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<document id="A4867D7F22F2500F0F0EAA9940EDC6C3" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A1341533CFD99FBE9FD54F86A" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus Dejean 1831" docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="10" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="56D45D70F513980E022A6F94A490F224">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
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<mods:namePart id="18FF9562C1684472146CF9009D42AB2D">Moret, Pierre</mods:namePart>
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<mods:affiliation id="2B660626FE44EEC579F9514AE8EA5E90">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="4842FFB39D33A11C5FCC061FA2252BD1" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
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<mods:namePart id="651D694B0F00A0DDE224002BCDC0E03B">Murienne, Jérôme</mods:namePart>
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<mods:affiliation id="1CE3ED2B76842D0632634BB3C7DBDFF2">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
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<mods:title id="77155D6443513B30A5971BDD653C86C3">European Journal of Taxonomy</mods:title>
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<mods:date id="E7B9FE7DF8D1D7D8625AE3C62B3D87C3">2020</mods:date>
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<treatment id="03B7726A1341533CFD99FBE9FD54F86A" ID-DOI="http://doi.org/10.5281/zenodo.3848411" ID-GBIF-Taxon="164160390" ID-Zenodo-Dep="3848411" LSID="urn:lsid:plazi:treatment:03B7726A1341533CFD99FBE9FD54F86A" httpUri="http://treatment.plazi.org/id/03B7726A1341533CFD99FBE9FD54F86A" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C30490F71341533CFD99FBE9FC4AFBAE" box="[616,972,1042,1069]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BA1C37C1341533CFD99FBE9FC4AFBAE" blockId="9.[616,972,1042,1069]" box="[616,972,1042,1069]" pageId="9" pageNumber="10">
<heading id="D0E974101341533CFD99FBE9FC4AFBAE" box="[616,972,1042,1069]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
Genus
<taxonomicName id="4C1EB8FF1341533CFD4BFBE8FC4AFBAE" ID-CoL="8KWS5" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" box="[698,972,1042,1069]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E1341533CFD4BFBE8FCA1FBAE" bold="true" box="[698,807,1042,1069]" italics="true" pageId="9" pageNumber="10">Dyscolus</emphasis>
<bibRefCitation id="EF8FBE8D1341533CFCDEFBE8FC4AFBAE" author="Dejean P. F." box="[815,972,1042,1069]" pageId="9" pageNumber="10" refId="ref26023" refString="Dejean P. F. 1831. Species general des Coleopteres de la collection de M. le comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris. https: // doi. org / 10.5962 / bhl. title. 8863" type="book" year="1831">Dejean, 1831</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30490F71341533CFF4CFBAEFADDFAFD" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BA1C37C1341533CFF4CFBAEFDF9FB12" blockId="9.[189,1398,1108,1205]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB0EE4521341533CFF4CFBAEFDF9FB12" pageId="9" pageNumber="10">
<treatmentCitation id="0ABFE56D1341533CFF4CFBAEFDAEFBED" author="Dejean P. F." box="[189,552,1108,1134]" page="437" pageId="9" pageNumber="10" year="1831">
<taxonomicName id="4C1EB8FF1341533CFF4CFBAEFDAEFBED" ID-CoL="8KWS5" authority="Dejean, 1831: 437" authorityName="Dejean" authorityPageNumber="437" authorityYear="1831" box="[189,552,1108,1134]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E1341533CFF4CFBAEFEAFFBED" box="[189,297,1108,1134]" italics="true" pageId="9" pageNumber="10">Dyscolus</emphasis>
<bibRefCitation id="EF8FBE8D1341533CFEC8FBAEFDAEFBED" author="Dejean P. F." box="[313,552,1108,1134]" pageId="9" pageNumber="10" refId="ref26023" refString="Dejean P. F. 1831. Species general des Coleopteres de la collection de M. le comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris. https: // doi. org / 10.5962 / bhl. title. 8863" type="book" year="1831">Dejean, 1831: 437</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54A57DDE1341533CFDCDFBAFFDF0FBEC" box="[572,630,1109,1135]" pageId="9" pageNumber="10">Type</typeStatus>
species:
<taxonomicName id="4C1EB8FF1341533CFD04FBAEFB36FBED" ID-CoL="388KS" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" box="[757,1200,1108,1134]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="memnonius">
<emphasis id="B96A1F6E1341533CFD04FBAEFC71FBED" box="[757,1015,1108,1134]" italics="true" pageId="9" pageNumber="10">Dyscolus memnonius</emphasis>
<bibRefCitation id="EF8FBE8D1341533CFBF9FBAEFB36FBED" author="Dejean P. F." box="[1032,1200,1108,1134]" pageId="9" pageNumber="10" refId="ref26023" refString="Dejean P. F. 1831. Species general des Coleopteres de la collection de M. le comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris. https: // doi. org / 10.5962 / bhl. title. 8863" type="book" year="1831">Dejean, 1831</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<treatmentCitation id="0ABFE56D1341533CFE70FB8DFDF7FB12" author="Chaudoir M. de" box="[385,625,1143,1169]" page="343" pageId="9" pageNumber="10" year="1878">
<bibRefCitation id="EF8FBE8D1341533CFE70FB8DFDF7FB12" author="Chaudoir M. de" box="[385,625,1143,1169]" pageId="9" pageNumber="10" pagination="275 - 382" refId="ref25990" refString="Chaudoir M. de 1878. Revision des genres Onychopterygia, Dicranoncus et Colpodes. Annales de la Societe entomologique de France (5 e Serie) 8: 275 - 382." type="journal article" year="1878">Chaudoir 1878: 343</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA1C37C1341533CFF4CFB60FD3DFB36" blockId="9.[189,1398,1108,1205]" box="[189,699,1178,1205]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB0EE4521341533CFF4CFB60FD3DFB36" box="[189,699,1178,1205]" pageId="9" pageNumber="10">
<treatmentCitation id="0ABFE56D1341533CFF4CFB60FDC2FB36" author="Moret P." box="[189,580,1178,1205]" page="498" pageId="9" pageNumber="10" year="1996">
<taxonomicName id="4C1EB8FF1341533CFF4CFB60FDC2FB36" authority="Moret, 1996: 498" authorityName="Moret" authorityYear="1996" box="[189,580,1178,1205]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Hydrodyscolus">
<emphasis id="B96A1F6E1341533CFF4CFB60FEEAFB37" box="[189,364,1178,1204]" italics="true" pageId="9" pageNumber="10">Hydrodyscolus</emphasis>
<bibRefCitation id="EF8FBE8D1341533CFE82FB61FDC2FB36" author="Moret P." box="[371,580,1178,1205]" pageId="9" pageNumber="10" pagination="491 - 503" refId="ref27041" refString="Moret P. 1996. Mise au point taxinomique sur le genre Dyscolus Dejean et description de cinq especes nouvelles (Caraboidea, Harpalidae, Platynini). Bollettino del Museo regionale di Scienze naturali - Torino 14 (2): 491 - 503." type="journal article" year="1996">Moret, 1996: 498</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B96A1F6E1341533CFDBFFB61FD3DFB36" bold="true" box="[590,699,1179,1205]" pageId="9" pageNumber="10">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA1C37C1341533CFF4CFB26FD94FB75" blockId="9.[189,530,1244,1270]" box="[189,530,1244,1270]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB0EE4521341533CFF4CFB26FD94FB75" box="[189,530,1244,1270]" pageId="9" pageNumber="10">
<taxonomicName id="4C1EB8FF1341533CFF4CFB26FEAFFB75" ID-CoL="8KWS5" authorityName="Dejean" authorityYear="1831" box="[189,297,1244,1270]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B96A1F6E1341533CFF4CFB26FEAFFB75" box="[189,297,1244,1270]" italics="true" pageId="9" pageNumber="10">Dyscolus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABFE56D1341533CFEB7FB26FD88FB75" author="Moret P." box="[326,526,1244,1270]" page="143" pageId="9" pageNumber="10" year="1989">
<bibRefCitation id="EF8FBE8D1341533CFEB7FB26FD88FB75" author="Moret P." box="[326,526,1244,1270]" pageId="9" pageNumber="10" pagination="133 - 148" refId="ref26951" refString="Moret P. 1989. Demembrement du genre Colpodes auctorum. I: Individualisation et definition des genres neotropicaux Dyscolus Dejean et Stenocnemion gen. nov. (Col. Caraboidea Platyninae). Bulletin de la Societe entomologique de France 93 (5 - 6): 133 - 148." type="journal article" year="1989">Moret 1989: 143</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA1C37C1341533CFF4CFAE7FADDFAFD" blockId="9.[189,1399,1309,1406]" pageId="9" pageNumber="10">
The revision of the sequenced species led us to re-examine related material and to describe three new species of which we do not have DNA data, but which can be readily separated from the sequenced taxa (
<taxonomicName id="4C1EB8FF1341533CFF36FA9EFE60FAFD" ID-CoL="8MDGD" authority="Moret" authorityName="Moret" authorityYear="2020" box="[199,486,1380,1406]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
<emphasis id="B96A1F6E1341533CFF36FA9EFE11FAFD" box="[199,407,1380,1406]" italics="true" pageId="9" pageNumber="10">Dyscolus placitus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151341533CFE1CFA9EFDCCFAFD" box="[493,586,1380,1406]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C1EB8FF1341533CFDA7FA9FFC9CFAFD" ID-CoL="8MCD9" authority="Moret" authorityName="Moret" authorityYear="2020" box="[598,794,1380,1406]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rivinus" status="sp. nov.">
<emphasis id="B96A1F6E1341533CFDA7FA9FFD4DFAFD" box="[598,715,1381,1406]" italics="true" pageId="9" pageNumber="10">D. rivinus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151341533CFCD0FA9EFCFAFAFD" box="[801,892,1380,1406]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1EB8FF1341533CFC44FA9FFB6FFAFD" ID-CoL="8MLKZ" authority="Moret" authorityName="Moret" authorityYear="2020" box="[949,1257,1380,1406]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="verecundissimus" status="sp. nov.">
<emphasis id="B96A1F6E1341533CFC44FA9FFB1FFAFD" box="[949,1177,1380,1406]" italics="true" pageId="9" pageNumber="10">D. verecundissimus</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151341533CFB01FA9EFAC8FAFD" box="[1264,1358,1380,1406]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C30490F71341533CFF4CFA5FFD54F86A" pageId="9" pageNumber="10" type="description">
<paragraph id="8BA1C37C1341533CFF4CFA5FFD12F935" blockId="9.[189,1399,1445,1718]" pageId="9" pageNumber="10">
All the species under study share the following characters, which are, therefore, not repeatedly mentioned in the descriptions of the new species: labrum with six setae near apical margin; clypeus with two pairs of setae; two supraorbital setae each side; mentum with one pair of setae, submentum with two pairs of setae (except in the species of the
<taxonomicName id="4C1EB8FF1341533CFDB2F9F5FD24F9AA" authorityName="Moret" authorityYear="2005" box="[579,674,1551,1577]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B96A1F6E1341533CFDB2F9F5FD24F9AA" box="[579,674,1551,1577]" italics="true" pageId="9" pageNumber="10">hirsutus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1EB8FF1341533CFD29F9F5FCDAF9AA" authorityName="Moret" authorityYear="1998" box="[728,860,1551,1577]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B96A1F6E1341533CFD29F9F5FCDAF9AA" box="[728,860,1551,1577]" italics="true" pageId="9" pageNumber="10">verecundus</emphasis>
</taxonomicName>
groups, which may have additional submental setae); pronotum with two pairs of lateral setae, the distal one located near the widest point of pronotum, the basal one near or at the laterobasal angle (except
<taxonomicName id="4C1EB8FF1341533CFCC9F9ACFB8DF9F3" authority="Moret" authorityName="Moret" authorityYear="2020" box="[824,1035,1621,1648]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="donosoi" status="sp. nov.">
<emphasis id="B96A1F6E1341533CFCC9F9ACFC3CF9EC" box="[824,954,1621,1647]" italics="true" pageId="9" pageNumber="10">D. donosoi</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151341533CFBE5F9ACFBF6F9F3" box="[1044,1136,1622,1648]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
which lacks the basal seta); parascutellar setiferous pore present; metathoracic wings reduced and metepisternum very short (except in
<taxonomicName id="4C1EB8FF1341533CFEC8F966FDA7F935" authority="Moret" authorityName="Moret" authorityYear="2020" box="[313,545,1692,1718]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="eleonorae" status="sp. nov.">
<emphasis id="B96A1F6E1341533CFEC8F966FE54F935" box="[313,466,1692,1718]" italics="true" pageId="9" pageNumber="10">D. eleonorae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A2151341533CFDD9F966FD01F935" box="[552,647,1692,1718]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BA1C37C1341533CFF4CF927FE2BF82B" blockId="9.[189,1399,1757,1960]" pageId="9" pageNumber="10">The order in which these 25 new species are presented below is based on broad biogeographic and ecological groupings and on resemblances in external morphology, in order to make comparisons and identifications easier. The five resulting sections are: 1/ full-winged montane forest species (pp. 11–13); 2/ riparian wingless species of the montane forest (pp. 13–18); 3/ wingless, non-riparian species of the montane forest (pp. 18–36); 4/ riparian species of the páramo (pp. 36–42); 5/ non-riparian species of the páramo (pp. 42–51).</paragraph>
<paragraph id="8BA1C37C1341533CFF4CF834FD54F86A" blockId="9.[189,722,1998,2025]" box="[189,722,1998,2025]" pageId="9" pageNumber="10">All the new species are authored by P. Moret.</paragraph>
</subSubSection>
</treatment>
</document>
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