135 lines
11 KiB
XML
135 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2018" docId="E33CD35ED69C227FD8FEE1BF7C00BC9A" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina bewani Shaverdo & Balke, sp. n." docType="treatment" docUuid="8B3BBF57-4E24-446C-BC66-44AB12E15B7E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="13" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="13" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>803</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
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<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125596" LSID="urn:lsid:zoobank.org:act:8B3BBF57-4E24-446C-BC66-44AB12E15B7E" httpUri="http://treatment.plazi.org/id/E33CD35ED69C227FD8FEE1BF7C00BC9A" lastPageNumber="13" pageId="6" pageNumber="13">
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<subSubSection pageId="6" pageNumber="13" type="nomenclature">
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<paragraph pageId="6" pageNumber="13">
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3.
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<taxonomicName LSID="http://zoobank.org/8B3BBF57-4E24-446C-BC66-44AB12E15B7E" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bewani" order="Coleoptera" pageId="6" pageNumber="13" phylum="Arthropoda" rank="species" species="bewani">Exocelina bewani Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="6" pageNumber="13">sp. n.</taxonomicNameLabel>
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Figs 13, 37
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="reference_group">
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<paragraph pageId="6" pageNumber="13">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="6" pageNumber="13" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB1296:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
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: supplementary figs 1-4, table 2;
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary figs S1-S2, table S3.
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</paragraph>
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<paragraph pageId="6" pageNumber="13">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bewani" order="Coleoptera" pageId="6" pageNumber="13" phylum="Arthropoda" rank="species" species="bewani">Exocelina bewani</taxonomicName>
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_New_Guinea_MB1296:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary information S5-S6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="type locality">
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<paragraph pageId="6" pageNumber="13">Type locality.</paragraph>
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<paragraph pageId="6" pageNumber="13">
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Papua New Guinea: Sandaun Province, Bewani Mts, approximately
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.0855">03°05.13'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="141.1705">141°10.23'E</geoCoordinate>
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., 400 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="type material">
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<paragraph pageId="6" pageNumber="13">Type material.</paragraph>
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<paragraph pageId="6" pageNumber="13">Holotype: male "Papua New Guinea: Sandaun, Bewani Stn., limestone creek@base of Bewani Mts., 400 m, 12.iv.2006, nr. 03.05.130S 141.10.227E, Balke & Sagata (PNG 39)", "M.Balke 1296" [green] (ZSM). Paratypes: 2 males, 1 female with the same label as the holotype (NHMW, ZSM).</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="description">
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<paragraph pageId="6" pageNumber="13">Description.</paragraph>
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<paragraph pageId="6" pageNumber="13">Body size and form: Beetle small: TL-H 3.05-3.25 mm, TL 3.35-3.55 mm, MW 1.8-1.9 mm (holotype: TL-H 3.05 mm, TL 3.35 mm, MW 1.8 mm), with broader, oval habitus.</paragraph>
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<paragraph pageId="6" pageNumber="13">Coloration: Brownish, with head and pronotum paler. Head reddish brown to brownish, sometimes paler anteriorly. Pronotum reddish brown on sides, brown to dark brown on disc. Elytra brown to dark brown, sometimes with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 13).</paragraph>
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<paragraph pageId="6" pageNumber="13">
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Surface sculpture: Shiny but with dense, distinct punctation dorsally. As in
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<taxonomicName lsidName="E. sumokedi" pageId="6" pageNumber="13" rank="species" species="sumokedi">E. sumokedi</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph pageId="6" pageNumber="13">Structures: Pronotum with lateral bead. Its lateral sides with shallow longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, narrowly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate.</paragraph>
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<paragraph pageId="6" pageNumber="13">
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Male: Antennae simple (Fig. 13). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior row of 21 and posterior row of 4 relatively short setae (Fig. 37D). Median lobe in lateral view slightly curved, with slightly thickened, angulate, and curved downwards apex; in ventral view, subparallel, with broad and slightly concave apex. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae sparser (Fig. 37
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Abdominal ventrite 6 with 3 or 4 lateral striae on each side.
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</paragraph>
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<paragraph pageId="6" pageNumber="13">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="affinities">
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<paragraph pageId="6" pageNumber="13">Affinities.</paragraph>
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<paragraph pageId="6" pageNumber="13">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina bewani" order="Coleoptera" pageId="6" pageNumber="13" phylum="Arthropoda" rank="species" species="bewani">Exocelina bewani</taxonomicName>
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sp. n. is very similar to
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<taxonomicName lsidName="E. sumokedi" pageId="6" pageNumber="13" rank="species" species="sumokedi">E. sumokedi</taxonomicName>
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sp. n. but is larger and has a less striated abdominal ventrite 6; also, its median lobe is not narrowed distally in ventral view and with apex broad and slightly concave ventrally and curved downwards in lateral view. The species is also similar to
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<taxonomicName lsidName="E. adelbertensis" pageId="6" pageNumber="13" rank="species" species="adelbertensis">E. adelbertensis</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="E. cyclops" pageId="6" pageNumber="13" rank="species" species="cyclops">E. cyclops</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="E. pseudopusilla" pageId="6" pageNumber="13" rank="species" species="pseudopusilla">E. pseudopusilla</taxonomicName>
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sp. n., and
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<taxonomicName lsidName="E. pusilla" pageId="6" pageNumber="13" rank="species" species="pusilla">E. pusilla</taxonomicName>
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sp. n., see their
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<normalizedToken originalValue="“Affinities”">"Affinities"</normalizedToken>
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and the
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<normalizedToken originalValue="“Key”">"Key"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="distribution">
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<paragraph pageId="6" pageNumber="13">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="13">Papua New Guinea: Sandaun Province. The species is known only from the type locality (Fig. 50).</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="13" type="etymology">
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<paragraph pageId="6" pageNumber="13">Etymology.</paragraph>
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<paragraph pageId="6" pageNumber="13">The species is named after the Bewani Mountains. The name is a noun in the nominative singular standing in apposition.</paragraph>
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</subSubSection>
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</treatment>
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</document> |