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<document id="60C6116EAA6759CB077E46D75DE792F2" ID-DOI="10.1093/zoolinnean/zlad043" ID-ISSN="0024-4082" ID-Zenodo-Dep="8426263" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696940821456" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan C. &amp; Cerdeño, Esperanza" docDate="2023" docId="03B17549E924476F9EF8FA7CFBDFFABA" docLanguage="en" docName="zlad043.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad043" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Protypotherium antiquum Ameghino 1885" docType="treatment" docVersion="2" lastPageNumber="428" masterDocId="FF880D31E92F47649E20FFB4FF9EFFC7" masterDocTitle="Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" masterLastPageNumber="444" masterPageNumber="417" pageNumber="428" updateTime="1696941306917" updateUser="ExternalLinkService">
<mods:mods id="0C006BECBDB6EF98727AC292F14E0C8E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4666E994195605BEC541674AC9B12839">
<mods:title id="FC0634D28AC32A1C6BFB68BD16754FE7">Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae</mods:title>
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<mods:namePart id="F0647CCC0303CB7568008D358BE71EF5">Fernández, Mercedes</mods:namePart>
<mods:affiliation id="65197A4830EA5A7CC115EC55EFACE568">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina &amp; Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina &amp; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
<mods:nameIdentifier id="B04B865FED5C1B59116FD0B9EDA7B5AB" type="email">mechisfernandezpaleo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A0DEFA3B942931FA2B85A4F84DAD1A9B">Fernicola, Juan C.</mods:namePart>
<mods:affiliation id="9E87B3030775AACA0566A267640D7E96">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales ‘ Bernardino Rivadavia’, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina &amp; Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina &amp; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="EB3B4B4F6A3D14657062D021F068E214">Cerdeño, Esperanza</mods:namePart>
<mods:affiliation id="FDE25021C37416C8BA51AC80A6E09EDB">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina &amp; Paleobiología y Paleoecología, Instituto de Nivología, Glaciología y Ciencias Ambientales, Centro Científico Tecnológico-CONICET-Mendoza, Avenida Ruiz Leal s / n, M 5500 Mendoza, Argentina</mods:affiliation>
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<mods:title id="D641A3167737BB668EB92B91FC575654">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="FEC7F940708177DB5B284D36C1AC9C34">2023</mods:date>
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<mods:number id="536477789498058CC060BC6D50D735A6">2023-07-11</mods:number>
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<mods:number id="D519E09C894C504EDBACE7B1F244B872">199</mods:number>
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<mods:number id="B5CED1D0017C5C142CA001367574AB82">2</mods:number>
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<mods:identifier id="9830882CBD94EA01F6FCF7D2B31E0680" type="DOI">10.1093/zoolinnean/zlad043</mods:identifier>
<mods:identifier id="26E5247DBD002E9024A3156A208A1B9C" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03B17549E924476F9EF8FA7CFBDFFABA" ID-DOI="http://doi.org/10.5281/zenodo.8426260" ID-Zenodo-Dep="8426260" LSID="urn:lsid:plazi:treatment:03B17549E924476F9EF8FA7CFBDFFABA" httpUri="http://treatment.plazi.org/id/03B17549E924476F9EF8FA7CFBDFFABA" lastPageNumber="428" pageId="11" pageNumber="428">
<subSubSection id="C30297D4E924476F9EF8FA7CFD0DFA25" box="[216,659,1480,1507]" pageId="11" pageNumber="428" type="nomenclature">
<paragraph id="8BA7C45FE924476F9EF8FA7CFD0DFA25" blockId="11.[216,659,1480,1507]" box="[216,659,1480,1507]" pageId="11" pageNumber="428">
<heading id="D0EF7333E924476F9EF8FA7CFD0DFA25" bold="true" box="[216,659,1480,1507]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="428" reason="2">
<taxonomicName id="4C18BFDCE924476F9EF8FA7CFD0DFA25" authority="Ameghino, 1885" authorityName="Ameghino" authorityYear="1885" box="[216,659,1480,1507]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9EF8FA7CFD0DFA25" bold="true" box="[216,659,1480,1507]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9EF8FA7CFE40FA25" bold="true" box="[216,478,1480,1507]" italics="true" pageId="11" pageNumber="428">Protypotherium antiquum</emphasis>
<bibRefCitation id="EF89B9AEE924476F9FC3FA7CFD0DFA25" author="Ameghino F" box="[483,659,1480,1506]" pageId="11" pageNumber="428" pagination="5 - 207" refId="ref21702" refString="Ameghino F. Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba 1885; 8: 5 - 207." type="journal article" year="1885">Ameghino, 1885</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E924476F9E51FA5BFBDFFABA" pageId="11" pageNumber="428" type="description">
<paragraph id="8BA7C45FE924476F9E51FA5BFDCCF9C0" blockId="11.[113,762,1519,1605]" box="[113,594,1519,1543]" pageId="11" pageNumber="428">
(
<figureCitation id="1323D8DAE924476F9E5CFA5BFF7CF9C0" box="[124,226,1519,1543]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">Fig. 4A–B</figureCitation>
; Supporting Information, Table S2)
</paragraph>
<paragraph id="8BA7C45FE924476F9EADF9BAFF68F982" blockId="11.[113,762,1519,1605]" pageId="11" pageNumber="428">
1885
<taxonomicName id="4C18BFDCE924476F9EE1F9BAFDABF9E1" authority="Ameghino" authorityName="Ameghino" authorityYear="1885" box="[193,565,1550,1574]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9EE1F9BAFE5FF9E1" box="[193,449,1550,1574]" italics="true" pageId="11" pageNumber="428">Protypotherium antiquum</emphasis>
Ameghino
</taxonomicName>
, p. 81–83 (original description).
</paragraph>
<paragraph id="8BA7C45FE924476F9E51F9D3FDC2F97A" blockId="11.[113,762,1639,1725]" pageId="11" pageNumber="428">
<materialsCitation id="3B70CE02E924476F9E51F9D3FDC2F97A" collectionCode="ZMK" collectorName="Ituzaingo Fm." country="Argentina" location="Entre Rios Province" pageId="11" pageNumber="428" specimenCode="ZMK 21" specimenCount="1" stateProvince="Entre Rios" typeStatus="lectotype">
<emphasis id="B96C184DE924476F9E51F9D3FF48F9B9" box="[113,214,1639,1662]" italics="true" pageId="11" pageNumber="428">
<typeStatus id="54A37AFDE924476F9E51F9D3FF4EF9B9" box="[113,208,1639,1662]" pageId="11" pageNumber="428">Lectotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBE6C24E924476F9EFDF9D3FEA8F9B8" box="[221,310,1639,1663]" collectionCode="ZMK" country="Germany" httpUri="http://grbio.org/cool/jqaq-6vk8" name="Zoologisches Museum der Universitaet Kiel" pageId="11" pageNumber="428">ZMK 21</specimenCode>
/1877 (
<figureCitation id="1323D8DAE924476F9FA8F9D3FE4CF9B8" box="[392,466,1639,1663]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">Fig. 4A</figureCitation>
), right mandibular fragment with p4–m3 (see
<bibRefCitation id="EF89B9AEE924476F9F0EF932FD8EF959" author="Fernandez M &amp; Fernicola JC &amp; Cerdeno E" box="[302,528,1670,1694]" pageId="11" pageNumber="428" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
Fernández
<emphasis id="B96C184DE924476F9F82F933FE4BF959" box="[418,469,1670,1694]" italics="true" pageId="11" pageNumber="428">et al.</emphasis>
2018
</bibRefCitation>
), from Late Miocene,
<collectorName id="26EDA189E924476F9E51F911FE9FF97A" box="[113,257,1701,1725]" pageId="11" pageNumber="428">Ituzaingó Fm.</collectorName>
,
<collectingRegion id="49DC0ABDE924476F9F29F912FE48F97A" box="[265,470,1701,1725]" country="Argentina" name="Entre Rios" pageId="11" pageNumber="428">Entre Ríos Province</collectingRegion>
(
<collectingCountry id="F30F84CFE924476F9FC2F911FDCEF97A" box="[482,592,1701,1725]" name="Argentina" pageId="11" pageNumber="428">Argentina</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA7C45FE924476F9E51F96BFE43F8F2" blockId="11.[113,762,1759,1845]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9E51F96BFEBCF930" box="[113,290,1759,1783]" italics="true" pageId="11" pageNumber="428">Referred material:</emphasis>
IBIGEO-P39 (
<figureCitation id="1323D8DAE924476F9FE9F96BFD8FF930" box="[457,529,1759,1783]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">Fig. 4B</figureCitation>
), less mandibular fragment with p4–m3; MCH-P 257, right mandibular fragment with p3–m1 (
<bibRefCitation id="EF89B9AEE924476F9EEBF8A9FE52F8F2" author="Armella MA &amp; Bonini T" box="[203,460,1821,1845]" pageId="11" pageNumber="428" pagination="102676" refId="ref22176" refString="Armella MA, Bonini T. Biostratigraphic significance of the presence of Protypotherium cf. P. antiquum Ameghino 1885 (Interatheriidae, Notoungulata) in the Late Miocene of northwestern Argentina. Journal of South American Earth Sciences 2020; 102: 102676. hups: // doi. org / 10.1016 / j. jsames. 2020.102676" type="journal article" year="2020">Armella and Bonini 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8BA7C45FE924476F9E51F8E3FC1DFE64" blockId="11.[113,765,1879,1965]" lastBlockId="11.[810,1459,144,419]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9E51F8E3FDD4F8A9" box="[113,586,1879,1903]" italics="true" pageId="11" pageNumber="428">
Extended diagnosis:
<taxonomicName id="4C18BFDCE924476F9F65F8E3FDD4F8A9" authorityName="Ameghino" authorityYear="1885" box="[325,586,1879,1903]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">Protypotherium antiquum</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C18BFDCE924476F9CFFF8E3FF23F849" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE924476F9CFFF8E3FF23F849" italics="true" pageId="11" pageNumber="428">Pr. australe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE924476F9EF0F8C3FEF2F849" authorityName="Ameghino" authorityYear="1887" box="[208,364,1910,1934]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE924476F9EF0F8C3FEF2F849" box="[208,364,1910,1934]" italics="true" pageId="11" pageNumber="428">Pr. praerutilum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE924476F9F5EF8C3FD95F849" authorityName="Ameghino" authorityYear="1902" box="[382,523,1910,1934]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE924476F9F5EF8C3FD95F849" box="[382,523,1910,1934]" italics="true" pageId="11" pageNumber="428">Pr. columnifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE924476F9C3DF8C3FD53F849" authorityName="Ameghino" authorityYear="1891" box="[541,717,1910,1934]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE924476F9C3DF8C3FD53F849" box="[541,717,1910,1934]" italics="true" pageId="11" pageNumber="428">Pr. compressidens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE924476F9CFFF8C3FF55F86A" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE924476F9CFFF8C3FF55F86A" italics="true" pageId="11" pageNumber="428">Pr. minutum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE924476F9EF5F822FED7F86A" authorityName="Ameghino" authorityYear="1889" box="[213,329,1941,1965]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE924476F9EF5F822FED7F86A" box="[213,329,1941,1965]" italics="true" pageId="11" pageNumber="428">Pr. claudum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C18BFDCE924476F9F5DF822FDBFF86A" box="[381,545,1941,1965]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE924476F9F5DF822FDBFF86A" box="[381,545,1941,1965]" italics="true" pageId="11" pageNumber="428">Pr. colloncurensis</emphasis>
</taxonomicName>
by having the talonid of m3 with a very deep longitudinal and transverse labial sulcus and a mesio-labial lobe distally straight; from
<taxonomicName id="4C18BFDCE924476F9B28FF04FA2DFF00" authorityName="Ameghino" authorityYear="1891" box="[1288,1459,175,199]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE924476F9B28FF04FA2DFF00" box="[1288,1459,175,199]" italics="true" pageId="11" pageNumber="428">Pr. compressidens</emphasis>
</taxonomicName>
by having relatively wider lower cheek teeth; from
<taxonomicName id="4C18BFDCE924476F9B03FF7BFA08FF20" authorityName="Ameghino" authorityYear="1889" box="[1315,1430,207,231]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE924476F9B03FF7BFA08FF20" box="[1315,1430,207,231]" italics="true" pageId="11" pageNumber="428">Pr. claudum</emphasis>
</taxonomicName>
by exhibiting a p4 with a transversely deeper, narrower, and more distally placed ectoflexid, and m1–3 with deeper entoflexid; from
<taxonomicName id="4C18BFDCE924476F9D45FE99FB93FE82" box="[869,1037,301,325]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE924476F9D45FE99FB93FE82" box="[869,1037,301,325]" italics="true" pageId="11" pageNumber="428">Pr. colloncurensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE924476F9A65FE99FB50FE82" authorityName="Ameghino" authorityYear="1902" box="[1093,1230,301,325]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE924476F9A65FE99FB50FE82" box="[1093,1230,301,325]" italics="true" pageId="11" pageNumber="428">Pr. columnifer</emphasis>
</taxonomicName>
by the sub-triangular talonid of p4; from
<taxonomicName id="4C18BFDCE924476F9DCDFEF9FB1EFEA3" authorityName="Ameghino" authorityYear="1887" box="[1005,1152,332,356]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE924476F9DCDFEF9FB1EFEA3" box="[1005,1152,332,356]" italics="true" pageId="11" pageNumber="428">Pr. praerutilum</emphasis>
</taxonomicName>
by having a slightly larger size; and from
<taxonomicName id="4C18BFDCE924476F9DB1FED8FB87FE44" authorityName="Ameghino" authorityYear="1902" box="[913,1049,363,387]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE924476F9DB1FED8FB87FE44" box="[913,1049,363,387]" italics="true" pageId="11" pageNumber="428">Pr. columnifer</emphasis>
</taxonomicName>
by the talonid of p4 narrower than the trigonid.
</paragraph>
<paragraph id="8BA7C45FE924476F9D0AFE70FC16FD9E" blockId="11.[810,1460,452,601]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9D0AFE70FB23FE1B" box="[810,1213,452,476]" italics="true" pageId="11" pageNumber="428">Geographic and temporal provenance:</emphasis>
<materialsCitation id="3B70CE02E924476F9AEEFE70FBDBFE3C" country="Argentina" location="Entre Rios Province" pageId="11" pageNumber="428" specimenCount="1" stateProvince="Entre Rios">
<collectingRegion id="49DC0ABDE924476F9AEEFE70FA2DFE1B" box="[1230,1459,452,476]" country="Argentina" name="Entre Rios" pageId="11" pageNumber="428">Entre Ríos Province</collectingRegion>
(
<collectingCountry id="F30F84CFE924476F9D11FE57FC01FE3C" box="[817,927,483,507]" name="Argentina" pageId="11" pageNumber="428">Argentina</collectingCountry>
), Ituzaingó Fm
</materialsCitation>
.;
<materialsCitation id="3B70CE02E924476F9A76FE57FBD0FDDD" collectorName="Palo Pintado Fm." country="Argentina" location="Salta Province" pageId="11" pageNumber="428" specimenCount="1" stateProvince="Salta">
<collectingRegion id="49DC0ABDE924476F9A76FE57FB73FE3C" box="[1110,1261,483,507]" country="Argentina" name="Salta" pageId="11" pageNumber="428">Salta Province</collectingRegion>
,
<collectorName id="26EDA189E924476F9ADBFE57FA2AFE3C" box="[1275,1460,483,507]" pageId="11" pageNumber="428">Palo Pintado Fm.</collectorName>
(
<bibRefCitation id="EF89B9AEE924476F9D15FDB6FB99FDDC" author="Zimicz N &amp; Payrola P &amp; Papa C" box="[821,1031,514,539]" pageId="11" pageNumber="428" pagination="31 - 44" refId="ref25896" refString="Zimicz N, Payrola P, del Papa C. New, Late Miocene mammalian assemblage from the Palo Pintado Formation (northwestern Argentina). Journal of South American Earth Sciences 2018; 81: 31 - 44. hups: // doi. org / 10.1016 / j. jsames. 2017.11.003" type="journal article" year="2018">
Zimicz
<emphasis id="B96C184DE924476F9DACFDB7FC5AFDDD" box="[908,964,514,538]" italics="true" pageId="11" pageNumber="428">et al.</emphasis>
2018
</bibRefCitation>
); and
</materialsCitation>
<materialsCitation id="3B70CE02E924476F9A7EFDB7FC1AFD9E" collectionCode="SALMA" collectorName="Chiquimil Fm. &amp; Late Miocene &amp; Huayquerian" country="Argentina" location="Catamarca Province" pageId="11" pageNumber="428" specimenCount="1" stateProvince="Catamarca">
<collectingRegion id="49DC0ABDE924476F9A7EFDB7FAA8FDDD" box="[1118,1334,514,539]" country="Argentina" name="Catamarca" pageId="11" pageNumber="428">Catamarca Province</collectingRegion>
,
<collectorName id="26EDA189E924476F9B6AFDB6FCCAFDFE" pageId="11" pageNumber="428">Chiquimil Fm.</collectorName>
(
<bibRefCitation id="EF89B9AEE924476F9D46FD96FBEDFDFD" author="Armella MA &amp; Bonini T" box="[870,1139,546,570]" pageId="11" pageNumber="428" pagination="102676" refId="ref22176" refString="Armella MA, Bonini T. Biostratigraphic significance of the presence of Protypotherium cf. P. antiquum Ameghino 1885 (Interatheriidae, Notoungulata) in the Late Miocene of northwestern Argentina. Journal of South American Earth Sciences 2020; 102: 102676. hups: // doi. org / 10.1016 / j. jsames. 2020.102676" type="journal article" year="2020">Armella and Bonini 2020</bibRefCitation>
);
<collectorName id="26EDA189E924476F9AAEFD96FABFFDFD" box="[1166,1313,546,570]" pageId="11" pageNumber="428">Late Miocene</collectorName>
,
<collectorName id="26EDA189E924476F9B0DFD96FA2CFDFD" box="[1325,1458,546,570]" pageId="11" pageNumber="428">Huayquerian</collectorName>
<collectionCode id="ED095C9AE924476F9D0AFDF5FC1AFD9E" box="[810,900,577,601]" pageId="11" pageNumber="428">SALMA</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA7C45FE924476F9D0AFDCFFABEFD55" blockId="11.[810,1312,634,659]" box="[810,1312,634,659]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9D0AFDCFFC22FD55" box="[810,956,635,658]" italics="true" pageId="11" pageNumber="428">Measurements:</emphasis>
Supporting Information, Table S2.
</paragraph>
<paragraph id="8BA7C45FE924476F9D0AFD00FA15FB38" blockId="11.[810,1459,692,1405]" pageId="11" pageNumber="428">
<emphasis id="B96C184DE924476F9D0AFD00FC16FD0B" box="[810,904,692,716]" italics="true" pageId="11" pageNumber="428">Remarks:</emphasis>
<bibRefCitation id="EF89B9AEE924476F9DAEFD00FBC4FD0B" author="Zimicz N &amp; Payrola P &amp; Papa C" box="[910,1114,692,716]" pageId="11" pageNumber="428" pagination="31 - 44" refId="ref25896" refString="Zimicz N, Payrola P, del Papa C. New, Late Miocene mammalian assemblage from the Palo Pintado Formation (northwestern Argentina). Journal of South American Earth Sciences 2018; 81: 31 - 44. hups: // doi. org / 10.1016 / j. jsames. 2017.11.003" type="journal article" year="2018">
Zimicz
<emphasis id="B96C184DE924476F9DFBFD01FB95FD0B" box="[987,1035,692,716]" italics="true" pageId="11" pageNumber="428">et al.</emphasis>
(2018)
</bibRefCitation>
identified IBIGEO-P39 (
<figureCitation id="1323D8DAE924476F9B41FD00FA36FD0B" box="[1377,1448,692,716]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">Fig. 4B</figureCitation>
) as
<taxonomicName id="4C18BFDCE924476F9D62FD60FC24FD2C" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[834,954,724,747]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE924476F9D62FD60FC24FD2C" box="[834,954,724,747]" italics="true" pageId="11" pageNumber="428">Pr.minutum</emphasis>
</taxonomicName>
mainly due to size and a trilobed m3. Nevertheless, these authors mentioned that it resembled the remaining species of the genus. Concerning these features, the presence of a trilobed m3 is common in
<taxonomicName id="4C18BFDCE924476F9A1AFC85FB4AFC8E" authorityName="Ameghino" authorityYear="1885" box="[1082,1236,817,841]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="genus">
<emphasis id="B96C184DE924476F9A1AFC85FB4AFC8E" box="[1082,1236,817,841]" italics="true" pageId="11" pageNumber="428">Protypotherium</emphasis>
</taxonomicName>
and it is not exclusive of
<taxonomicName id="4C18BFDCE924476F9D68FCE5FC58FCAF" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[840,966,849,872]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE924476F9D68FCE5FC58FCAF" box="[840,966,849,872]" italics="true" pageId="11" pageNumber="428">Pr. minutum</emphasis>
</taxonomicName>
(
<typeStatus id="54A37AFDE924476F9DF7FCE4FBA6FCAF" box="[983,1080,848,872]" pageId="11" pageNumber="428">holotype</typeStatus>
MLP 12-2176) (see above). When comparing the m3 of MLP 12-2176 and IBIGEO-P39, the labial groove of the former is transversally shallow, but persistent, and in both the distal face of the mesio-labial lobe and the disto-labial lobe of the talonid are rounded. Instead, this groove is transversally deeper and persistent in IBIGEO-P39 (
<figureCitation id="1323D8DAE924476F9B7DFC59FA36FBC2" box="[1373,1448,1005,1029]" captionStart="Figure 4" captionStartId="7.[113,178,1701,1725]" captionTargetBox="[147,1428,146,1672]" captionTargetId="figure-66@7.[145,1428,144,1673]" captionTargetPageId="7" captionText="Figure 4. Protypotherium antiquum, in occlusal view: (A) ZMK 21/1877, right mandibular fragment with p4–m3; (B) IBIGEO-P 39, less mandibular fragment with p4–m3. Protypotherium australe, in occlusal view: (C) MACN-A 1338 (cast), less maxillary fragment with P3 (broken)–M3. Protypotherium praerutilum, in occlusal view:(D) MACN-A 1081, incomplete skull; (E) MACN-A 3920, complete skull; (F) AMNH 9565, incomplete skull.Protypotherium australe, in occlusal view: (G) MACN-A 9644, incomplete skull. ’ indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426274" httpUri="https://zenodo.org/record/8426274/files/figure.png" pageId="11" pageNumber="428">Fig. 4B</figureCitation>
) and the distal face of the mesio-labial lobe is straight, which is coincident with the diagnostic features of
<taxonomicName id="4C18BFDCE924476F9AFCFB99FAC3FB84" authorityName="Ameghino" authorityYear="1885" box="[1244,1373,1068,1092]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9AFCFB99FAC3FB84" box="[1244,1373,1068,1092]" italics="true" pageId="11" pageNumber="428">Pr. antiquum</emphasis>
</taxonomicName>
. In addition, IBIGEO-P39 (e.g. Lm1 =
<quantity id="4CE069BAE924476F9A5BFBFFFB42FBA4" box="[1147,1244,1099,1123]" metricMagnitude="-3" metricUnit="m" metricValue="6.26" pageId="11" pageNumber="428" unit="mm" value="6.26">6.26 mm</quantity>
; Wm1 ≈
<quantity id="4CE069BAE924476F9B60FBFFFA0DFBA4" box="[1344,1427,1099,1123]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="428" unit="mm" value="3.2">3.2 mm</quantity>
) is intermediate in size between
<taxonomicName id="4C18BFDCE924476F9A78FBDFFB4AFB45" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[1112,1236,1131,1154]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE924476F9A78FBDFFB4AFB45" box="[1112,1236,1131,1154]" italics="true" pageId="11" pageNumber="428">Pr. minutum</emphasis>
</taxonomicName>
(e.g. Lm1 ≈
<quantity id="4CE069BAE924476F9B7CFBDEFA30FB45" box="[1372,1454,1130,1154]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="428" unit="mm" value="4.6">4.6 mm</quantity>
; Wm1 ≈
<quantity id="4CE069BAE924476F9DA1FB3EFC76FB65" box="[897,1000,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="428" unit="mm" value="2.6">2.60 mm</quantity>
) and
<taxonomicName id="4C18BFDCE924476F9A14FB3EFB23FB66" authorityName="Ameghino" authorityYear="1885" box="[1076,1213,1162,1185]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9A14FB3EFB23FB66" box="[1076,1213,1162,1185]" italics="true" pageId="11" pageNumber="428">Pr. antiquum</emphasis>
</taxonomicName>
(e.g. Lm1 =
<quantity id="4CE069BAE924476F9B74FB3EFA30FB66" box="[1364,1454,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="428" unit="mm" value="7.4">7.4 mm</quantity>
; Wm1 ≈
<quantity id="4CE069BAE924476F9DA1FB1DFC4EFB06" box="[897,976,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="428" unit="mm" value="4.1">4.1 mm</quantity>
(see Supporting Information, Table S2), being even closer to the laưer. Therefore, mainly based on the morphological features, we re-identify IBIGEO-P39 as
<taxonomicName id="4C18BFDCE924476F9B27FB5CFA18FB38" authorityName="Ameghino" authorityYear="1885" box="[1287,1414,1256,1279]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9B27FB5CFA18FB38" box="[1287,1414,1256,1279]" italics="true" pageId="11" pageNumber="428">Pr. antiquum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA7C45FE924476F9D65FAB3FBDFFABA" blockId="11.[810,1459,692,1405]" pageId="11" pageNumber="428">
On the other hand, we state that MCH-P 257, mentioned as
<taxonomicName id="4C18BFDCE924476F9D0AFA92FC5AFAF9" authorityName="Ameghino" authorityYear="1885" box="[810,964,1318,1342]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="genus">
<emphasis id="B96C184DE924476F9D0AFA92FC5AFAF9" box="[810,964,1318,1342]" italics="true" pageId="11" pageNumber="428">Protypotherium</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C18BFDCE924476F9DCAFA93FBF4FAF9" authorityName="Ameghino" authorityYear="1885" box="[1002,1130,1319,1342]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9DCAFA93FBF4FAF9" box="[1002,1130,1319,1342]" italics="true" pageId="11" pageNumber="428">Pr. antiquum</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF89B9AEE924476F9AB1FA92FA30FAF9" author="Armella MA &amp; Bonini T" box="[1169,1454,1318,1342]" pageId="11" pageNumber="428" pagination="102676" refId="ref22176" refString="Armella MA, Bonini T. Biostratigraphic significance of the presence of Protypotherium cf. P. antiquum Ameghino 1885 (Interatheriidae, Notoungulata) in the Late Miocene of northwestern Argentina. Journal of South American Earth Sciences 2020; 102: 102676. hups: // doi. org / 10.1016 / j. jsames. 2020.102676" type="journal article" year="2020">Armella and Bonini (2020)</bibRefCitation>
, also corresponds to
<taxonomicName id="4C18BFDCE924476F9DDDFAF2FBE0FA9A" authorityName="Ameghino" authorityYear="1885" box="[1021,1150,1350,1373]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="11" pageNumber="428" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE924476F9DDDFAF2FBE0FA9A" box="[1021,1150,1350,1373]" italics="true" pageId="11" pageNumber="428">Pr. antiquum</emphasis>
</taxonomicName>
, sharing the same mentioned features with IBIGEO-P39.
</paragraph>
</subSubSection>
</treatment>
</document>
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