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<document id="EAA70F4CDE4E06F712ADCDA62E53B07D" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A136B5311FDA9F92FFBE5F935" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus placitus Moret 2020, sp. nov." docType="treatment" docUuid="02B83689-1D33-4C9B-85D2-8D6316CBD0FE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="37" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="36" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="542D5939FCF27D1B076D2E10DDA45E7A">
<mods:title id="EC6E7F3416CA4D2F7618F6694A5CE1E4">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="8E818C293C2F7BD9EF3B7E5D13ED0858" type="personal">
<mods:role id="DEDCAA9084F7A3E208150FF6A2F8A360">
<mods:roleTerm id="A98B2699CB971294428A8BB682576FEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="865AB5B23C66659B61E98EEF141337D9">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="4CD0FBA0F0206212BAE00EA16FBB72B7" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="056191B789263220766631963D11D677">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="A44C22E99C00CEA512E2268D03382478" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
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<mods:role id="73FE1B50EE5A5FA424540E469FDA2770">
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<mods:namePart id="6727112737BE2522C9100A42BE59AB0C">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="5D04222A00C13D541E163FFC9A2D5428" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="F041791B607DBCED23EED1DAF1ECBA15">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="B673A14E3D0AFFB67D4B4B418A60795F" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4EA570BCCDD24FCB36A2F469052EC173">text</mods:typeOfResource>
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<mods:titleInfo id="8EE895AC22E39FFFE98DFB455C187863">
<mods:title id="C6A8F16CD3F1F9BD925ECBCB9D77DE6F">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9AEB44757A8A0214EA8449A4080E933B">
<mods:date id="0C0F1B902BBD041686FFD12C903888E4">2020</mods:date>
<mods:detail id="AEA16FF9F70058E90787FB673B01E9B4" type="pubDate">
<mods:number id="3A88415332270D3DBA45DD5FAEE10DAB">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="C683828FC4556DFFEA6D43BF06A8F9AA" type="volume">
<mods:number id="F7C766D9C5A6E60F940285BD80135D20">646</mods:number>
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<mods:start id="415E54263C79C7B5EB9C386D699442A0">1</mods:start>
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<mods:classification id="8B5700B3F9153BFF1401132CEE9F2DFA">journal article</mods:classification>
<mods:identifier id="DF9DF50F74C19F842E96271360E80691" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="FA064C212EDE1183020735724FBB80E6" type="DOI">10.5852/ejt.2020.646</mods:identifier>
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<treatment id="03B7726A136B5311FDA9F92FFBE5F935" ID-DOI="http://doi.org/10.5281/zenodo.3848403" ID-GBIF-Taxon="164160394" ID-Zenodo-Dep="3848403" LSID="urn:lsid:zoobank.org:act:02B83689-1D33-4C9B-85D2-8D6316CBD0FE" httpUri="http://treatment.plazi.org/id/03B7726A136B5311FDA9F92FFBE5F935" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="C30490F7136B5316FDA9F92FFB40F897" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BA1C37C136B5316FDA9F92FFC5AF973" blockId="35.[365,1222,1749,1847]" box="[600,988,1749,1776]" pageId="35" pageNumber="36">
<heading id="D0E97410136B5316FDA9F92FFC5AF973" box="[600,988,1749,1776]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
<taxonomicName id="4C1EB8FF136B5316FDA9F92FFCFCF973" ID-CoL="8MDGD" authority="Moret" authorityName="Moret" authorityYear="2020" box="[600,890,1749,1776]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136B5316FDA9F92FFCADF973" bold="true" box="[600,811,1749,1776]" italics="true" pageId="35" pageNumber="36">Dyscolus placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136B5316FC70F92CFC5AF973" box="[897,988,1750,1776]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C136B5316FE9CF903FB40F897" blockId="35.[365,1222,1749,1847]" box="[365,1222,1785,1812]" pageId="35" pageNumber="36">
<uri id="FF8FCF7E136B5316FE9CF903FB40F897" box="[365,1222,1785,1812]" pageId="35" pageNumber="36">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A9136B5316FD61F903FB40F897" box="[656,1222,1785,1812]" pageId="35" pageNumber="36">02B83689-1D33-4C9B-85D2-8D6316CBD0FE</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136B5316FD59F8E6FC0AF8B4" box="[680,908,1820,1847]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BA1C37C136B5316FD59F8E6FC0AF8B4" blockId="35.[365,1222,1749,1847]" box="[680,908,1820,1847]" pageId="35" pageNumber="36">
<figureCitation id="1325DFF9136B5316FD59F8E6FCADF8B5" box="[680,811,1820,1846]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="35" pageNumber="36">Figs 38, 41</figureCitation>
(above)
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136B5316FF4CF8A4FD63F8FB" box="[189,741,1885,1912]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8BA1C37C136B5316FF4CF8A4FD63F8FB" blockId="35.[189,741,1885,1912]" box="[189,741,1885,1912]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB0EE452136B5316FF4CF8A4FD63F8FB" box="[189,741,1885,1912]" pageId="35" pageNumber="36">
<treatmentCitation id="0ABFE56D136B5316FF4CF8A4FDE2F8FB" author="Moret P." box="[189,612,1885,1912]" page="151" pageId="35" pageNumber="36" year="2005">
<taxonomicName id="4C1EB8FF136B5316FF4CF8A4FDE2F8FB" ID-CoL="388RK" authority="Moret, 2005: 151" authorityName="Moret" authorityPageNumber="151" authorityYear="2005" box="[189,612,1885,1912]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B96A1F6E136B5316FF4CF8A4FE09F8FB" box="[189,399,1886,1912]" italics="true" pageId="35" pageNumber="36">Dyscolus hirsutus</emphasis>
<bibRefCitation id="EF8FBE8D136B5316FE67F8A4FDE2F8FB" author="Moret P." box="[406,612,1885,1912]" pageId="35" pageNumber="36" refId="ref27190" refString="Moret P. 2005. Los Coleopteros Carabidae del Paramo en los Andes del Ecuador: Sistematica, Ecologia y Biogeografia. Pontificia Universidad Catolica del Ecuador, Centro de Biodiversidad y Ambiente, Monografia 2, Quito." type="book" year="2005">Moret, 2005: 151</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B96A1F6E136B5316FD80F8A5FD58F8FB" box="[625,734,1887,1912]" italics="true" pageId="35" pageNumber="36">pro parte</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136B5316FF4CF865FD1DF86B" box="[189,667,1951,2024]" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8BA1C37C136B5316FF4CF865FEC2F83A" blockId="35.[189,324,1951,1977]" box="[189,324,1951,1977]" pageId="35" pageNumber="36">
<heading id="D0E97410136B5316FF4CF865FEC2F83A" bold="true" box="[189,324,1951,1977]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B96A1F6E136B5316FF4CF865FEC2F83A" bold="true" box="[189,324,1951,1977]" pageId="35" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136B5316FF4CF834FD1DF86B" blockId="35.[189,667,1998,2024]" box="[189,667,1998,2024]" pageId="35" pageNumber="36">Latin adjective meaning ‘nice, pleasant’.</paragraph>
</subSubSection>
<subSubSection id="C30490F7136C5311FF4CFEF1FBE5F935" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BA1C37C136C5311FF4CFEF1FEEAFEA6" blockId="36.[189,364,267,293]" box="[189,364,267,293]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFEF1FEEAFEA6" bold="true" box="[189,364,267,293]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFEF1FEEAFEA6" bold="true" box="[189,364,267,293]" pageId="36" pageNumber="37">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136C5311FF1BFEC0FEDAFED7" blockId="36.[189,1399,314,410]" box="[234,348,314,340]" pageId="36" pageNumber="37">
<emphasis id="B96A1F6E136C5311FF1BFEC0FEDAFED7" bold="true" box="[234,348,314,340]" pageId="36" pageNumber="37">
<typeStatus id="54A57DDE136C5311FF1BFEC0FEDAFED7" box="[234,348,314,340]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFEA7FC47FE19" blockId="36.[189,1399,314,410]" pageId="36" pageNumber="37">
<materialsCitation id="3B76C921136C5311FF4CFEA7FC47FE19" ID-GBIF-Occurrence="2620303382" collectingDate="2017-03-11" collectionCode="MNHN" collectorName="P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="Waypoint 283" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo Province" typeStatus="holotype">
<collectingCountry id="F30983EC136C5311FF4CFEA7FECDFEF4" box="[189,331,349,375]" name="Ecuador" pageId="36" pageNumber="37">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F5136C5311FEAEFEA1FEF0FEF4" box="[351,374,347,375]" pageId="36" pageNumber="37" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E136C5311FE73FEA7FDB7FEF4" box="[386,561,349,375]" country="Ecuador" name="Napo" pageId="36" pageNumber="37">Napo Province</collectingRegion>
,
<collectingCounty id="62C0BBF0136C5311FDCAFEA7FD2CFEF4" box="[571,682,349,375]" pageId="36" pageNumber="37">Guamaní</collectingCounty>
,
<collectingMunicipality id="6BC55906136C5311FD42FEA7FCFDFEF4" box="[691,891,349,375]" pageId="36" pageNumber="37">Paso de la Virgen</collectingMunicipality>
,
<location id="8EC195A7136C5311FC75FEA7FBAFFEF4" LSID="urn:lsid:plazi:treatment:03B7726A136B5311FDA9F92FFBE5F935:8EC195A7136C5311FC75FEA7FBAFFEF4" box="[900,1065,349,375]" country="Ecuador" county="Guamani" latitude="-0.35016665" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" name="Waypoint 283" pageId="36" pageNumber="37" stateProvince="Napo Province">Waypoint 283</location>
;
<geoCoordinate id="EE2AA5BB136C5311FBC2FEA1FB3FFEF4" box="[1075,1209,347,375]" degrees="0" direction="south" minutes="21" orientation="latitude" pageId="36" pageNumber="37" precision="1" seconds="0.6" value="-0.35016665">0°21′0.6″ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB136C5311FB35FEA1FAF4FEF4" box="[1220,1394,347,375]" degrees="78" direction="west" minutes="11" orientation="longitude" pageId="36" pageNumber="37" precision="1" seconds="52.4" value="-78.19789">78°11′52.4″ W</geoCoordinate>
;
<elevation id="0033244F136C5311FF4CFE7AFED0FE19" box="[189,342,384,410]" metricMagnitude="3" metricUnit="m" metricValue="3.8899999999999997" pageId="36" pageNumber="37" unit="m" value="3890.0">
<quantity id="4CE66E99136C5311FF4CFE7AFE90FE19" box="[189,278,384,410]" metricMagnitude="3" metricUnit="m" metricValue="3.8899999999999997" pageId="36" pageNumber="37" unit="m" value="3890.0">3890 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136C5311FE92FE7AFD85FE19" box="[355,515,384,410]" pageId="36" pageNumber="37" value="2017-03-11">
<collectingDate id="EFE41C54136C5311FE92FE7AFD85FE19" box="[355,515,384,410]" pageId="36" pageNumber="37" value="2017-03-11">11 Mar. 2017</collectingDate>
</date>
;
<collectorName id="26EBA6AA136C5311FDFEFE7AFDF5FE19" box="[527,627,384,410]" pageId="36" pageNumber="37">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136C5311FD5AFE7AFC9BFE19" box="[683,797,384,410]" pageId="36" pageNumber="37">M. Gobbi</collectorName>
leg.;
<collectionCode id="ED0F5BB9136C5311FCAEFE7AFC3BFE19" box="[863,957,384,410]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="36" pageNumber="37">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C136C5311FF1BFE40FE79FE57" blockId="36.[189,1398,440,609]" box="[234,511,440,468]" pageId="36" pageNumber="37">
<emphasis id="B96A1F6E136C5311FF1BFE40FEE1FE57" bold="true" box="[234,359,442,468]" pageId="36" pageNumber="37">
<typeStatus id="54A57DDE136C5311FF1BFE40FEE1FE57" box="[234,359,442,468]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 ♂♂,
<specimenCount id="9D1808F5136C5311FE3BFE40FE72FE57" box="[458,500,442,468]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFE27FD9DFDE2" blockId="36.[189,1398,440,609]" pageId="36" pageNumber="37">
<materialsCitation id="3B76C921136C5311FF4CFE27FC04FE74" ID-GBIF-Occurrence="2620303417" box="[189,898,475,503]" collectingDate="2017-03-11" collectionCode="CPM" collectorName="Paso de la Virgen &amp; P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="ECUADOR" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<collectingCountry id="F30983EC136C5311FF4CFE27FECDFE74" box="[189,331,477,503]" name="Ecuador" pageId="36" pageNumber="37">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F5136C5311FE91FE27FE0AFE74" box="[352,396,475,503]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB9136C5311FCB5FE27FC04FE74" box="[836,898,477,503]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="36" pageNumber="37">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136C5311FC69FE27FEEDFD98" ID-GBIF-Occurrence="2620303353" collectingDate="2017-03-11" collectionCode="QCAZ" collectorName="Paso de la Virgen &amp; P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="Waypoint" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCode="QCAZ 256725" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="9D1808F5136C5311FC69FE27FC45FE74" box="[920,963,477,503]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBB86B07136C5311FF4CFDFAFEEDFD98" box="[189,363,512,539]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="36" pageNumber="37" type="Museum">QCAZ 256725</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B76C921136C5311FE70FDFAFD91FDE2" ID-GBIF-Occurrence="2620303334" collectingDate="1982-05-13" collectionCode="UASM" collectorName="H. Frania &amp; F. Sperling" country="Ecuador" elevation="3810" location="Paso de Papallacta" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo Province" typeStatus="paratype">
<specimenCount id="9D1808F5136C5311FE70FDFAFE2AFD99" box="[385,428,510,538]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
(red label: “
<taxonomicName id="4C1EB8FF136C5311FDB3FDFAFBADFD98" authority="Moret" authorityName="Moret" authorityYear="2005" box="[578,1067,512,539]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hirsutus" subGenus="Hydrodyscolus">
<emphasis id="B96A1F6E136C5311FDB3FDFAFD28FD99" box="[578,686,512,538]" italics="true" pageId="36" pageNumber="37">Dyscolus</emphasis>
(
<emphasis id="B96A1F6E136C5311FD4CFDFAFCEAFD99" box="[701,876,512,538]" italics="true" pageId="36" pageNumber="37">Hydrodyscolus</emphasis>
)
<emphasis id="B96A1F6E136C5311FC8DFDFAFC5DFD99" box="[892,987,512,538]" italics="true" pageId="36" pageNumber="37">hirsutus</emphasis>
Moret
</taxonomicName>
, Paratype”);
<collectingRegion id="49DA0D9E136C5311FB30FDFBFAF4FD99" box="[1217,1394,512,539]" country="Ecuador" name="Napo" pageId="36" pageNumber="37">Napo Province</collectingRegion>
,
<location id="8EC195A7136C5311FF4CFDDEFE11FDBE" LSID="urn:lsid:plazi:treatment:03B7726A136B5311FDA9F92FFBE5F935:8EC195A7136C5311FF4CFDDEFE11FDBE" box="[189,407,547,574]" country="Ecuador" name="Paso de Papallacta" pageId="36" pageNumber="37" stateProvince="Napo Province">Paso de Papallacta</location>
(Guamaní),
<elevation id="0033244F136C5311FDDAFDD9FD45FDBE" box="[555,707,547,574]" metricMagnitude="3" metricUnit="m" metricValue="3.81" pageId="36" pageNumber="37" unit="m" value="3810.0">
<quantity id="4CE66E99136C5311FDDAFDD9FD02FDBD" box="[555,644,547,574]" metricMagnitude="3" metricUnit="m" metricValue="3.81" pageId="36" pageNumber="37" unit="m" value="3810.0">3810 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC136C5311FD21FDD9FCE9FDBE" box="[720,879,547,574]" pageId="36" pageNumber="37" value="1982-05-13">
<collectingDate id="EFE41C54136C5311FD21FDD9FCE9FDBE" box="[720,879,547,574]" pageId="36" pageNumber="37" value="1982-05-13">13 May 1982</collectingDate>
</date>
;
<collectorName id="26EBA6AA136C5311FC8AFDDEFC6CFDBE" box="[891,1002,547,574]" pageId="36" pageNumber="37">H. Frania</collectorName>
and
<collectorName id="26EBA6AA136C5311FBD3FDDEFB27FDBE" box="[1058,1185,547,574]" pageId="36" pageNumber="37">F. Sperling</collectorName>
leg.; under stones and logs near stream;
<collectionCode id="ED0F5BB9136C5311FE4FFDBDFD91FDE2" box="[446,535,583,609]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="36" pageNumber="37">UASM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFD7AFE52FD19" blockId="36.[189,468,640,666]" box="[189,468,640,666]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFD7AFE52FD19" bold="true" box="[189,468,640,666]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFD7AFE52FD19" bold="true" box="[189,468,640,666]" pageId="36" pageNumber="37">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFD55FB10FBEC" blockId="36.[189,1399,687,1136]" pageId="36" pageNumber="37">
Habitus:
<figureCitation id="1325DFF9136C5311FED9FD55FE06FD4A" box="[296,384,687,713]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="36" pageNumber="37">Fig. 38</figureCitation>
. Wingless. Body length:
<quantity id="4CE66E99136C5311FD56FD55FCBEFD4A" box="[679,824,687,713]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" metricValueMax="9.6" metricValueMin="8.8" pageId="36" pageNumber="37" unit="mm" value="9.2" valueMax="9.6" valueMin="8.8">8.8–9.6 mm</quantity>
. Dorsal surface of the body entirely dark brown; ventral surface mostly dark brown, with reddish areas on the sides of the abdominal sclerites and on most parts of the coxae and trochanters; legs, antennae and mouthparts reddish brown. Elytral microsculpture made of slightly oblong sculpticells. Head convex, collar constriction strong, eyes moderately bulging, genae flat in dorsal view, mandibles moderately long and acute; submentum with two or three pairs of setae, mentum with one pair of setae. Pronotum cordiform, hind angles right angled, blunt; two pairs of lateral setae. Elytra elongate-oval, narrow, without subapical sinuation, rounded at apex; humeri almost entirely effaced. Striae well impressed, intervals feebly convex. Third interval with 3–6 setae, fifth interval with 0–3 setae, seventh interval with 0 or 1 seta; 17 to 20 umbilicate setae along the lateral margin. Last visible abdominal ventrite with one pair (♂) or two pairs (
<specimenCount id="9D1808F5136C5311FBDDFC16FBB9FB85" box="[1068,1087,1004,1030]" pageId="36" pageNumber="37" type="female">♀</specimenCount>
) of setae along its apical margin. Legs slender; fourth metatarsomere without subapical dorsolateral setae, apical lobes parallel, the outer lobe 1.5 times longer than the inner lobe.
<emphasis id="B96A1F6E136C5311FCF8FBC8FC32FBCF" box="[777,948,1074,1100]" italics="true" pageId="36" pageNumber="37">Male genitalia</emphasis>
:
<figureCitation id="1325DFF9136C5311FC33FBC8FB9EFBCF" box="[962,1048,1074,1100]" captionStart="Figs 36–41" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 36–41. Dyscolus spp., habitus (36–38) and aedeagus, median lobe in lateral view (39–40) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="36" pageNumber="37">Fig. 41</figureCitation>
. Apex of the median lobe bent downwards; endophallus without sclerotized structure.
<emphasis id="B96A1F6E136C5311FCB4FBAFFB8BFBEC" box="[837,1037,1109,1135]" italics="true" pageId="36" pageNumber="37">Female genitalia</emphasis>
: unstudied.
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFB6DFEE5FB32" blockId="36.[189,355,1175,1201]" box="[189,355,1175,1201]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFB6DFEE5FB32" bold="true" box="[189,355,1175,1201]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFB6DFEE5FB32" bold="true" box="[189,355,1175,1201]" pageId="36" pageNumber="37">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFB3FFAF1FA31" blockId="36.[189,1399,1221,1459]" pageId="36" pageNumber="37">
<taxonomicName id="4C1EB8FF136C5311FF4CFB3FFE08FB5C" authorityName="Moret" authorityYear="2005" box="[189,398,1221,1247]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B96A1F6E136C5311FF4CFB3FFE08FB5C" box="[189,398,1221,1247]" italics="true" pageId="36" pageNumber="37">Dyscolus hirsutus</emphasis>
</taxonomicName>
, now geographically restricted to the El Altar volcano, differs from
<taxonomicName id="4C1EB8FF136C5311FB59FB3CFAF1FB63" authority="Moret" authorityName="Moret" authorityYear="2020" box="[1192,1399,1221,1248]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136C5311FB59FB3CFAAEFB5C" box="[1192,1320,1221,1247]" italics="true" pageId="36" pageNumber="37">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136C5311FF4CFB13FE9EFA80" box="[189,280,1257,1283]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
by the following characters: upper surface bicolorous with the head and the elytra dark brown, the pronotum reddish; ventral surface (including the epipleura) entirely reddish, except for the head which is dark brown; mandibles with a long and very acute tip; eyes more bulging; hind angles of the pronotum broadly rounded; humeri more effaced; seventh interval with 1 or 2 setae; apex of the median lobe broader, apically rounded (slightly hooked in
<taxonomicName id="4C1EB8FF136C5311FCD4FA8CFC7DFA13" authority="Moret" authorityName="Moret" authorityYear="2020" box="[805,1019,1397,1424]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E136C5311FCD4FA8CFC2FFA0C" box="[805,937,1397,1423]" italics="true" pageId="36" pageNumber="37">D. placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136C5311FBF4FA8CFBE1FA13" box="[1029,1127,1398,1424]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
). The differences with
<taxonomicName id="4C1EB8FF136C5311FF4CFA63FE08FA30" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,398,1433,1459]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="arauzae" status="sp. nov.">
<emphasis id="B96A1F6E136C5311FF4CFA63FEC6FA31" box="[189,320,1433,1458]" italics="true" pageId="36" pageNumber="37">D. arauzae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136C5311FE64FA63FE69FA30" box="[405,495,1433,1459]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF136C5311FDD6FA63FD7CFA30" authority="Moret" authorityName="Moret" authorityYear="2020" box="[551,762,1433,1459]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
<emphasis id="B96A1F6E136C5311FDD6FA63FD2DFA31" box="[551,683,1433,1458]" italics="true" pageId="36" pageNumber="37">D. piscator</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A215136C5311FCF0FA63FCDDFA30" box="[769,859,1433,1459]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
are set forth in the diagnoses of these species.
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CFA20FE9BFA77" blockId="36.[189,285,1498,1524]" box="[189,285,1498,1524]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFA20FE9BFA77" bold="true" box="[189,285,1498,1524]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFA20FE9BFA77" bold="true" box="[189,285,1498,1524]" pageId="36" pageNumber="37">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CF9F2FA93F9C5" blockId="36.[189,1399,1544,1606]" pageId="36" pageNumber="37">
Hygrophilous in the upper montane forest, at around
<quantity id="4CE66E99136C5311FCB0F9F3FC60F9A0" box="[833,998,1545,1571]" metricMagnitude="3" metricUnit="m" metricValue="3.85" metricValueMax="3.9" metricValueMin="3.8" pageId="36" pageNumber="37" unit="m" value="3850.0" valueMax="3900.0" valueMin="3800.0">3800–3900 m</quantity>
a.s.l. The specimens collected in 2017 were found in the understory of a
<taxonomicName id="4C1EB8FF136C5311FD7CF9D6FD7CF9C5" box="[653,762,1580,1606]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E136C5311FD7CF9D6FD7CF9C5" box="[653,762,1580,1606]" italics="true" pageId="36" pageNumber="37">Polylepis</emphasis>
</taxonomicName>
forest, beneath soaked mosses on the ground.
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CF997FE6DF904" blockId="36.[189,491,1645,1671]" box="[189,491,1645,1671]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CF997FE6DF904" bold="true" box="[189,491,1645,1671]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CF997FE6DF904" bold="true" box="[189,491,1645,1671]" pageId="36" pageNumber="37">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C136C5311FF4CF966FBE5F935" blockId="36.[189,1123,1692,1718]" box="[189,1123,1692,1718]" pageId="36" pageNumber="37">Microendemic species, restricted to the Guamaní area in the Eastern Cordillera.</paragraph>
</subSubSection>
</treatment>
</document>
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