location>
(
<geoCoordinate id="EE2A10ABFFC5FFF1BDB7FE8D0F6C0580" box="[492,625,373,397]" degrees="07" direction="south" minutes="08" orientation="latitude" pageId="5" pageNumber="28" precision="15" seconds="27" value="-7.1408334">07°08’27’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFC5FFF1BEDAFE8D0E080580" box="[641,789,373,397]" degrees="106" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="28" precision="15" seconds="46" value="106.62945">106°37’46’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFC5FFF1BCE6FE6E0C1E05A0" box="[189,259,406,430]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="28" unit="m" value="710.0">
<elevation id="0033915FFFC5FFF1BCE6FE6E0C1E05A0" box="[189,259,406,430]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="28" unit="m" value="710.0">710 m</elevation>
</quantity>
,
<collectingDate id="EFE4A944FFC5FFF1BD55FE6E0CEE05A3" box="[270,499,406,430]" isEnumeration="true" pageId="5" pageNumber="28">
<date id="FFA050ACFFC5FFF1BD55FE6E0C6605A3" box="[270,379,406,430]" pageId="5" pageNumber="28" value="2015-09-24">24.9.2015</date>
,
<date id="FFA050ACFFC5FFF1BDDDFE6E0CEE05A3" box="[390,499,406,430]" pageId="5" pageNumber="28" value="2015-09-27">27.9.2015</date>
</collectingDate>
, leg.
<collectorName id="26EB13BAFFC5FFF1BE77FE6E0F7205A3" box="[556,623,406,430]" pageId="5" pageNumber="28">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFC5FFF1BE26FE6E0FD005A3" box="[637,717,406,430]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="5" pageNumber="28">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFC5FFF1BE8CFE6E0E0805A3" box="[727,789,406,430]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="28" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFC5FFF1BCE6FE4F0FBC05FD" collectingDate="1989-06-11" collectionCode="MNHG" collectorName="Agosti &amp; Lobl &amp; Burckhardt" elevation="1400" location="Cibodas" pageId="5" pageNumber="28" specimenCount="1" stateProvince="Jawa Barat">
<locationDeviation id="9A136F4EFFC5FFF1BCE6FE4F0C8205C2" box="[189,415,439,463]" location="Bogor" pageId="5" pageNumber="28">
<quantity id="4CE6DB89FFC5FFF1BCE6FE4F0C1505C2" box="[189,264,439,463]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="28" unit="km" value="50.0">50 km</quantity>
NEE Bogor
</locationDeviation>
:
<location id="8EC120B7FFC5FFF1BDEBFE4F0F1705C2" LSID="urn:lsid:plazi:treatment:03B7C77AFFC3FFF2BF14FD440C1005A3:8EC120B7FFC5FFF1BDEBFE4F0F1705C2" box="[432,522,439,463]" name="Cibodas" pageId="5" pageNumber="28" stateProvince="Jawa Barat">Cibodas</location>
,
<quantity id="4CE6DB89FFC5FFF1BE47FE4F0F6B05C3" box="[540,630,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="28" unit="m" value="1400.0">
<elevation id="0033915FFFC5FFF1BE47FE4F0F6B05C3" box="[540,630,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="28" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 3. –
<date id="FFA050ACFFC5FFF1BEEAFE4F0E0005C2" box="[689,797,439,463]" pageId="5" pageNumber="28" value="1989-06-11">
<collectingDate id="EFE4A944FFC5FFF1BEEAFE4F0E0005C2" box="[689,797,439,463]" pageId="5" pageNumber="28" value="1989-06-11">6.11.1989</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFC5FFF1BCA9FE200C2105FD" box="[242,316,472,496]" pageId="5" pageNumber="28">Agosti</collectorName>
,
<collectorName id="26EB13BAFFC5FFF1BD14FE200C9C05FD" box="[335,385,472,496]" pageId="5" pageNumber="28">Löbl</collectorName>
&amp;
<collectorName id="26EB13BAFFC5FFF1BDEFFE200F3105FD" box="[436,556,472,496]" pageId="5" pageNumber="28">Burckhardt</collectorName>
(
<collectionCode id="ED0FEEA9FFC5FFF1BE19FE200F8705FD" box="[578,666,472,496]" pageId="5" pageNumber="28">MNHG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFC5FFF1BEEFFE200E0C061C" collectingDate="1997-05-28" collectionCode="MHNG" collectorName="Kurbatov" elevation="1450" location="Mt Gede" pageId="5" pageNumber="28" specimenCount="1" stateProvince="Jawa Barat">
<location id="8EC120B7FFC5FFF1BEEFFE200E0005FD" LSID="urn:lsid:plazi:treatment:03B7C77AFFC3FFF2BF14FD440C1005A3:8EC120B7FFC5FFF1BEEFFE200E0005FD" box="[692,797,472,496]" name="Mt Gede" pageId="5" pageNumber="28" stateProvince="Jawa Barat">Mt Gede</location>
,
<quantity id="4CE6DB89FFC5FFF1BCE6FE010C44061D" box="[189,345,504,529]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="5" pageNumber="28" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<elevation id="0033915FFFC5FFF1BCE6FE010C44061D" box="[189,345,504,529]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="5" pageNumber="28" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400 – 1500 m</elevation>
</quantity>
, 24. –
<date id="FFA050ACFFC5FFF1BDC1FE000F1A061D" box="[410,519,504,528]" pageId="5" pageNumber="28" value="1997-05-28">
<collectingDate id="EFE4A944FFC5FFF1BDC1FE000F1A061D" box="[410,519,504,528]" pageId="5" pageNumber="28" value="1997-05-28">28.5.1997</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFC5FFF1BE1BFE000FB8061D" box="[576,677,504,528]" pageId="5" pageNumber="28">Kurbatov</collectorName>
(
<collectionCode id="ED0FEEA9FFC5FFF1BEE9FE010E17061C" box="[690,778,505,529]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="28" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC5FFF2BC8FFDE10FE10561" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28" type="description">
<paragraph id="8BA1766CFFC5FFF1BC8FFDE1090D05A0" blockId="5.[189,802,241,857]" lastBlockId="5.[824,1437,241,857]" pageId="5" pageNumber="28">
<emphasis id="B96AAA7EFFC5FFF1BC8FFDE10C66063C" bold="true" box="[212,379,537,561]" pageId="5" pageNumber="28">Redescription:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFC5FFF1BEECFDE10E00063F" box="[695,797,537,562]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="5" pageNumber="28">Figs 1, 2</figureCitation>
. Pterodimorphous; brachypterous specimens (
<figureCitation id="13256AE9FFC5FFF1BE91FDC20E04065F" box="[714,793,570,594]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="5" pageNumber="28">Fig. 2</figureCitation>
) with reduced humeral angles and abdominal tergite VII without palisade fringe. Head subquadrate. Penultimate antennal segment quadrate. Mesotibia slender. Body surface with distinct, dense, setose punctation, without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body orange-brown with light, yellow-brown appendages (
<figureCitation id="13256AE9FFC5FFF1BEADFCD80DD60754" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="5" pageNumber="28">Fig. 2</figureCitation>
) to reddish medium brown with orange-brown posterior fifth of elytra, often somewhat lighter brown shoulders or basis of elytra, light brown tip of abdomen, light brown maxillary palpi and legs (
<figureCitation id="13256AE9FFC5FFF1B83DFECB09B60546" box="[1126,1195,307,331]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="5" pageNumber="28">Fig. 1</figureCitation>
); antennae gradually lighter from medium brown scapus to light brown segment 11. Total body length
<quantity id="4CE6DB89FFC5FFF1B8FAFE8D08360580" box="[1185,1323,373,397]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="5" pageNumber="28" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4 – 2.9 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFC5FFF1BFDAFE6E091105A0" box="[897,1036,406,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="5" pageNumber="28" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4 – 1.6 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFC5FFF2BF14FE4F0DE60546" blockId="5.[824,1437,241,857]" lastBlockId="6.[151,764,241,430]" lastPageId="6" lastPageNumber="29" pageId="5" pageNumber="28">
Male:Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with posterior margin evenly concave in posterior fifth (
<figureCitation id="13256AE9FFC5FFF1B8CAFE0009F6061C" box="[1169,1259,504,529]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="5" pageNumber="28">Fig. 119</figureCitation>
).Aedeagus (
<figureCitation id="13256AE9FFC5FFF1B936FE000E48063F" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="5" pageNumber="28">Figs 27</figureCitation>
,
<figureCitation id="13256AE9FFC5FFF1BF3BFDE10EB6063C" box="[864,939,537,561]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">50 – 52</figureCitation>
) about
<quantity id="4CE6DB89FFC5FFF1BFA7FDE20950063C" box="[1020,1101,538,562]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="28" unit="mm" value="0.7">0.7 mm</quantity>
long; apical lobes with convex ventral margins (
<figureCitation id="13256AE9FFC5FFF1BFB5FDC20921065F" box="[1006,1084,570,594]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">Fig. 50</figureCitation>
) and, in ventral and dorsal view, round apicolateral margins (
<figureCitation id="13256AE9FFC5FFF1B834FDA309DE067E" box="[1135,1219,603,627]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">Fig. 51</figureCitation>
: arrow
<collectionCode id="ED0FEEA9FFC5FFF1B941FDA40833067E" box="[1306,1326,604,627]" pageId="5" pageNumber="28">II</collectionCode>
,
<figureCitation id="13256AE9FFC5FFF1B966FDA30892067E" box="[1341,1423,603,627]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">Fig. 52</figureCitation>
); dorsal lobe strongly curved ventrad, strongly projecting from apical lobes ventrally, with round, lobiform end (
<figureCitation id="13256AE9FFC5FFF1B92AFD650E4B06DB" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">Fig. 50</figureCitation>
: arrow I); flagellum short, inconspicuous; lateral lobes evenly convex, not much projecting (
<figureCitation id="13256AE9FFC5FFF1B88EFD27084A06FA" box="[1237,1367,735,759]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">Figs 51, 52</figureCitation>
), with group of numerous, apicad and ventromediad pointing setae [setae broken off in
<figureCitation id="13256AE9FFC5FFF1B80CFCD809B10734" box="[1111,1196,800,825]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="5" pageNumber="28">Figs 27</figureCitation>
,
<figureCitation id="13256AE9FFC5FFF1B8E1FCD808180735" box="[1210,1285,800,824]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="5" pageNumber="28">50 – 52</figureCitation>
, but insertion points visible (for setation compare with aedeagus of
<taxonomicName id="4C1E0DEFFFC6FFF2BCCCFF0A0C5A0504" authorityName="Frisch" authorityYear="2022" box="[151,327,241,265]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="batukaruensis">
<emphasis id="B96AAA7EFFC6FFF2BCCCFF0A0C5A0504" box="[151,327,241,265]" italics="true" pageId="6" pageNumber="29">S. batukaruensis</emphasis>
</taxonomicName>
,
<figureCitation id="13256AE9FFC6FFF2BD08FF090CC80504" box="[339,469,241,265]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="6" pageNumber="29">Figs 53 – 55</figureCitation>
)]; transverse ridge of small, semicircular median foramen narrow, curved proximad (
<figureCitation id="13256AE9FFC6FFF2BCC4FECB0DF20546" box="[159,239,307,331]" captionStart="Figures 50–55" captionStartId="20.[150,223,1886,1908]" captionTargetBox="[222,1327,582,1831]" captionTargetId="figure-271@20.[151,1398,568,1866]" captionTargetPageId="20" captionText="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." figureDoi="http://doi.org/10.5281/zenodo.10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="6" pageNumber="29">Fig. 51</figureCitation>
).
</paragraph>
<caption id="DF6126E4FFC5FFF1BCE6F8720EFC03BB" ID-DOI="http://doi.org/10.5281/zenodo.10724240" ID-Zenodo-Dep="10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="5" pageNumber="28" startId="5.[189,262,1930,1952]" targetBox="[237,1406,902,1874]" targetPageId="5" targetType="figure">
<paragraph id="8BA1766CFFC5FFF1BCE6F8720EFC03BB" blockId="5.[189,1436,1930,1975]" pageId="5" pageNumber="28">
<emphasis id="B96AAA7EFFC5FFF1BCE6F8720C220392" bold="true" box="[189,319,1930,1952]" pageId="5" pageNumber="28">Figures 8–10.</emphasis>
Habitus of
<taxonomicName id="4C1E0DEFFFC5FFF1BDF6F8720FB30392" authorityName="Frisch" authorityYear="2022" box="[429,686,1930,1951]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="tortuosiflagellatus">
<emphasis id="B96AAA7EFFC5FFF1BDF6F8720FB30392" box="[429,686,1930,1951]" italics="true" pageId="5" pageNumber="28">Scopaeus tortuosiflagellatus</emphasis>
</taxonomicName>
, paratype, Jawa Barat, Sukabumi: Cisalimas (
<emphasis id="B96AAA7EFFC5FFF1B80CF872097F0392" bold="true" box="[1111,1122,1930,1951]" pageId="5" pageNumber="28">8</emphasis>
);
<taxonomicName id="4C1E0DEFFFC5FFF1B82DF87209F903AD" authorityName="Frisch" authorityYear="2022" box="[1142,1252,1930,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="posoanus">
<emphasis id="B96AAA7EFFC5FFF1B82DF872099C0392" box="[1142,1153,1930,1951]" italics="true" pageId="5" pageNumber="28">S</emphasis>
.
<emphasis id="B96AAA7EFFC5FFF1B8D5F87309F903AD" box="[1166,1252,1931,1952]" italics="true" pageId="5" pageNumber="28">posoanus</emphasis>
</taxonomicName>
, paratype, Sulawesi Tengah: Lake Poso (
<emphasis id="B96AAA7EFFC5FFF1BD23F8590C9E03BB" bold="true" box="[376,387,1953,1974]" pageId="5" pageNumber="28">9</emphasis>
);
<taxonomicName id="4C1E0DEFFFC5FFF1BDCDF8590F5A03BB" authorityName="Frisch" authorityYear="2022" box="[406,583,1953,1974]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="spiraliflagellatus">
<emphasis id="B96AAA7EFFC5FFF1BDCDF8590CBC03BB" box="[406,417,1953,1974]" italics="true" pageId="5" pageNumber="28">S</emphasis>
.
<emphasis id="B96AAA7EFFC5FFF1BDF6F8590F5A03BB" box="[429,583,1953,1974]" italics="true" pageId="5" pageNumber="28">spiraliflagellatus</emphasis>
</taxonomicName>
, paratype, Sulawesi Tengah: Lake Poso (
<emphasis id="B96AAA7EFFC5FFF1BFE6F8590EC903BB" bold="true" box="[957,980,1953,1974]" pageId="5" pageNumber="28">10</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BA1766CFFC6FFF2BCF5FEAC0FE10561" blockId="6.[151,764,241,430]" box="[174,764,340,364]" pageId="6" pageNumber="29">
Female: Sperm pump as in
<figureCitation id="13256AE9FFC6FFF2BD92FEAC0F3E0561" box="[457,547,340,364]" captionStart="Figures 146–154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146–154. Sperm pump (146–154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150–152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="6" pageNumber="29">Fig. 146</figureCitation>
; bursa membranous.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFC6FFF2BCF5FE8D0C1005A3" pageId="6" pageNumber="29" type="distribution">
<paragraph id="8BA1766CFFC6FFF2BCF5FE8D0C1005A3" blockId="6.[151,764,241,430]" pageId="6" pageNumber="29">
<emphasis id="B96AAA7EFFC6FFF2BCF5FE8D0C5F0580" bold="true" box="[174,322,373,397]" pageId="6" pageNumber="29">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFC6FFF2BD17FE8D0F130580" authorityName="Cameron" authorityYear="1936" box="[332,526,373,397]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96AAA7EFFC6FFF2BD17FE8D0F130580" box="[332,526,373,397]" italics="true" pageId="6" pageNumber="29">Scopaeus javanus</emphasis>
</taxonomicName>
is known from
<collectingRegion id="49DAB88EFFC6FFF2BE9DFE8E0DCC05A0" country="Indonesia" name="Jawa Barat" pageId="6" pageNumber="29">Jawa Barat</collectingRegion>
only.
</paragraph>
</subSubSection>
</treatment>
</document>
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