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<document id="F7FDE55D34C21B6EB2B402A00E1F8FD1" ID-DOI="10.5281/zenodo.10396362" ID-ISSN="1942-1354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707168658757" checkinUser="felipe" docAuthor="Zhang, Jing, Cong, Qian &amp; Grishin, Nick V." docDate="2023" docId="03810139FFCCBB44C0CAFA69E297B5E5" docLanguage="en" docName="InsectaMundi.2023.1026.1-115.pdf" docOrigin="Insecta Mundi 2023 (26)" docSource="http://dx.doi.org/10.5281/zenodo.10396362" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Pythonides latemarginatus Zhang &amp; Cong &amp; Grishin 2023, new species" docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="FFB87941FFDFBB50C00AFF56E355B262" masterDocTitle="Supplementary Materials and Appendix" masterLastPageNumber="115" masterPageNumber="1" pageNumber="18" updateTime="1707169288913" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="4E21D5EDFD14A91021FA299CC5454523">Supplementary Materials and Appendix</mods:title>
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<mods:name id="1EF40D8B24A0CD8536DEE0C3B53A1151" type="personal">
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<mods:namePart id="23C4D69D871A8424BDB5FF09882984CA">Zhang, Jing</mods:namePart>
<mods:affiliation id="E6D7ABD9706818AB6103C86FB5BBEDC1">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:namePart id="BB1853BE56399E352DEDAB3E10E4A8C9">Cong, Qian</mods:namePart>
<mods:affiliation id="3BEB3F120400224C332043997F1FD012">McDermott Center for Human Growth and Development and Department of Biophysics University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 8816 USA</mods:affiliation>
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<mods:namePart id="1877F8897914A9F95C443556166D328E">Grishin, Nick V.</mods:namePart>
<mods:affiliation id="AAB55E4D60E4996B276C555049B1B20D">Departments of Biophysics and Biochemistry University of Texas Southwestern Medical Center 5323 Harry Hines Blvd., Dallas, TX, 75390 - 9050 USA</mods:affiliation>
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<mods:title id="DCBF9A502BF2437935B0EEF078BE4B7C">Insecta Mundi</mods:title>
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<mods:date id="5FC6C58883EF1F99BF1BA4E7BEB1610F">2023</mods:date>
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<treatment id="03810139FFCCBB44C0CAFA69E297B5E5" LSID="urn:lsid:plazi:treatment:03810139FFCCBB44C0CAFA69E297B5E5" httpUri="http://treatment.plazi.org/id/03810139FFCCBB44C0CAFA69E297B5E5" lastPageId="20" lastPageNumber="19" pageId="19" pageNumber="18">
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<paragraph id="8B97B02FFFCCBB43C0CAFA69E04AB73E" blockId="19.[192,1442,1343,1924]" box="[192,799,1343,1372]" pageId="19" pageNumber="18">
<heading id="D0DF0743FFCCBB43C0CAFA69E04AB73E" box="[192,799,1343,1372]" fontSize="12" level="2" pageId="19" pageNumber="18" reason="2">
<emphasis id="B95C6C3DFFCCBB43C0CAFA69E04AB73E" bold="true" box="[192,799,1343,1372]" pageId="19" pageNumber="18">
<taxonomicName id="4C28CBACFFCCBB43C0CAFA69E12AB73E" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[192,639,1343,1372]" class="Insecta" family="Hesperiidae" genus="Pythonides" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="latemarginatus" status="sp. nov.">
<emphasis id="B95C6C3DFFCCBB43C0CAFA69E146B73E" bold="true" box="[192,531,1343,1372]" italics="true" pageId="19" pageNumber="18">Pythonides latemarginatus</emphasis>
Grishin
</taxonomicName>
,
<taxonomicNameLabel id="A26FD146FFCCBB43C280FA16E04AB73E" box="[650,799,1343,1372]" pageId="19" pageNumber="18" rank="species">new species</taxonomicNameLabel>
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https://zoobank.org/
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</paragraph>
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<subSubSection id="C332E3A4FFCCBB43C0CAFAD3E156B7C2" box="[192,515,1413,1440]" pageId="19" pageNumber="18" type="description">
<paragraph id="8B97B02FFFCCBB43C0CAFAD3E156B7C2" blockId="19.[192,1442,1343,1924]" box="[192,515,1413,1440]" pageId="19" pageNumber="18">
(
<figureCitation id="1313ACAAFFCCBB43C0C2FAD3E252B7FD" box="[200,263,1413,1439]" captionStart="Figure 2" captionStartId="20.[192,261,1419,1443]" captionTargetBox="[193,1440,268,1379]" captionTargetId="figure-157@20.[192,1440,266,1382]" captionTargetPageId="20" captionText="Figure 2. Phylogenetic trees of Achlyodini inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="19" pageNumber="18">Fig. 2</figureCitation>
part, 3738, 250251)
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFCCBB43C0CAFAFAE6F5B5E6" pageId="19" pageNumber="18" type="diagnosis">
<paragraph id="8B97B02FFFCCBB43C0CAFAFAE290B494" blockId="19.[192,1442,1343,1924]" pageId="19" pageNumber="18">
<emphasis id="B95C6C3DFFCCBB43C0CAFAFAE2BDB7A4" bold="true" box="[192,488,1452,1478]" pageId="19" pageNumber="18">Definition and diagnosis.</emphasis>
Specimens from
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identified as
<taxonomicName id="4C28CBACFFCCBB43C392FAFAE6F5B7A4" authority="Staudinger, 1876" authorityName="Staudinger" authorityYear="1876" box="[920,1440,1452,1478]" class="Insecta" family="Hesperiidae" genus="Pythonides" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="jovianus" subSpecies="amaryllis">
<emphasis id="B95C6C3DFFCCBB43C392FAFAE7B5B7A4" box="[920,1248,1452,1478]" italics="true" pageId="19" pageNumber="18">Pythonides jovianus amaryllis</emphasis>
Staudinger, 1876
</taxonomicName>
(
<typeStatus id="54930E8DFFCCBB43C0C2FA98E3AFB78A" box="[200,250,1486,1512]" pageId="19" pageNumber="18">type</typeStatus>
locality in
<collectingCountry id="F33FF0BFFFCCBB43C17DFA98E284B78A" box="[375,465,1486,1512]" name="Panama" pageId="19" pageNumber="18">Panama</collectingCountry>
and
<collectingCountry id="F33FF0BFFFCCBB43C20DFA98E12DB78A" box="[519,632,1486,1512]" name="Colombia" pageId="19" pageNumber="18">Colombia</collectingCountry>
) are sister to all
<taxonomicName id="4C28CBACFFCCBB43C325FA98E7CFB78A" authority="(Stoll, 1782)" baseAuthorityName="Stoll" baseAuthorityYear="1782" box="[815,1178,1486,1512]" class="Insecta" family="Hesperiidae" genus="Pythonides" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="jovianus">
<emphasis id="B95C6C3DFFCCBB43C325FA98E75EB785" box="[815,1035,1486,1512]" italics="true" pageId="19" pageNumber="18">Pythonides jovianus</emphasis>
(Stoll, 1782)
</taxonomicName>
and are genetically differentiated from it (
<figureCitation id="1313ACAAFFCCBB43C190FAB9E289B46B" box="[410,476,1519,1545]" captionStart="Figure 2" captionStartId="20.[192,261,1419,1443]" captionTargetBox="[193,1440,268,1379]" captionTargetId="figure-157@20.[192,1440,266,1382]" captionTargetPageId="20" captionText="Figure 2. Phylogenetic trees of Achlyodini inferred from protein-coding regions in a) the Z chromosome and b) the mitochondrial genome (in segments). See Fig. 1 legend for other notations." pageId="19" pageNumber="18">Fig. 2</figureCitation>
): e.g., their COI barcodes differ by 3.5% (24 bp) from
<emphasis id="B95C6C3DFFCCBB43C43EFAA6E792B46B" box="[1076,1223,1519,1545]" italics="true" pageId="19" pageNumber="18">P. j. amaryllis</emphasis>
, and therefore they represent a new species. This new species keys to
<emphasis id="B95C6C3DFFCCBB43C2F2F944E0DAB449" box="[760,911,1553,1579]" italics="true" pageId="19" pageNumber="18">P. j. amaryllis</emphasis>
(E.41.3(a)) in
<bibRefCitation id="EFB9CDDEFFCCBB43C43FF944E79DB44E" author="Evans WH" box="[1077,1224,1553,1580]" pageId="19" pageNumber="18" pagination="26 - 53" refId="ref77110" refString="Evans WH. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. The Trustees of the British Museum (Natural History); London. v + 246 p., pl. 26 - 53." type="book chapter" year="1953">Evans (1953)</bibRefCitation>
and differs from it by a broader brown marginal area on the hindwing and correspondingly smaller blue patch, but more extensive forewing postdiscal blue spotting, in particular having a blue streak by the inner margin and two prominent blue streaks near forewing tornus beneath (
<figureCitation id="1313ACAAFFCCBB43C263F920E1B6B4F2" box="[617,739,1654,1680]" captionStart="Figures 3348" captionStartId="96.[192,271,1841,1865]" captionTargetBox="[198,1434,372,1745]" captionTargetId="figure-67@96.[192,1440,361,1751]" captionTargetPageId="96" captionText="Figures 3348. Holotypes, data in text. 3334) Ridens angulinea sp. n. 3536) Pythonides lera sp. n. 3738) Pythonides latemarginatus sp. n. 3940) Gindanes variegatus sp. n. 4142) Milanion (Milanion) virga sp. n. 4344) Milanion (Milanion) furvus sp. n. 4546) Milanion (Milanion) laricus sp. n. 4748) Charidia ronda sp. n." pageId="19" pageNumber="18">Fig. 3738</figureCitation>
). This species is not cryptic and is diagnosed reliably by phenotype. In DNA, a combination of the following base pairs is diagnostic in the nuclear genome: aly
<date id="FF9696EFFFCCBB43C4E7F9CEE609B4D0" box="[1261,1372,1688,1714]" pageId="19" pageNumber="18" value="2178-10-01">2178.10.1</date>
:C78T, aly
<date id="FF9696EFFFCCBB43C0EBF9ECE205B4B6" box="[225,336,1722,1748]" pageId="19" pageNumber="18" value="2178-10-01">2178.10.1</date>
:C96T, aly363.24.5:G81A, aly522.12.2:G39A, aly9588.7.2:A27G, and COI barcode: T49A, T268C, T385C, T457C, T499C.
</paragraph>
<paragraph id="8B97B02FFFCCBB43C0CAF855E6F5B5E6" blockId="19.[192,1442,1343,1924]" pageId="19" pageNumber="18">
<emphasis id="B95C6C3DFFCCBB43C0CAF855E11BB57F" bold="true" box="[192,590,1794,1821]" pageId="19" pageNumber="18">
Barcode sequence of the
<typeStatus id="54930E8DFFCCBB43C1EAF855E11CB57F" box="[480,585,1795,1821]" pageId="19" pageNumber="18" type="holotype">holotype</typeStatus>
.
</emphasis>
Sample NVG-18018B04, GenBank OR837638, 658 base pairs: AACTTTATATTTTATTTTTGGAATTTGAGCTGGAATAGTAGGAACATCATTAAGTTTATTAATTCGAACTGAATTAGGTAATCCCGGTTCATTAATT GGAGATGATCAAATTTATAATACTATTGTAACTGCTCATGCTTTTATTATAATTTTCTTTATAGTAATACCAATTATAATTGGAGGATTTGGAAATT GACTAGTACCCCTTATACTAGGAGCTCCTGATATAGCATTCCCTCGAATAAATAATATAAGTTTTTGATTACTCCCCCCTTCTCTTACATTATTAAT TTCAAGAAGTATTGTTGAAAATGGAACTGGAACAGGATGAACAGTATACCCCCCTCTTTCCTCTAATATTGCTCATCAAGGATCTTCTGTTGACTTA
</paragraph>
</subSubSection>
<caption id="DF57E0A7FFCBBB44C0CAFADDE0D3B7A0" pageId="20" pageNumber="19" startId="20.[192,261,1419,1443]" targetBox="[193,1440,268,1379]" targetPageId="20" targetType="figure">
<paragraph id="8B97B02FFFCBBB44C0CAFADDE0D3B7A0" blockId="20.[192,1440,1419,1474]" pageId="20" pageNumber="19">
<emphasis id="B95C6C3DFFCBBB44C0CAFADDE24BB7C1" bold="true" box="[192,286,1419,1443]" pageId="20" pageNumber="19">Figure 2.</emphasis>
Phylogenetic trees of
<taxonomicName id="4C28CBACFFCBBB44C20AFADDE124B7C1" authorityName="Burmeister" authorityYear="1878" box="[512,625,1419,1443]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Achlyodini">Achlyodini</taxonomicName>
inferred from protein-coding regions in
<emphasis id="B95C6C3DFFCBBB44C41EFADDE77DB7C1" bold="true" box="[1044,1064,1419,1443]" pageId="20" pageNumber="19">a)</emphasis>
the Z chromosome and
<emphasis id="B95C6C3DFFCBBB44C52EFADDE66EB7C1" bold="true" box="[1316,1339,1419,1443]" pageId="20" pageNumber="19">b)</emphasis>
the mitochondrial genome (in segments). See Fig. 1 legend for other notations.
</paragraph>
</caption>
<subSubSection id="C332E3A4FFCBBB44C0CAFAACE7D8B45F" box="[192,1440,1530,1597]" pageId="20" pageNumber="19" type="description">
<paragraph id="8B97B02FFFCBBB44C0CAFAACE7D8B45F" blockId="20.[192,1440,1530,1597]" box="[192,1440,1530,1597]" pageId="20" pageNumber="19">GCTATTTTTTCCTTACACCTAGCTGGTATTTCATCAATTTTAGGAGCTATTAATTTTATTACTACAATCATTAATATACGAATTAGAAATTTATCAT TTGATCAAATACCCTTATTTGTTTGAGCTGTAGGAATTACAGCATTATTATTATTATTATCATTACCTGTTTTAGCCGGAGCTATTACCATATTATT AACTGACCGAAATTTAAATACATCATTTTTTGATCCAGCTGGTGGAGGAGATCCAATTTTATATCAACATTTATTT</paragraph>
</subSubSection>
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<paragraph id="8B97B02FFFCBBB44C0CAF919E6F5B40B" blockId="20.[191,1441,1619,1887]" box="[192,1440,1615,1645]" pageId="20" pageNumber="19">
<emphasis id="B95C6C3DFFCBBB44C0CAF919E239B40B" bold="true" box="[192,364,1615,1641]" pageId="20" pageNumber="19">Type material.</emphasis>
<materialsCitation id="3B40BA72FFCBBB44C17FF919E6F5B40B" box="[373,1440,1615,1645]" location="National Museum of Natural History" municipality="Smithsonian Institution" pageId="20" pageNumber="19" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B95C6C3DFFCBBB44C17FF919E2BCB40B" bold="true" box="[373,489,1615,1641]" pageId="20" pageNumber="19">
<typeStatus id="54930E8DFFCBBB44C17FF919E2B1B40B" box="[373,484,1615,1641]" pageId="20" pageNumber="19" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D2E7BA6FFCBBB44C1F9F907E15DB40F" box="[499,520,1617,1645]" count="1" pageId="20" pageNumber="19" type="male"></specimenCount>
deposited in the
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,
<collectingMunicipality id="6BF32A55FFCBBB44C486F919E6C9B40B" box="[1164,1436,1615,1641]" pageId="20" pageNumber="19">Smithsonian Institution</collectingMunicipality>
,
</materialsCitation>
</paragraph>
<paragraph id="8B97B02FFFCBBB44C0CAF927E030B573" blockId="20.[191,1441,1619,1887]" pageId="20" pageNumber="19">
<collectingRegion id="49EC7ECDFFCBBB44C0CAF927E21CB4E9" box="[192,329,1649,1675]" country="United States of America" name="Washington" pageId="20" pageNumber="19">Washington</collectingRegion>
,
<collectionCode id="ED3928EAFFCBBB44C159F927E228B4E9" box="[339,381,1649,1675]" pageId="20" pageNumber="19">DC</collectionCode>
,
<collectingCountry id="F33FF0BFFFCBBB44C18CF927E2EEB4E9" box="[390,443,1649,1675]" name="United States of America" pageId="20" pageNumber="19">USA</collectingCountry>
(
<collectionCode id="ED3928EAFFCBBB44C1C3F927E14BB4E9" box="[457,542,1649,1675]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="19" type="Museum">USNM</collectionCode>
), illustrated in
<figureCitation id="1313ACAAFFCBBB44C2CBF927E06DB4E9" box="[705,824,1649,1675]" captionStart="Figures 3348" captionStartId="96.[192,271,1841,1865]" captionTargetBox="[198,1434,372,1745]" captionTargetId="figure-67@96.[192,1440,361,1751]" captionTargetPageId="96" captionText="Figures 3348. Holotypes, data in text. 3334) Ridens angulinea sp. n. 3536) Pythonides lera sp. n. 3738) Pythonides latemarginatus sp. n. 3940) Gindanes variegatus sp. n. 4142) Milanion (Milanion) virga sp. n. 4344) Milanion (Milanion) furvus sp. n. 4546) Milanion (Milanion) laricus sp. n. 4748) Charidia ronda sp. n." pageId="20" pageNumber="19">Fig. 3738</figureCitation>
, bears the following four rectangular labels, three white: [
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:
<collectingCountry id="F33FF0BFFFCBBB44C15BF9C5E2FEB4CF" box="[337,427,1683,1709]" name="Panama" pageId="20" pageNumber="19">Panama</collectingCountry>
| Cerro Campana 2500 |
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<geoCoordinate id="EE1CD6E8FFCBBB44C329F9C5E0D1B4CF" box="[803,900,1682,1711]" degrees="79" direction="west" minutes="55" orientation="longitude" pageId="20" pageNumber="19" precision="925" value="-79.916664">79°55W</geoCoordinate>
|
<date id="FF9696EFFFCBBB44C393F9C5E773B4CF" box="[921,1062,1683,1709]" pageId="20" pageNumber="19" value="1970-08">August 1970</date>
| leg. G.B.Small], [DNA sample ID: | NVG-18018B04 | c/o Nick
<collectionCode id="ED3928EAFFCBBB44C23CF9E3E11CB4AD" box="[566,585,1717,1743]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="20" pageNumber="19" type="Museum">V</collectionCode>
. Grishin], [USNMENT | {QR Code} | 01450934], and one red [
<typeStatus id="54930E8DFFCBBB44C542F9E2E256B492" pageId="20" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D2E7BA6FFCBBB44C100F981E24AB491" box="[266,287,1751,1779]" count="1" pageId="20" pageNumber="19" type="male"></specimenCount>
|
<taxonomicName id="4C28CBACFFCBBB44C139F980E2E4B492" authorityName="Hubner" authorityYear="1819" box="[307,433,1750,1776]" class="Insecta" family="Hesperiidae" genus="Pythonides" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus">Pythonides</taxonomicName>
|
<taxonomicName id="4C28CBACFFCBBB44C1CFF980E193B492" authority="Grishin" authorityName="Zhang &amp; Cong &amp; Grishin" authorityYear="2023" box="[453,710,1750,1776]" class="Insecta" family="Hesperiidae" genus="Pythonides" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="latemarginatus">latemarginatus Grishin</taxonomicName>
].
<emphasis id="B95C6C3DFFCBBB44C2E9F980E007B492" bold="true" box="[739,850,1750,1776]" pageId="20" pageNumber="19">
<typeStatus id="54930E8DFFCBBB44C2E9F980E018B492" box="[739,845,1750,1776]" pageId="20" pageNumber="19" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D2E7BA6FFCBBB44C352F980E02FB491" box="[856,890,1750,1779]" count="1" pageId="20" pageNumber="19" type="male">1♂</specimenCount>
NVG-18018B03 USNMENT_01450933
<collectingCountry id="F33FF0BFFFCBBB44C54AF980E6C8B492" box="[1344,1437,1750,1776]" name="Panama" pageId="20" pageNumber="19">Panama</collectingCountry>
:
<collectingRegion id="49EC7ECDFFCBBB44C0CAF9A1E245B573" box="[192,272,1783,1809]" country="Panama" name="Darien" pageId="20" pageNumber="19">Darien</collectingRegion>
, Cana,
<quantity id="4CD01DCAFFCBBB44C168F9A1E2FFB570" box="[354,426,1783,1810]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="19" unit="m" value="750.0">750 m</quantity>
,
<date id="FF9696EFFFCBBB44C1BEF9A1E117B570" box="[436,578,1783,1810]" pageId="20" pageNumber="19" value="1983-06-15">15-Jun-1983</date>
, G. B. Small leg. [
<collectionCode id="ED3928EAFFCBBB44C308F9A1E00FB573" box="[770,858,1783,1809]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="19" type="Museum">USNM</collectionCode>
].
</paragraph>
<paragraph id="8B97B02FFFCBBB44C0CAF848E64DB55A" blockId="20.[191,1441,1619,1887]" box="[192,1304,1822,1849]" pageId="20" pageNumber="19">
<emphasis id="B95C6C3DFFCBBB44C0CAF848E20EB55A" bold="true" box="[192,347,1822,1848]" pageId="20" pageNumber="19">
<typeStatus id="54930E8DFFCBBB44C0CAF848E3ACB55A" box="[192,249,1822,1848]" pageId="20" pageNumber="19">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33FF0BFFFCBBB44C16BF849E2EBB55B" box="[353,446,1823,1849]" name="Panama" pageId="20" pageNumber="19">Panama</collectingCountry>
:
<collectingRegion id="49EC7ECDFFCBBB44C1CDF849E1D9B55A" box="[455,652,1822,1849]" country="Panama" name="Panama" pageId="20" pageNumber="19">Panama Province</collectingRegion>
, Cerro Campana, elevation
<quantity id="4CD01DCAFFCBBB44C3C9F848E747B55A" box="[963,1042,1822,1849]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="20" pageNumber="19" unit="ft" value="2500.0">2500 ft</quantity>
,
<collectionCode id="ED3928EAFFCBBB44C416F848E718B55A" box="[1052,1101,1822,1848]" pageId="20" pageNumber="19">GPS</collectionCode>
<geoCoordinate id="EE1CD6E8FFCBBB44C459F848E7CAB55A" box="[1107,1183,1822,1848]" degrees="8.6833" direction="north" orientation="latitude" pageId="20" pageNumber="19" precision="5" value="8.6833">8.6833</geoCoordinate>
,
<geoCoordinate id="EE1CD6E8FFCBBB44C4A0F848E641B55A" box="[1194,1300,1822,1848]" degrees="79.9167" direction="west" orientation="longitude" pageId="20" pageNumber="19" precision="5" value="-79.9167">79.9167</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C332E3A4FFCBBB44C0CAF813E297B5E5" pageId="20" pageNumber="19" type="etymology">
<paragraph id="8B97B02FFFCBBB44C0CAF813E297B5E5" blockId="20.[191,1441,1619,1887]" lastBlockId="20.[192,450,1904,1927]" pageId="20" pageNumber="19">
<emphasis id="B95C6C3DFFCBBB44C0CAF813E211B53D" bold="true" box="[192,324,1861,1887]" pageId="20" pageNumber="19">Etymology.</emphasis>
The name is Latin for the broad (dark) margin and is a masculine adjective.
<emphasis id="B95C6C3DFFCBBB44C0CAF83AE20EB5E4" bold="true" box="[192,347,1900,1926]" pageId="20" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="F33FF0BFFFCBBB44C16BF83BE2EBB5E5" box="[353,446,1901,1927]" name="Panama" pageId="20" pageNumber="19">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>