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<document id="72255D8E999D5CC4CF1ED21091227B70" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A1366531AFDCEF901FAB4F806" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus verecundior Moret 2020, sp. nov." docType="treatment" docUuid="DAB1A22E-25D1-438C-AE2E-25542AF7E50F" docUuidSource="ZooBank" docVersion="8" lastPageNumber="48" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="47" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F32F5C5DF965CBA13A3DB3DEBB8B06F5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C1453AA8E4DAAF27FB3CBDE5EF0BEFB">
<mods:title id="9B161C3C29C69CC7A38124D71A0A2131">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="C87FFF301AD3E7A555CFEB63099FE515" type="personal">
<mods:role id="2D1C803D6F9880A1CB96343B56FF8BF9">
<mods:roleTerm id="0FE5BF455EC4A4909632833B4CD03CD8">Author</mods:roleTerm>
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<mods:namePart id="39174298D93402AA22C53ACF5E4B8147">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="5849E9D23568B327FD97AD8651FFF556" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="E2493F27BF9B34B290CE261EEEB0C036">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="EAC95B8CBAB073BCD945DC3F1DF6B7ED" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="3529181F072E6F01774976A175AC667A" type="personal">
<mods:role id="16E4559E370064DE94886D7A7E50C323">
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</mods:role>
<mods:namePart id="987859F3CEC88BFFE025047FA416AF02">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="412E8839030F98E7C4B5C476014875BF" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="B573F72F92F6905ED8BDA8D3D86A43FA">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="06725A00A8573320FBEAA4441F684495" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="6835B53ECF3EA12ED128EC18F84A0D46">text</mods:typeOfResource>
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<mods:titleInfo id="2BAA9392B32E800EA5C097DCA4658FD6">
<mods:title id="5C9C2DA5D9A5D42341A5CEB58F0DCFA3">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="3E14CA03B31F48B7BDA7A44952084EBE">
<mods:date id="DC56138BA858E97C60E4C8CB4C33BE19">2020</mods:date>
<mods:detail id="6F90BD4A3DF4CEFB1EF778B4AA170624" type="pubDate">
<mods:number id="7D757943CF24B4A9F94268CE409DA125">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="D146582FA4AEABF93AEEB71AE322C5BB" type="volume">
<mods:number id="3994F09F71B3C3C7E978654539FB1BF7">646</mods:number>
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<mods:start id="68E6209C55893AF07EFBECAAF7795741">1</mods:start>
<mods:end id="D7616D86A6B8CB91B820F7FF38BA814B">55</mods:end>
</mods:extent>
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<mods:classification id="CB12D47FE10C490053E64C7B606231D9">journal article</mods:classification>
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<mods:identifier id="B23C531729E09B20BE5BBEAEEB581856" type="DOI">10.5852/ejt.2020.646</mods:identifier>
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<treatment id="03B7726A1366531AFDCEF901FAB4F806" ID-DOI="http://doi.org/10.5281/zenodo.3848415" ID-GBIF-Taxon="164160399" ID-Zenodo-Dep="3848415" LSID="urn:lsid:zoobank.org:act:DAB1A22E-25D1-438C-AE2E-25542AF7E50F" httpUri="http://treatment.plazi.org/id/03B7726A1366531AFDCEF901FAB4F806" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C30490F71366531BFDCEF901FB4DF8B9" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA1C37C1366531BFDCEF901FC73F895" blockId="46.[360,1227,1787,1885]" box="[575,1013,1787,1814]" pageId="46" pageNumber="47">
<heading id="D0E974101366531BFDCEF901FC73F895" box="[575,1013,1787,1814]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="4C1EB8FF1366531BFDCEF901FC12F895" ID-CoL="8M6SR" authority="Moret" authorityName="Moret" authorityYear="2020" box="[575,916,1787,1814]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="verecundior" status="sp. nov.">
<emphasis id="B96A1F6E1366531BFDCEF901FCC2F895" bold="true" box="[575,836,1787,1814]" italics="true" pageId="46" pageNumber="47">Dyscolus verecundior</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151366531BFC6AF906FC73F895" box="[923,1013,1788,1814]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C1366531BFE99F8E5FB4DF8B9" blockId="46.[360,1227,1787,1885]" box="[360,1227,1823,1850]" pageId="46" pageNumber="47">
<uri id="FF8FCF7E1366531BFE99F8E5FB4DF8B9" box="[360,1227,1823,1850]" pageId="46" pageNumber="47">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A91366531BFD76F8E5FB4DF8B9" box="[647,1227,1823,1850]" pageId="46" pageNumber="47">DAB1A22E-25D1-438C-AE2E-25542AF7E50F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71366531BFD29F8B8FCDDF8DE" box="[728,859,1858,1885]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA1C37C1366531BFD29F8B8FCDDF8DE" blockId="46.[360,1227,1787,1885]" box="[728,859,1858,1885]" pageId="46" pageNumber="47">
<figureCitation id="1325DFF91366531BFD29F8B8FCB4F8DE" box="[728,818,1858,1885]" captionStart="Figs 54–55" captionStartId="48.[189,241,1964,1990]" captionTargetBox="[254,1364,851,1900]" captionTargetId="figure@48.[203,1385,843,1920]" captionTargetPageId="48" captionText="Figs 54–55. Dyscolus spp., habitus. 54. D. verecundior Moret sp. nov., male holotype (MNHN, COI voucher PM136-04). 55. D. verecundissimus Moret sp. nov., male holotype (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.3829716" httpUri="https://zenodo.org/record/3829716/files/figure.png" pageId="46" pageNumber="47">Figs 54</figureCitation>
,
<figureCitation id="1325DFF91366531BFCCFF8B8FCDDF8DE" box="[830,859,1858,1885]" captionStart="Figs 56–59" captionStartId="49.[189,241,1887,1913]" captionTargetBox="[209,1382,913,1794]" captionTargetId="figure@49.[189,1399,907,1816]" captionTargetPageId="49" captionText="Figs 56–59. Group of Dyscolus verecundus Moret, 1998, male aedeagus, median lobe in lateral and dorsal view. 56. D. verecundus, paratype from Rucu Pichincha. 57. D. verecundus, paratype from Guagua Pichincha. 58. D. verecundior Moret sp. nov., holotype (MNHN, COI voucher PM136- 04);.59. D. verecundissimus Moret sp. nov., holotype (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.3829718" httpUri="https://zenodo.org/record/3829718/files/figure.png" pageId="46" pageNumber="47">58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71366531BFF4CF879FC27F85C" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8BA1C37C1366531BFF4CF879FD7BF81E" blockId="46.[189,765,1923,1950]" box="[189,765,1923,1950]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB0EE4521366531BFF4CF879FD7BF81E" box="[189,765,1923,1950]" pageId="46" pageNumber="47">
<treatmentCitation id="0ABFE56D1366531BFF4CF879FDFAF81D" author="Moret P." box="[189,636,1923,1950]" page="20" pageId="46" pageNumber="47" year="1998">
<taxonomicName id="4C1EB8FF1366531BFF4CF879FDFAF81D" ID-CoL="388PZ" authority="Moret, 1998: 20" authorityName="Moret" authorityPageNumber="20" authorityYear="1998" box="[189,636,1923,1950]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1366531BFF4CF879FE32F81E" box="[189,436,1923,1949]" italics="true" pageId="46" pageNumber="47">Dyscolus verecundus</emphasis>
<bibRefCitation id="EF8FBE8D1366531BFE4DF87EFDFAF81D" author="Moret P." box="[444,636,1924,1950]" pageId="46" pageNumber="47" pagination="11 - 28" refId="ref27087" refString="Moret P. 1998. Les Dyscolus de la zone periglaciaire des Andes equatoriennes (Coleoptera, Harpalidae, Platyninae). Bulletin de la Societe entomologique de France 103 (1): 11 - 28." type="journal article" year="1998">Moret, 1998: 20</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="B96A1F6E1366531BFD79F87EFD7BF81E" box="[648,765,1924,1949]" italics="true" pageId="46" pageNumber="47">pro parte.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CF83FFC27F85C" blockId="46.[189,929,1989,2016]" box="[189,929,1989,2016]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB0EE4521366531BFF4CF83FFC27F85C" box="[189,929,1989,2016]" pageId="46" pageNumber="47">
<taxonomicName id="4C1EB8FF1366531BFF4CF83FFDBCF85C" ID-CoL="388PZ" authorityName="Moret" authorityYear="1998" box="[189,570,1989,2015]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="verecundus" subGenus="Dyscolus">
<emphasis id="B96A1F6E1366531BFF4CF83FFEAFF85C" box="[189,297,1989,2015]" italics="true" pageId="46" pageNumber="47">Dyscolus</emphasis>
(
<emphasis id="B96A1F6E1366531BFECBF83FFE20F85C" box="[314,422,1989,2015]" italics="true" pageId="46" pageNumber="47">Dyscolus</emphasis>
)
<emphasis id="B96A1F6E1366531BFE47F83FFDBCF85C" box="[438,570,1989,2015]" italics="true" pageId="46" pageNumber="47">verecundus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABFE56D1366531BFDA9F83FFCA6F85C" author="Moret P." box="[600,800,1989,2016]" page="142" pageId="46" pageNumber="47" year="2005">
<bibRefCitation id="EF8FBE8D1366531BFDA9F83FFCA6F85C" author="Moret P." box="[600,800,1989,2016]" pageId="46" pageNumber="47" refId="ref27190" refString="Moret P. 2005. Los Coleopteros Carabidae del Paramo en los Andes del Ecuador: Sistematica, Ecologia y Biogeografia. Pontificia Universidad Catolica del Ecuador, Centro de Biodiversidad y Ambiente, Monografia 2, Quito." type="book" year="2005">Moret 2005: 142</bibRefCitation>
</treatmentCitation>
,
<emphasis id="B96A1F6E1366531BFCDDF83CFC27F85C" box="[812,929,1990,2015]" italics="true" pageId="46" pageNumber="47">pro parte.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30490F71367531AFF4CFEF1FAE4FED0" box="[189,1378,267,340]" pageId="47" pageNumber="48" type="etymology">
<paragraph id="8BA1C37C1367531AFF4CFEF1FEC2FEA6" blockId="47.[189,324,267,293]" box="[189,324,267,293]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CFEF1FEC2FEA6" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CFEF1FEC2FEA6" bold="true" box="[189,324,267,293]" pageId="47" pageNumber="48">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFEC3FAE4FED0" blockId="47.[189,1378,313,340]" box="[189,1378,313,340]" pageId="47" pageNumber="48">
Comparative form of the Latin adjective ‘
<taxonomicName id="4C1EB8FF1367531AFD56FEC3FCADFED0" authorityName="Moret" authorityYear="1998" box="[679,811,313,339]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFD56FEC3FCADFED0" box="[679,811,313,339]" italics="true" pageId="47" pageNumber="48">verecundus</emphasis>
</taxonomicName>
’, which is the name of a closely related species.
</paragraph>
</subSubSection>
<subSubSection id="C30490F71367531AFF4CFE79FAB4F806" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BA1C37C1367531AFF4CFE79FEEAFE1F" blockId="47.[189,364,386,413]" box="[189,364,386,413]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CFE79FEEAFE1F" bold="true" box="[189,364,386,413]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CFE79FEEAFE1F" bold="true" box="[189,364,386,413]" pageId="47" pageNumber="48">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF1BFE4BFEDAFE48" blockId="47.[189,1399,433,565]" box="[234,348,433,459]" pageId="47" pageNumber="48">
<emphasis id="B96A1F6E1367531AFF1BFE4BFEDAFE48" bold="true" box="[234,348,433,459]" pageId="47" pageNumber="48">
<typeStatus id="54A57DDE1367531AFF1BFE4BFEDAFE48" box="[234,348,433,459]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFE2EFC91FDB6" blockId="47.[189,1399,433,565]" pageId="47" pageNumber="48">
<materialsCitation id="3B76C9211367531AFF4CFE2EFC91FDB6" ID-GBIF-Occurrence="2620303429" collectingDate="2016-07-15" collectionCode="MNHN" collectorName="P. Moret &amp; A. Barragan" country="Ecuador" elevation="4335" latitude="-0.644404" location="Waypoint 136" longLatPrecision="1" longitude="-78.70347" municipality="Volcan Illiniza" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Pichincha Province" typeStatus="holotype">
<collectingCountry id="F30983EC1367531AFF4CFE2EFECDFE6D" box="[189,331,468,494]" name="Ecuador" pageId="47" pageNumber="48">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F51367531AFE87FE28FE0BFE6D" box="[374,397,466,494]" pageId="47" pageNumber="48" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E1367531AFE55FE2EFD15FE6D" box="[420,659,468,494]" country="Ecuador" name="Pichincha" pageId="47" pageNumber="48">Pichincha Province</collectingRegion>
,
<collectingMunicipality id="6BC559061367531AFD56FE2EFCD8FE6D" box="[679,862,468,494]" pageId="47" pageNumber="48">Volcán Illiniza</collectingMunicipality>
, ENE slope,
<location id="8EC195A71367531AFBE0FE2EFB44FE6D" LSID="urn:lsid:plazi:treatment:03B7726A1366531AFDCEF901FAB4F806:8EC195A71367531AFBE0FE2EFB44FE6D" box="[1041,1218,468,494]" country="Ecuador" latitude="-0.644404" longLatPrecision="1" longitude="-78.70347" municipality="Volcan Illiniza" name="Waypoint 136" pageId="47" pageNumber="48" stateProvince="Pichincha Province">Waypoint 136</location>
;
<geoCoordinate id="EE2AA5BB1367531AFB26FE2EFAF7FE6D" box="[1239,1393,468,494]" degrees="0.644404" direction="south" orientation="latitude" pageId="47" pageNumber="48" precision="1" value="-0.644404">0.644404º S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB1367531AFF4CFE0DFEEDFD91" box="[189,363,503,530]" degrees="78.703469" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="1" value="-78.70347">78.703469º W</geoCoordinate>
;
<elevation id="0033244F1367531AFE8AFE0DFD9DFD92" box="[379,539,503,530]" metricMagnitude="3" metricUnit="m" metricValue="4.335" pageId="47" pageNumber="48" unit="m" value="4335.0">
<quantity id="4CE66E991367531AFE8AFE0DFE51FD91" box="[379,471,503,530]" metricMagnitude="3" metricUnit="m" metricValue="4.335" pageId="47" pageNumber="48" unit="m" value="4335.0">4335 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC1367531AFDDDFE0DFD40FD92" box="[556,710,503,530]" pageId="47" pageNumber="48" value="2016-07-15">
<collectingDate id="EFE41C541367531AFDDDFE0DFD40FD92" box="[556,710,503,530]" pageId="47" pageNumber="48" value="2016-07-15">15 Jul. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA1367531AFD27FE02FCBBFD91" box="[726,829,504,530]" pageId="47" pageNumber="48">P. Moret</collectorName>
and
<collectorName id="26EBA6AA1367531AFC8DFE02FB89FD92" box="[892,1039,503,530]" pageId="47" pageNumber="48">A. Barragán</collectorName>
leg.; COI voucher in ethanol PM136-04, BOLD sequence SUM062-18;
<collectionCode id="ED0F5BB91367531AFD44FDE1FC95FDB6" box="[693,787,539,565]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="48">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C1367531AFF1BFD9EFE43FDFD" blockId="47.[189,1399,610,779]" box="[234,453,610,638]" pageId="47" pageNumber="48">
<emphasis id="B96A1F6E1367531AFF1BFD9EFEE1FDFD" bold="true" box="[234,359,612,638]" pageId="47" pageNumber="48">
<typeStatus id="54A57DDE1367531AFF1BFD9EFEE1FDFD" box="[234,359,612,638]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 ♂♂)
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFD7DFD37FC89" blockId="47.[189,1399,610,779]" pageId="47" pageNumber="48">
<materialsCitation id="3B76C9211367531AFF4CFD7DFAE3FD47" ID-GBIF-Occurrence="2620303328" collectedFrom="Polylepis forest" collectingDate="1985-08-07" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="4080" location="Volcan Illiniza" municipality="Volcan Illiniza" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha Province">
<collectingCountry id="F30983EC1367531AFF4CFD7DFECDFD22" box="[189,331,647,673]" name="Ecuador" pageId="47" pageNumber="48">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E1367531AFE84FD7DFDEAFD22" bold="true" box="[373,620,647,673]" pageId="47" pageNumber="48">
<collectingRegion id="49DA0D9E1367531AFE84FD7DFDEAFD22" box="[373,620,647,673]" country="Ecuador" name="Pichincha" pageId="47" pageNumber="48">Pichincha Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F51367531AFD63FD7DFD43FD22" box="[658,709,645,673]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
(red printed label: “
<taxonomicName id="4C1EB8FF1367531AFC3DFD7DFAF7FD22" authority="Moret, 1998" authorityName="Moret" authorityYear="1998" box="[972,1393,647,673]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFC3DFD7DFB4FFD22" box="[972,1225,647,673]" italics="true" pageId="47" pageNumber="48">Dyscolus verecundus</emphasis>
<bibRefCitation id="EF8FBE8D1367531AFB29FD7DFAF7FD22" author="Moret P." box="[1240,1393,647,673]" pageId="47" pageNumber="48" pagination="11 - 28" refId="ref27087" refString="Moret P. 1998. Les Dyscolus de la zone periglaciaire des Andes equatoriennes (Coleoptera, Harpalidae, Platyninae). Bulletin de la Societe entomologique de France 103 (1): 11 - 28." type="journal article" year="1998">Moret, 1998</bibRefCitation>
</taxonomicName>
, Paratype”);
<collectingMunicipality id="6BC559061367531AFEB8FD50FE70FD47" box="[329,502,682,708]" pageId="47" pageNumber="48">Volcán Illiniza</collectingMunicipality>
, NE slope;
<quantity id="4CE66E991367531AFD8DFD50FD53FD47" box="[636,725,682,708]" metricMagnitude="3" metricUnit="m" metricValue="4.08" pageId="47" pageNumber="48" unit="m" value="4080.0">
<elevation id="0033244F1367531AFD8DFD50FD53FD47" box="[636,725,682,708]" metricMagnitude="3" metricUnit="m" metricValue="4.08" pageId="47" pageNumber="48" unit="m" value="4080.0">4080 m</elevation>
</quantity>
a.s.l.;
<date id="FFA0E5BC1367531AFCD0FD50FC35FD46" box="[801,947,682,709]" pageId="47" pageNumber="48" value="1985-08-07">
<collectingDate id="EFE41C541367531AFCD0FD50FC35FD46" box="[801,947,682,709]" pageId="47" pageNumber="48" value="1985-08-07">7 Aug. 1985</collectingDate>
</date>
;
<collectorName id="26EBA6AA1367531AFC4FFD50FBA7FD47" box="[958,1057,682,708]" pageId="47" pageNumber="48">P. Moret</collectorName>
leg.;
<collectedFrom id="380280811367531AFB93FD50FA9BFD47" box="[1122,1309,682,708]" pageId="47" pageNumber="48">
<taxonomicName id="4C1EB8FF1367531AFB93FD50FB49FD47" box="[1122,1231,682,708]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E1367531AFB93FD50FB49FD47" box="[1122,1231,682,708]" italics="true" pageId="47" pageNumber="48">Polylepis</emphasis>
</taxonomicName>
forest
</collectedFrom>
;
<collectionCode id="ED0F5BB91367531AFAD6FD50FAE3FD47" box="[1319,1381,682,708]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="47" pageNumber="48">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C9211367531AFF4CFD37FD37FC89" ID-GBIF-Occurrence="2620303316" collectingDate="1986-01-24" collectionCode="CPM" collectorName="P. Moret" pageId="47" pageNumber="48" specimenCount="4451" specimenCount-male="4451">
<specimenCount id="9D1808F51367531AFF4CFD37FF6EFD64" box="[189,232,715,743]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
(red printed label: “
<taxonomicName id="4C1EB8FF1367531AFE23FD37FCE7FD64" authority="Moret, 1998" authorityName="Moret" authorityYear="1998" box="[466,865,717,744]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFE23FD37FD4EFD64" box="[466,712,717,743]" italics="true" pageId="47" pageNumber="48">Dyscolus verecundus</emphasis>
<bibRefCitation id="EF8FBE8D1367531AFD3FFD34FCE7FD64" author="Moret P." box="[718,865,717,744]" pageId="47" pageNumber="48" pagination="11 - 28" refId="ref27087" refString="Moret P. 1998. Les Dyscolus de la zone periglaciaire des Andes equatoriennes (Coleoptera, Harpalidae, Platyninae). Bulletin de la Societe entomologique de France 103 (1): 11 - 28." type="journal article" year="1998">Moret, 1998</bibRefCitation>
</taxonomicName>
, Paratype”); E slope of Volcán Corazón;
<quantity id="4CE66E991367531AFACDFD37FF52FC88" metricMagnitude="3" metricUnit="m" metricValue="4.45" pageId="47" pageNumber="48" unit="m" value="4450.0">
<specimenCount id="9D1808F51367531AFACDFD37FF52FC88" pageId="47" pageNumber="48" type="male">4450 m</specimenCount>
</quantity>
a.s.l.;
<date id="FFA0E5BC1367531AFED0FD0AFE3CFC89" box="[289,442,752,779]" pageId="47" pageNumber="48" value="1986-01-24">
<collectingDate id="EFE41C541367531AFED0FD0AFE3CFC89" box="[289,442,752,779]" pageId="47" pageNumber="48" value="1986-01-24">24 Jan. 1986</collectingDate>
</date>
;
<collectorName id="26EBA6AA1367531AFE37FD0BFDACFC88" box="[454,554,753,779]" pageId="47" pageNumber="48">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB91367531AFD9AFD0AFD2BFC89" box="[619,685,752,778]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="47" pageNumber="48">CPM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFCC3FE52FCD0" blockId="47.[189,468,825,851]" box="[189,468,825,851]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CFCC3FE52FCD0" bold="true" box="[189,468,825,851]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CFCC3FE52FCD0" bold="true" box="[189,468,825,851]" pageId="47" pageNumber="48">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFC92FE2CFAC8" blockId="47.[189,1399,872,1356]" pageId="47" pageNumber="48">
Habitus:
<figureCitation id="1325DFF91367531AFEDAFC92FE03FC00" box="[299,389,872,899]" captionStart="Figs 54–55" captionStartId="48.[189,241,1964,1990]" captionTargetBox="[254,1364,851,1900]" captionTargetId="figure@48.[203,1385,843,1920]" captionTargetPageId="48" captionText="Figs 54–55. Dyscolus spp., habitus. 54. D. verecundior Moret sp. nov., male holotype (MNHN, COI voucher PM136-04). 55. D. verecundissimus Moret sp. nov., male holotype (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.3829716" httpUri="https://zenodo.org/record/3829716/files/figure.png" pageId="47" pageNumber="48">Fig. 54</figureCitation>
. Wingless. Body length: 10.8–11.0 mm. Body colour variable, from reddish brown to piceous black, the vertex and the sutural zone of the elytra reddish; legs, antennae and mouthparts reddish. Elytral microsculpture isodiametric, distinctly impressed. Head robust, convex, neck strongly constricted, eyes moderately bulging, genae almost flat in dorsal view; antennae short. Pronotum transverse, subquadrate; sides moderately arcuate distally, weakly sinuate in basal third; hind angles variable, obtuse or almost quadrate, but always blunt; two pairs of lateral setae. Elytra subparallel, disc almost depressed; humeri rounded but not effaced, subapical sinuation obsolete; striae thin, shallowly impressed; intervals flat; third elytral interval with 3–5 discal setae. Fourth metatarsomere triangular, with one pair of lateroapical setae and without subapical dorsolateral setae; apical lobes very short, the inner one not extending beyond the insertion point of the lateroapical seta; fifth metatarsomere asetose ventrally. Last visible abdominal ventrite of the male with one pair of setae along its apical margin (♂).
<emphasis id="B96A1F6E1367531AFF08FB11FE2FFA86" box="[249,425,1259,1285]" italics="true" pageId="47" pageNumber="48">Male genitalia</emphasis>
:
<figureCitation id="1325DFF91367531AFE4CFB11FD9EFA85" box="[445,536,1259,1286]" captionStart="Figs 56–59" captionStartId="49.[189,241,1887,1913]" captionTargetBox="[209,1382,913,1794]" captionTargetId="figure@49.[189,1399,907,1816]" captionTargetPageId="49" captionText="Figs 56–59. Group of Dyscolus verecundus Moret, 1998, male aedeagus, median lobe in lateral and dorsal view. 56. D. verecundus, paratype from Rucu Pichincha. 57. D. verecundus, paratype from Guagua Pichincha. 58. D. verecundior Moret sp. nov., holotype (MNHN, COI voucher PM136- 04);.59. D. verecundissimus Moret sp. nov., holotype (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.3829718" httpUri="https://zenodo.org/record/3829718/files/figure.png" pageId="47" pageNumber="48">Fig. 58</figureCitation>
. Median lobe weakly arcuate; apical blade bluntly triangular in dorsal view; endophallus with two sclerotized squamose areas, ovoid in shape, median and subapical.
<emphasis id="B96A1F6E1367531AFAEEFAF4FEA3FAC8" italics="true" pageId="47" pageNumber="48">Female genitalia</emphasis>
: unknown.
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFA80FEE5FA17" blockId="47.[189,355,1402,1428]" box="[189,355,1402,1428]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CFA80FEE5FA17" bold="true" box="[189,355,1402,1428]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CFA80FEE5FA17" bold="true" box="[189,355,1402,1428]" pageId="47" pageNumber="48">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CFA53FE88F915" blockId="47.[189,1399,1449,1686]" pageId="47" pageNumber="48">
This species closely resembles
<taxonomicName id="4C1EB8FF1367531AFDBBFA50FD7FFA40" authorityName="Moret" authorityYear="1998" box="[586,761,1449,1475]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFDBBFA50FD7FFA40" box="[586,761,1449,1475]" italics="true" pageId="47" pageNumber="48">D. verecundus</emphasis>
</taxonomicName>
, now restricted to Mt
<collectingRegion id="49DA0D9E1367531AFBD2FA53FB1CFA40" box="[1059,1178,1449,1475]" country="Ecuador" name="Pichincha" pageId="47" pageNumber="48">Pichincha</collectingRegion>
, its
<typeStatus id="54A57DDE1367531AFB29FA50FA8DFA47" box="[1240,1291,1450,1476]" pageId="47" pageNumber="48">type</typeStatus>
locality.
<taxonomicName id="4C1EB8FF1367531AFF4CFA36FD8FFA64" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,521,1484,1511]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundior" status="sp. nov.">
<emphasis id="B96A1F6E1367531AFF4CFA36FE3DFA65" box="[189,443,1484,1510]" italics="true" pageId="47" pageNumber="48">Dyscolus verecundior</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151367531AFDFEFA37FDEEFA64" box="[527,616,1485,1511]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C1EB8FF1367531AFD0FFA37FC23FA65" authorityName="Moret" authorityYear="1998" box="[766,933,1484,1510]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFD0FFA37FC23FA65" box="[766,933,1484,1510]" italics="true" pageId="47" pageNumber="48">D. verecundus</emphasis>
</taxonomicName>
in a few external and genital characters: genae almost flat (convex in
<taxonomicName id="4C1EB8FF1367531AFDE4FA0AFD38F98A" authorityName="Moret" authorityYear="1998" box="[533,702,1519,1545]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFDE4FA0AFD38F98A" box="[533,702,1519,1545]" italics="true" pageId="47" pageNumber="48">D. verecundus</emphasis>
</taxonomicName>
); antennae slightly thicker, the second antennomere of the male 1.41 to 1.49 times as long as wide (1.50 to 1.65 times in
<taxonomicName id="4C1EB8FF1367531AFC42F9E9FBDBF9AE" authorityName="Moret" authorityYear="1998" box="[947,1117,1555,1581]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFC42F9E9FBDBF9AE" box="[947,1117,1555,1581]" italics="true" pageId="47" pageNumber="48">D. verecundus</emphasis>
</taxonomicName>
); fourth metatarsomere thinner, 1.16 to 1.20 times as long as wide (1.07 to 1.13 times in
<taxonomicName id="4C1EB8FF1367531AFC22F9CDFBFBF9D3" authorityName="Moret" authorityYear="1998" box="[979,1149,1590,1616]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1367531AFC22F9CDFBFBF9D3" box="[979,1149,1590,1616]" italics="true" pageId="47" pageNumber="48">D. verecundus</emphasis>
</taxonomicName>
); apical blade of the median lobe thinner in lateral view (compare
<figureCitation id="1325DFF91367531AFD2BF9A3FC3CF9F7" box="[730,954,1625,1652]" captionStart="Figs 56–59" captionStartId="49.[189,241,1887,1913]" captionTargetBox="[209,1382,913,1794]" captionTargetId="figure@49.[189,1399,907,1816]" captionTargetPageId="49" captionText="Figs 56–59. Group of Dyscolus verecundus Moret, 1998, male aedeagus, median lobe in lateral and dorsal view. 56. D. verecundus, paratype from Rucu Pichincha. 57. D. verecundus, paratype from Guagua Pichincha. 58. D. verecundior Moret sp. nov., holotype (MNHN, COI voucher PM136- 04);.59. D. verecundissimus Moret sp. nov., holotype (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.3829718" httpUri="https://zenodo.org/record/3829718/files/figure.png" pageId="47" pageNumber="48">Figs 56–57 and 58</figureCitation>
); sclerotized areas of the endophallus bigger.
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CF93FFE9BF95C" blockId="47.[189,285,1733,1759]" box="[189,285,1733,1759]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CF93FFE9BF95C" bold="true" box="[189,285,1733,1759]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CF93FFE9BF95C" bold="true" box="[189,285,1733,1759]" pageId="47" pageNumber="48">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CF90EFD61F88D" blockId="47.[189,743,1779,1806]" box="[189,743,1779,1806]" pageId="47" pageNumber="48">
Lower superpáramo from
<quantity id="4CE66E991367531AFE01F90EFD2FF88D" box="[496,681,1779,1806]" metricMagnitude="3" metricUnit="m" metricValue="4.265" metricValueMax="4.45" metricValueMin="4.08" pageId="47" pageNumber="48" unit="m" value="4265.0" valueMax="4450.0" valueMin="4080.0">4080 to 4450 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CF8C6FE6DF8D5" blockId="47.[189,491,1852,1878]" box="[189,491,1852,1878]" pageId="47" pageNumber="48">
<heading id="D0E974101367531AFF4CF8C6FE6DF8D5" bold="true" box="[189,491,1852,1878]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96A1F6E1367531AFF4CF8C6FE6DF8D5" bold="true" box="[189,491,1852,1878]" pageId="47" pageNumber="48">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1367531AFF4CF891FAB4F806" blockId="47.[189,1330,1899,1926]" box="[189,1330,1899,1926]" pageId="47" pageNumber="48">Microendemic species, restricted to the Illiniza and Corazón volcanoes in the Western Cordillera.</paragraph>
</subSubSection>
</treatment>
</document>
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