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<document id="8D619E05E49A9A28E9D09C6B55700491" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A1364531BFDA2FA6EFAD7FCCB" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus silvestris Moret 2020, sp. nov." docType="treatment" docUuid="8DA5B9B7-F332-4D75-AC2D-CEBBD34ADA3C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="47" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="45" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="AC25902F215F52EDD0B13DFD3844BC5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B144B973645D13F94EB9E54D42CB9E40">
<mods:title id="746DF46F1106AD1ACA318E43EFF8E487">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="8CC4BE03336588350C9CF755B9FA8341" type="personal">
<mods:role id="0C2CA1CD516527C51538B28B5A996E27">
<mods:roleTerm id="7FCC2ED30F455C137FF0F54E96C30D88">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68F2B49E407DFF6FE8C27EFEC875D6A7">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="3B4052002237196CDF3231AD5BAE96CE" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="C48A051545A4797D08C942AE907AF47A">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="61D11A2F61DDD17B44B13ACB593BE658" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="8C468DC3DE1A18FAFDEA584D19B16406" type="personal">
<mods:role id="183DB38666618A605CC24733608660C2">
<mods:roleTerm id="F77ABB464600DB2EB36E8E417243348A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F487A45947568F854F681C4A1FDF446">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="7A33360E7A96F4EA00DD07270120DE0A" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="7C6C0C09F7182FEB3133F6CAE528EA89">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="891AF75492D0C9A3E3F073AF1F2DC93F" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="363464A4663DED0CEBCE22E8868908F0">text</mods:typeOfResource>
<mods:relatedItem id="6390BA27AB4715497005C6E2A50FF289" type="host">
<mods:titleInfo id="695437AED73CEDED059464F4E88BA872">
<mods:title id="D8F819A9E27281E9B802C5CFBA8670E8">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="5B71F7A6F41D54C96AC5D3B4448BADF2">
<mods:date id="3E43ED69223D220C37AB1394745FE9F7">2020</mods:date>
<mods:detail id="F09A3BC856B4E69FD206387708F18C1F" type="pubDate">
<mods:number id="089BB4E5B3EB88E0816D9ACF0DF23BED">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="687F75780490EBA2FA4A2F9B315AB7D3" type="volume">
<mods:number id="C254232CDB56792335A07576CB41E69D">646</mods:number>
</mods:detail>
<mods:extent id="084D265C08C43AE36CA7AFF845E037F9" unit="page">
<mods:start id="4F0A1BA5F67A63D08837B40ED09DFC81">1</mods:start>
<mods:end id="464D0816487CCA3EA7227A24EE6B39F9">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B3F1B0FFDE5C2F91D9E9A5487CECA4C4">journal article</mods:classification>
<mods:identifier id="4C2CD6A1EF6B5B96D57730A6471ED3A2" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="D95DF73F863B01F829CE7F0213797752" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="E4FFAE54110F20487CC91A0A62D25B9E" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="FA7BB9F2161F0379646989AEC25C3BEC" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="2BF42E865BBE6BE7BF4BCDD5A8A90166" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A1364531BFDA2FA6EFAD7FCCB" ID-DOI="http://doi.org/10.5281/zenodo.3848391" ID-GBIF-Taxon="164160398" ID-Zenodo-Dep="3848391" LSID="urn:lsid:zoobank.org:act:8DA5B9B7-F332-4D75-AC2D-CEBBD34ADA3C" httpUri="http://treatment.plazi.org/id/03B7726A1364531BFDA2FA6EFAD7FCCB" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45">
<subSubSection id="C30490F713645319FDA2FA6EFB5DFA51" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BA1C37C13645319FDA2FA6EFC67FA2C" blockId="44.[344,1243,1428,1526]" box="[595,993,1428,1455]" pageId="44" pageNumber="45">
<heading id="D0E9741013645319FDA2FA6EFC67FA2C" box="[595,993,1428,1455]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="45" reason="2">
<taxonomicName id="4C1EB8FF13645319FDA2FA6EFCF9FA2C" ID-CoL="8MH8B" authority="Moret" authorityName="Moret" authorityYear="2020" box="[595,895,1428,1455]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="silvestris" status="sp. nov.">
<emphasis id="B96A1F6E13645319FDA2FA6EFCB6FA2C" bold="true" box="[595,816,1428,1455]" italics="true" pageId="44" pageNumber="45">Dyscolus silvestris</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513645319FC77FA6FFC67FA2C" box="[902,993,1429,1455]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C13645319FEA9FA4DFB5DFA51" blockId="44.[344,1243,1428,1526]" box="[344,1243,1463,1490]" pageId="44" pageNumber="45">
<uri id="FF8FCF7E13645319FEA9FA4DFB5DFA51" box="[344,1243,1463,1490]" pageId="44" pageNumber="45">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A913645319FD89FA4DFB5DFA51" box="[632,1243,1463,1490]" pageId="44" pageNumber="45">8DA5B9B7-F332-4D75-AC2D-CEBBD34ADA3C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713645319FD29FA21FCDDFA75" box="[728,859,1499,1526]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BA1C37C13645319FD29FA21FCDDFA75" blockId="44.[344,1243,1428,1526]" box="[728,859,1499,1526]" pageId="44" pageNumber="45">
<figureCitation id="1325DFF913645319FD29FA21FCDDFA75" box="[728,859,1499,1526]" captionStart="Figs 52–53" captionStartId="45.[189,241,1955,1981]" captionTargetBox="[351,1238,831,1892]" captionTargetId="figure@45.[334,1253,818,1914]" captionTargetPageId="45" captionText="Figs 52–53. Dyscolus silvestris Moret sp. nov., male holotype (QCAZ). 52. Habitus. 53. Male aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829714" httpUri="https://zenodo.org/record/3829714/files/figure.png" pageId="44" pageNumber="45">Figs 52–53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713645319FF4CF9E6FD13F9E6" box="[189,661,1564,1637]" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8BA1C37C13645319FF4CF9E6FEC2F9B5" blockId="44.[189,324,1564,1590]" box="[189,324,1564,1590]" pageId="44" pageNumber="45">
<heading id="D0E9741013645319FF4CF9E6FEC2F9B5" bold="true" box="[189,324,1564,1590]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B96A1F6E13645319FF4CF9E6FEC2F9B5" bold="true" box="[189,324,1564,1590]" pageId="44" pageNumber="45">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13645319FF4CF9B1FD13F9E6" blockId="44.[189,661,1611,1637]" box="[189,661,1611,1637]" pageId="44" pageNumber="45">Latin adjective meaning ‘forest related’.</paragraph>
</subSubSection>
<subSubSection id="C30490F71364531BFF4CF976FAD7FCCB" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BA1C37C13645319FF4CF976FEEAF925" blockId="44.[189,364,1676,1702]" box="[189,364,1676,1702]" pageId="44" pageNumber="45">
<heading id="D0E9741013645319FF4CF976FEEAF925" bold="true" box="[189,364,1676,1702]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B96A1F6E13645319FF4CF976FEEAF925" bold="true" box="[189,364,1676,1702]" pageId="44" pageNumber="45">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13645319FF1BF941FEDAF956" blockId="44.[189,1398,1723,1819]" box="[234,348,1723,1749]" pageId="44" pageNumber="45">
<emphasis id="B96A1F6E13645319FF1BF941FEDAF956" bold="true" box="[234,348,1723,1749]" pageId="44" pageNumber="45">
<typeStatus id="54A57DDE13645319FF1BF941FEDAF956" box="[234,348,1723,1749]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C13645319FF4CF924FEE4F898" blockId="44.[189,1398,1723,1819]" pageId="44" pageNumber="45">
<materialsCitation id="3B76C92113645319FF4CF924FEE4F898" ID-GBIF-Occurrence="2620303319" collectingDate="2001-07-01" collectionCode="QCAZ" collectorName="P. Moret" country="Ecuador" elevation="3300" location="Termas de Papallacta" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Napo Province" typeStatus="holotype">
<collectingCountry id="F30983EC13645319FF4CF924FECDF97B" box="[189,331,1758,1784]" name="Ecuador" pageId="44" pageNumber="45">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F513645319FE95F926FEFDF97B" box="[356,379,1756,1784]" pageId="44" pageNumber="45" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E13645319FE78F924FDBDF97B" box="[393,571,1758,1784]" country="Ecuador" name="Napo" pageId="44" pageNumber="45">Napo Province</collectingRegion>
,
<location id="8EC195A713645319FDB6F924FCC5F97B" LSID="urn:lsid:plazi:treatment:03B7726A1364531BFDA2FA6EFAD7FCCB:8EC195A713645319FDB6F924FCC5F97B" box="[583,835,1758,1784]" country="Ecuador" name="Termas de Papallacta" pageId="44" pageNumber="45" stateProvince="Napo Province">Termas de Papallacta</location>
;
<elevation id="0033244F13645319FCBEF924FC6EF97B" box="[847,1000,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="44" pageNumber="45" unit="m" value="3300.0">
<quantity id="4CE66E9913645319FCBEF924FC2EF97B" box="[847,936,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="44" pageNumber="45" unit="m" value="3300.0">3300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13645319FC04F924FBFCF97B" box="[1013,1146,1758,1784]" pageId="44" pageNumber="45" value="2001-07-01">
<collectingDate id="EFE41C5413645319FC04F924FBFCF97B" box="[1013,1146,1758,1784]" pageId="44" pageNumber="45" value="2001-07-01">1 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA13645319FB77F924FB6CF97B" box="[1158,1258,1758,1784]" pageId="44" pageNumber="45">P. Moret</collectorName>
leg.; “sousbois”;
<collectionCode id="ED0F5BB913645319FEFBF8FBFED8F898" box="[266,350,1793,1819]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="44" pageNumber="45" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C13645319FF1BF8B9FDA2F8DE" blockId="44.[189,1399,1857,2026]" box="[234,548,1857,1885]" pageId="44" pageNumber="45">
<emphasis id="B96A1F6E13645319FF1BF8B9FEE1F8DE" bold="true" box="[234,359,1859,1885]" pageId="44" pageNumber="45">
<typeStatus id="54A57DDE13645319FF1BF8B9FEE1F8DE" box="[234,359,1859,1885]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
</emphasis>
(5 ♂♂,
<specimenCount id="9D1808F513645319FE3BF8B9FD9EF8DE" box="[458,536,1859,1885]" pageId="44" pageNumber="45" type="female">10 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C13645318FF4CF89CFB75FE0C" blockId="44.[189,1399,1857,2026]" lastBlockId="45.[189,1399,267,399]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<materialsCitation id="3B76C92113645319FF4CF89CFB5AF803" ID-GBIF-Occurrence="2620303318" box="[189,1244,1892,1920]" collectingDate="2001-07-01" collectionCode="CPM" collectorName="de Papallacta &amp; P. Moret" country="Ecuador" elevation="3300" location="Napo Province" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo Province">
<collectingCountry id="F30983EC13645319FF4CF89CFECDF803" box="[189,331,1894,1920]" name="Ecuador" pageId="44" pageNumber="45">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E13645319FE81F89CFDAAF803" bold="true" box="[368,556,1894,1920]" pageId="44" pageNumber="45">
<collectingRegion id="49DA0D9E13645319FE81F89CFDAAF803" box="[368,556,1894,1920]" country="Ecuador" name="Napo" pageId="44" pageNumber="45">Napo Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F513645319FDBDF89CFDFAF803" box="[588,636,1892,1920]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F513645319FD7FF89CFD39F803" box="[654,703,1894,1920]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB913645319FB6FF89CFB5AF803" box="[1182,1244,1894,1920]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="44" pageNumber="45">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113645319FB0DF89CFE3DF844" ID-GBIF-Occurrence="2620303370" collectingDate="2015-10-25" collectionCode="MNHN" collectorName="P. Moret" country="Ecuador" county="Papallacta" elevation="3410" latitude="-0.37350002" location="Waypoint 41" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" pageId="44" pageNumber="45" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo">
<specimenCount id="9D1808F513645319FB0DF89CFAAAF803" box="[1276,1324,1892,1920]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F513645319FACEF89CFAE9F803" box="[1343,1391,1894,1920]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62C0BBF013645319FF4CF873FEBFF820" box="[189,313,1929,1955]" pageId="44" pageNumber="45">Papallacta</collectingCounty>
,
<collectingMunicipality id="6BC5590613645319FEB7F870FD96F827" box="[326,528,1930,1956]" pageId="44" pageNumber="45">Termas Jamanco</collectingMunicipality>
,
<location id="8EC195A713645319FDECF873FD3EF820" LSID="urn:lsid:plazi:treatment:03B7726A1364531BFDA2FA6EFAD7FCCB:8EC195A713645319FDECF873FD3EF820" box="[541,696,1929,1955]" country="Ecuador" county="Papallacta" latitude="-0.37350002" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" name="Waypoint 41" pageId="44" pageNumber="45" stateProvince="Napo">Waypoint 41</location>
;
<geoCoordinate id="EE2AA5BB13645319FD36F87DFCE7F820" box="[711,865,1927,1955]" degrees="0" direction="south" minutes="22" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="24.6" value="-0.37350002">0°22′24.6″ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB13645319FC81F87DFB93F827" box="[880,1045,1927,1956]" degrees="78" direction="west" minutes="10" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="5.5" value="-78.16819">78°10′5.5″ W</geoCoordinate>
;
<elevation id="0033244F13645319FBD5F873FB47F820" box="[1060,1217,1929,1956]" metricMagnitude="3" metricUnit="m" metricValue="3.41" pageId="44" pageNumber="45" unit="m" value="3410.0">
<quantity id="4CE66E9913645319FBD5F873FBF9F827" box="[1060,1151,1929,1956]" metricMagnitude="3" metricUnit="m" metricValue="3.41" pageId="44" pageNumber="45" unit="m" value="3410.0">3410 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13645319FB20F873FAF4F827" box="[1233,1394,1929,1956]" pageId="44" pageNumber="45" value="2015-10-25">
<collectingDate id="EFE41C5413645319FB20F873FAF4F827" box="[1233,1394,1929,1956]" pageId="44" pageNumber="45" value="2015-10-25">25 Oct. 2015</collectingDate>
</date>
;
<collectorName id="26EBA6AA13645319FF4CF857FEA6F844" box="[189,288,1965,1991]" pageId="44" pageNumber="45">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB913645319FE90F857FE3DF844" box="[353,443,1965,1991]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="44" pageNumber="45">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113645319FE22F856FB98F845" ID-GBIF-Occurrence="2620303391" box="[467,1054,1964,1991]" collectingDate="2015-10-25" collectionCode="QCAZ" collectorName="P. Moret" country="Ecuador" county="Papallacta" elevation="3410" latitude="-0.37350002" location="Waypoint" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="9D1808F513645319FE22F856FE79F844" box="[467,511,1964,1991]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB913645319FC3FF856FB98F845" box="[974,1054,1964,1990]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="44" pageNumber="45" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113645319FBC4F856FD88F869" ID-GBIF-Occurrence="2620303313" collectingDate="2015-10-25" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" county="Papallacta" elevation="3410" latitude="-0.37350002" location="Waypoint" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" pageId="44" pageNumber="45" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Napo">
<specimenCount id="9D1808F513645319FBC4F856FBE7F845" box="[1077,1121,1962,1990]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F513645319FB9EF856FB34F844" box="[1135,1202,1964,1991]" pageId="44" pageNumber="45" type="female">3 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB913645319FE21F82AFD88F869" box="[464,526,2000,2026]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="44" pageNumber="45">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113645318FDDEF82AFDF2FEA6" ID-GBIF-Occurrence="2620303308" collectingDate="2015-10-25" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" county="Papallacta" elevation="3410" lastPageId="45" lastPageNumber="46" latitude="-0.37350002" location="Waypoint" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<specimenCount id="9D1808F513645319FDDEF82AFDD9F869" box="[559,607,1998,2026]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
; same collection data as for preceding; COI voucher PM041-02, BOLD sequence SUM103-18;
<collectionCode id="ED0F5BB913655318FDC7FEF1FDF2FEA6" box="[566,628,267,293]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="45" pageNumber="46">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113655318FD67FEF1FCD7FECB" ID-GBIF-Occurrence="2620303333" collectingDate="2015-10-25" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" county="Papallacta" elevation="3410" latitude="-0.37350002" location="Waypoint" longLatPrecision="1" longitude="-78.16819" municipality="Termas Jamanco" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="9D1808F513655318FD67FEF1FD41FEA5" box="[662,711,267,294]" pageId="45" pageNumber="46" type="female">1 ♀</specimenCount>
; same collection data as for preceding; COI voucher in ethanol PM041-06, BOLD sequence SUM104-18;
<collectionCode id="ED0F5BB913655318FCE2FED4FCD7FECB" box="[787,849,302,328]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="45" pageNumber="46">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113655318FC9AFED4FE3FFEEF" ID-GBIF-Occurrence="2620303405" collectingDate="1986-04-09" collectionCode="CPM" collectorName="A. Vigna" country="Ecuador" elevation="3370" location="Lago Papallacta" pageId="45" pageNumber="46" specimenCount="2" specimenCount-female="2" stateProvince="Napo">
<specimenCount id="9D1808F513655318FC9AFED4FC28FECA" box="[875,942,302,329]" pageId="45" pageNumber="46" type="female">2 ♀♀</specimenCount>
;
<location id="8EC195A713655318FC4DFED5FBFAFECB" LSID="urn:lsid:plazi:treatment:03B7726A1364531BFDA2FA6EFAD7FCCB:8EC195A713655318FC4DFED5FBFAFECB" box="[956,1148,302,329]" country="Ecuador" name="Lago Papallacta" pageId="45" pageNumber="46" stateProvince="Napo">Lago Papallacta</location>
;
<elevation id="0033244F13655318FB79FED4FAA7FECB" box="[1160,1313,302,329]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="45" pageNumber="46" unit="m" value="3370.0">
<quantity id="4CE66E9913655318FB79FED4FB67FECA" box="[1160,1249,302,329]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="45" pageNumber="46" unit="m" value="3370.0">3370 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13655318FADEFED4FF7DFEE8" pageId="45" pageNumber="46" value="1986-04-09">
<collectingDate id="EFE41C5413655318FADEFED4FF7DFEE8" pageId="45" pageNumber="46" value="1986-04-09">9 Apr. 1986</collectingDate>
</date>
;
<collectorName id="26EBA6AA13655318FEF5FEA8FEF7FEE8" box="[260,369,337,364]" pageId="45" pageNumber="46">A. Vigna</collectorName>
;
<collectionCode id="ED0F5BB913655318FE8AFEA8FE3FFEEF" box="[379,441,338,364]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="45" pageNumber="46">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113655318FE21FEA8FB6CFEEF" ID-GBIF-Occurrence="2620303375" box="[464,1258,337,364]" collectingDate="1979-04-06" collectingDateMax="1979-04-08" collectingDateMin="1979-04-06" collectionCode="UASM" collectorName="H. Frania" country="Ecuador" elevation="2700" location="Papallacta" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Napo">
<specimenCount id="9D1808F513655318FE21FEA8FE7AFEEF" box="[464,508,338,364]" pageId="45" pageNumber="46" type="female">1 ♀</specimenCount>
;
<location id="8EC195A713655318FDFBFEABFD00FEE8" LSID="urn:lsid:plazi:treatment:03B7726A1364531BFDA2FA6EFAD7FCCB:8EC195A713655318FDFBFEABFD00FEE8" box="[522,646,337,363]" country="Ecuador" name="Papallacta" pageId="45" pageNumber="46" stateProvince="Napo">Papallacta</location>
;
<elevation id="0033244F13655318FD60FEA8FCA1FEE8" box="[657,807,337,364]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="45" pageNumber="46" unit="m" value="2700.0">
<quantity id="4CE66E9913655318FD60FEA8FD6FFEEF" box="[657,745,338,364]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="45" pageNumber="46" unit="m" value="2700.0">2700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13655318FCC5FEABFC5AFEEF" box="[820,988,337,364]" pageId="45" pageNumber="46" value="1979-04-06" valueMax="1979-04-08" valueMin="1979-04-06">
<collectingDate id="EFE41C5413655318FCC5FEABFC5AFEEF" box="[820,988,337,364]" pageId="45" pageNumber="46" value="1979-04-06" valueMax="1979-04-08" valueMin="1979-04-06">6–8 Apr. 1979</collectingDate>
</date>
;
<collectorName id="26EBA6AA13655318FC16FEA8FBD3FEE8" box="[999,1109,337,364]" pageId="45" pageNumber="46">H. Frania</collectorName>
leg.;
<collectionCode id="ED0F5BB913655318FB64FEA8FB6CFEEF" box="[1173,1258,338,364]" collectionName="Canada, Alberta, Edmonton, University of Alberta, E.H. Strickland Entomological Museum" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" lsid="urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="45" pageNumber="46">UASM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113655318FAF0FEA8FB69FE0C" ID-GBIF-Occurrence="2620303359" collectingDate="1999-11-06" collectionCode="CMNC" collectorName="R. Anderson" country="Ecuador" elevation="3400" latitude="-0.3413889" location="Lago Papallacta" longLatPrecision="21" longitude="-78.17305" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<specimenCount id="9D1808F513655318FAF0FEA8FAABFEEF" box="[1281,1325,336,364]" pageId="45" pageNumber="46" type="male">1 ♂</specimenCount>
;
<location id="8EC195A713655318FACBFEA8FEBFFE0C" LSID="urn:lsid:plazi:treatment:03B7726A1364531BFDA2FA6EFAD7FCCB:8EC195A713655318FACBFEA8FEBFFE0C" country="Ecuador" latitude="-0.3413889" longLatPrecision="21" longitude="-78.17305" name="Lago Papallacta" pageId="45" pageNumber="46" stateProvince="Napo">Lago Papallacta</location>
;
<geoCoordinate id="EE2AA5BB13655318FEB4FE89FE40FE0C" box="[325,454,371,399]" degrees="0" direction="south" minutes="20" orientation="latitude" pageId="45" pageNumber="46" precision="15" seconds="29" value="-0.3413889">0°20′29″ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB13655318FE22FE89FDE8FE0C" box="[467,622,371,399]" degrees="78" direction="west" minutes="10" orientation="longitude" pageId="45" pageNumber="46" precision="15" seconds="23" value="-78.17305">78°10′23″ W</geoCoordinate>
;
<elevation id="0033244F13655318FD8BFE8FFC95FE0C" box="[634,787,373,399]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="45" pageNumber="46" unit="m" value="3400.0">
<quantity id="4CE66E9913655318FD8BFE8FFD55FE0C" box="[634,723,373,399]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="45" pageNumber="46" unit="m" value="3400.0">3400 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13655318FCD1FE8FFC32FE0C" box="[800,948,373,399]" pageId="45" pageNumber="46" value="1999-11-06">
<collectingDate id="EFE41C5413655318FCD1FE8FFC32FE0C" box="[800,948,373,399]" pageId="45" pageNumber="46" value="1999-11-06">6 Nov. 1999</collectingDate>
</date>
;
<collectorName id="26EBA6AA13655318FC31FE8FFBD5FE0C" box="[960,1107,373,399]" pageId="45" pageNumber="46">R. Anderson</collectorName>
leg.;
<collectionCode id="ED0F5BB913655318FB64FE8FFB69FE0C" box="[1173,1263,373,399]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="45" pageNumber="46">CMNC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C13655318FF4CFE4CFE52FE53" blockId="45.[189,468,438,464]" box="[189,468,438,464]" pageId="45" pageNumber="46">
<heading id="D0E9741013655318FF4CFE4CFE52FE53" bold="true" box="[189,468,438,464]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B96A1F6E13655318FF4CFE4CFE52FE53" bold="true" box="[189,468,438,464]" pageId="45" pageNumber="46">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1365531BFF4CFE1FFD88FECB" blockId="45.[189,1399,485,758]" lastBlockId="46.[189,1399,267,329]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
Habitus:
<figureCitation id="1325DFF913655318FEDCFE1FFE0EFD83" box="[301,392,485,512]" captionStart="Figs 52–53" captionStartId="45.[189,241,1955,1981]" captionTargetBox="[351,1238,831,1892]" captionTargetId="figure@45.[334,1253,818,1914]" captionTargetPageId="45" captionText="Figs 52–53. Dyscolus silvestris Moret sp. nov., male holotype (QCAZ). 52. Habitus. 53. Male aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829714" httpUri="https://zenodo.org/record/3829714/files/figure.png" pageId="45" pageNumber="46">Fig. 52</figureCitation>
. Wingless. Body length: 9.1–10.0 mm. Body colour variable, predominantly dark brown with reddish brown areas in some specimens; legs, antennae and mouthparts reddish brown. Elytral microsculpture isodiametric. Head broad and convex, neck slightly constricted, eyes moderately bulging. Pronotum subcordiform, the base much narrower than the apex; sides arcuate anterad, markedly sinuate posterad; hind angles obtuse, sharp; two pairs of lateral setae. Elytra subovate, moderately convex; striae entire, well impressed; intervals flat. Third elytral interval with 3–4 discal setae. Legs: protarsi of the male much broader than those of the female. Fifth metatarsomere asetose ventrally. Last visible abdominal ventrite with one pair (♂) or two pairs (
<specimenCount id="9D1808F513655318FC74FD21FC1EFD76" box="[901,920,731,757]" pageId="45" pageNumber="46" type="female">♀</specimenCount>
) of setae along its apical margin.
<emphasis id="B96A1F6E1365531BFACBFD21FEA3FEA6" italics="true" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">Male genitalia</emphasis>
:
<figureCitation id="1325DFF91366531BFEC6FEF1FE16FEA5" box="[311,400,267,294]" captionStart="Figs 52–53" captionStartId="45.[189,241,1955,1981]" captionTargetBox="[351,1238,831,1892]" captionTargetId="figure@45.[334,1253,818,1914]" captionTargetPageId="45" captionText="Figs 52–53. Dyscolus silvestris Moret sp. nov., male holotype (QCAZ). 52. Habitus. 53. Male aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829714" httpUri="https://zenodo.org/record/3829714/files/figure.png" pageId="46" pageNumber="47">Fig. 53</figureCitation>
. Median lobe arcuate, shortly and bluntly triangular at apex, endophallus unarmed.
<emphasis id="B96A1F6E1366531BFF4CFED5FE02FECA" box="[189,388,303,329]" italics="true" pageId="46" pageNumber="47">Female genitalia</emphasis>
: unstudied.
</paragraph>
<caption id="DF6193F413655318FF4CF859FD84F863" ID-DOI="http://doi.org/10.5281/zenodo.3829714" ID-Zenodo-Dep="3829714" httpUri="https://zenodo.org/record/3829714/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,241,1955,1981]" targetBox="[351,1238,831,1892]" targetPageId="45">
<paragraph id="8BA1C37C13655318FF4CF859FD84F863" blockId="45.[189,1398,1955,2016]" pageId="45" pageNumber="46">
<emphasis id="B96A1F6E13655318FF4CF859FECEF83E" bold="true" box="[189,328,1955,1981]" pageId="45" pageNumber="46">Figs 52–53.</emphasis>
<taxonomicName id="4C1EB8FF13655318FEBEF859FDF1F83E" authority="Moret" authorityName="Moret" authorityYear="2020" box="[335,631,1955,1981]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="silvestris" status="sp. nov.">
<emphasis id="B96A1F6E13655318FEBEF859FDAFF83E" box="[335,553,1955,1981]" italics="true" pageId="45" pageNumber="46">Dyscolus silvestris</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513655318FD8CF859FD5FF83E" box="[637,729,1955,1981]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (QCAZ).
<emphasis id="B96A1F6E13655318FBF3F859FB99F83E" bold="true" box="[1026,1055,1955,1981]" pageId="45" pageNumber="46">52</emphasis>
. Habitus.
<emphasis id="B96A1F6E13655318FB67F859FB35F83E" bold="true" box="[1174,1203,1955,1981]" pageId="45" pageNumber="46">53</emphasis>
. Male aedeagus, median lobe in lateral view.
</paragraph>
</caption>
<paragraph id="8BA1C37C1366531BFF4CFE8AFEE5FE09" blockId="46.[189,355,368,394]" box="[189,355,368,394]" pageId="46" pageNumber="47">
<heading id="D0E974101366531BFF4CFE8AFEE5FE09" bold="true" box="[189,355,368,394]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96A1F6E1366531BFF4CFE8AFEE5FE09" bold="true" box="[189,355,368,394]" pageId="46" pageNumber="47">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CFE65FC6BFDC5" blockId="46.[189,1399,414,582]" pageId="46" pageNumber="47">
<taxonomicName id="4C1EB8FF1366531BFF4CFE65FE65FE3A" authority="Moret" authorityName="Moret" authorityYear="2020" box="[189,483,415,441]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="silvestris" status="sp. nov.">
<emphasis id="B96A1F6E1366531BFF4CFE65FE13FE3A" box="[189,405,415,441]" italics="true" pageId="46" pageNumber="47">Dyscolus silvestris</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151366531BFE16FE65FDC6FE3A" box="[487,576,415,441]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
is similar to the shorter and broader individuals of
<taxonomicName id="4C1EB8FF1366531BFB70FE65FAE9FE3A" baseAuthorityName="Bates" baseAuthorityYear="1891" box="[1153,1391,415,441]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="denigratus">
<emphasis id="B96A1F6E1366531BFB70FE65FAE9FE3A" box="[1153,1391,415,441]" italics="true" pageId="46" pageNumber="47">Dyscolus denigratus</emphasis>
</taxonomicName>
. The main differences are: head slightly bigger and broader; sides of the pronotum sinuate basally, hind angles sharply obtuse (completely rounded in
<taxonomicName id="4C1EB8FF1366531BFD01FE1CFC12FE7C" baseAuthorityName="Bates" baseAuthorityYear="1891" box="[752,916,485,511]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="denigratus">
<emphasis id="B96A1F6E1366531BFD01FE1CFC12FE7C" box="[752,916,485,511]" italics="true" pageId="46" pageNumber="47">D. denigratus</emphasis>
</taxonomicName>
); femora of the male not swollen as in
<taxonomicName id="4C1EB8FF1366531BFF4CFDF3FEDBFDA1" baseAuthorityName="Bates" baseAuthorityYear="1891" box="[189,349,520,546]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="denigratus">
<emphasis id="B96A1F6E1366531BFF4CFDF3FEDBFDA1" box="[189,349,520,546]" italics="true" pageId="46" pageNumber="47">D. denigratus</emphasis>
</taxonomicName>
. The apex of the median lobe of the male genitalia has a shorter apex, and the endophallus is entirely unarmed (with a heavily sclerotized area in
<taxonomicName id="4C1EB8FF1366531BFCCAFDD6FC5AFDC6" baseAuthorityName="Bates" baseAuthorityYear="1891" box="[827,988,555,581]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="denigratus">
<emphasis id="B96A1F6E1366531BFCCAFDD6FC5AFDC6" box="[827,988,555,581]" italics="true" pageId="46" pageNumber="47">D. denigratus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CFD97FE9BFD04" blockId="46.[189,285,621,647]" box="[189,285,621,647]" pageId="46" pageNumber="47">
<heading id="D0E974101366531BFF4CFD97FE9BFD04" bold="true" box="[189,285,621,647]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96A1F6E1366531BFF4CFD97FE9BFD04" bold="true" box="[189,285,621,647]" pageId="46" pageNumber="47">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CFD66FE30FD5B" blockId="46.[189,1399,667,728]" pageId="46" pageNumber="47">
Upper montane forest on the Amazonian slope of the
<taxonomicName id="4C1EB8FF1366531BFCB1FD61FC08FD36" authorityName="Stal" authorityYear="1866" box="[832,910,667,693]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="54" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
, from
<quantity id="4CE66E991366531BFC2FFD61FB1AFD35" box="[990,1180,667,694]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="46" pageNumber="47" unit="m" value="3050.0" valueMax="3400.0" valueMin="2700.0">2700 to 3400 m</quantity>
a.s.l. This species lives in the leaf litter.
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CFCFAFE6DFC99" blockId="46.[189,491,768,794]" box="[189,491,768,794]" pageId="46" pageNumber="47">
<heading id="D0E974101366531BFF4CFCFAFE6DFC99" bold="true" box="[189,491,768,794]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B96A1F6E1366531BFF4CFCFAFE6DFC99" bold="true" box="[189,491,768,794]" pageId="46" pageNumber="47">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C1366531BFF4CFCD4FAD7FCCB" blockId="46.[189,1361,814,841]" box="[189,1361,814,841]" pageId="46" pageNumber="47">
Only known from the
<typeStatus id="54A57DDE1366531BFE33FCD5FE73FCCA" box="[450,501,815,841]" pageId="46" pageNumber="47">type</typeStatus>
locality around Papallacta in Northern
<collectingCountry id="F30983EC1366531BFC32FCD4FBA1FCCB" box="[963,1063,814,840]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
, probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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