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<document id="BD5CCA8D9D03871BE8C4D5323C53BC1A" ID-CLB-Dataset="2463" ID-DOI="10.11646/phytotaxa.531.3.5" ID-GBIF-Dataset="6bee07be-4cc1-4f28-8c30-ee299dd9cf5f" ID-ISSN="1179-3163" ID-Zenodo-Dep="5886338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642750472595" checkinUser="plazi" docAuthor="Loeuille, Benoît, Bringel Jr, João Bernardo De A., Faria Jr, Jair E. Q. &amp; Valls, José F. M." docDate="2022" docId="E30487A8FF98905BFF62F9BDFEF7FE69" docLanguage="en" docName="phytotaxa.531.3.5.pdf" docOrigin="Phytotaxa 531 (3)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Eremanthus tomentosus Loeuille &amp; Bringel 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="253" masterDocId="1F3DFFD0FF99905DFFEAFFC9FFC0FFDD" masterDocTitle="Three new species of Lychnophorinae from the Brazilian Central Plateau (Asteraceae: Vernonieae)" masterLastPageNumber="266" masterPageNumber="249" pageNumber="250" updateTime="1699303671397" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2297295B7C061E856B2414C6BFADAD1E">Three new species of Lychnophorinae from the Brazilian Central Plateau (Asteraceae: Vernonieae)</mods:title>
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<mods:namePart id="6F95D6DA6DE061EAF0F6CDA8C3C5827E">Loeuille, Benoît</mods:namePart>
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<mods:namePart id="C42BB2699019C653A0373DF524D44005">Bringel Jr, João Bernardo De A.</mods:namePart>
<mods:affiliation id="A4651CC33EF9C39BDB43D897E7442F4C">Departamento de Botânica, Instituto de Biologia, Universidade de Brasília- UnB, Asa Norte, Campus Darcy Ribeiro, CEP 70910 - 900, Brasília, DF, Brazil</mods:affiliation>
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<mods:namePart id="945C4B69146F48B336C4B70DFB070058">Faria Jr, Jair E. Q.</mods:namePart>
<mods:affiliation id="003CDB55685B3D89562B1104A3179078">Superintendência de Conservação. Jardim Botânico de Brasília, Área Especial SMDB Estação Ecológica Jardim Botânico de Brasília, CEP 71680 - 001, Brasília, DF, Brazil</mods:affiliation>
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<mods:namePart id="0A7301232DA7A3F3CB30ED53790BB646">Valls, José F. M.</mods:namePart>
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<mods:date id="42152676F2BB1AE1F07CE8CC2AB834E3">2022</mods:date>
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<emphasis id="59D9EAACFF98905CFF62F9BDFE61F953" bold="true" box="[136,417,1652,1679]" italics="true" pageId="1" pageNumber="250">Eremanthus tomentosus</emphasis>
Loeuille &amp; Bringel
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(
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, 
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<emphasis id="59D9EAACFF98905CFF62F955FD95F924" italics="true" pageId="1" pageNumber="250">Species Eremantho cincto caulibus rufo-brunneo tomentosis, syncephaliis hemisphaericis, bracteis secundariis conspicuis similis, sed foliis petiolatis (non sessilibus), supra tomentosis (non glabratis), capitulis per 3/4 longitudinem connatis (non per totam longitudinem) et cypselis rubro-glanduliferis basi differt.</emphasis>
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— 
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: 
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, seguindo pela GO-020 atravessando o 
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, elev. 
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1713
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(
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!; isotypes 
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!, 
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!, 
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!)
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—Treelet up to 
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. Stems moderately branched, reddish-brown tomentose with semicircular leaf scars following leaf fall,younger branches furrowed.Leaves alternate, simple;petiole 
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long, upper leaves subsessile, base enlarged; blade oblong or elliptic, sometimes narrow obovate, 5.7–10.1 × 
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, venation brochidodromous, midrib and secondary veins prominent abaxially, midrib furrowed, adaxial surface willow green, tomentose, abaxial surface green-greyish, sometimes slightly reddish-brown, densely tomentose, tomentum of unbranched, long, thin trichomes, 3- to 5-armed, bladder-like trichomes and 3- to 5-armed not swollen trichomes, coriaceous, margins entire, flat, apex obtuse or slightly acute, base acute, sometimes obtuse. Capitula fused in a syncephalium (secondary order inflorescence), sometimes 2–5 syncephalia juxtaposed into a tertiary order inflorescence, organized in a terminal cyme (not compound) of 3–7 syncephalia. Syncephalium 
<quantity id="AC559B5BFF9B905FFD2FFEE7FC81FE94" box="[709,833,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.4" metricValueMin="1.1" pageId="2" pageNumber="251" unit="cm" value="1.75" valueMax="2.4" valueMin="1.1">1.1–2.4 cm</quantity>
tall, 
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diam., hemispherical; secondary bracts ovate, base obtuse, leaf-like, 0.5–2 × 
<quantity id="AC559B5BFF9B905FFDC5FE9BFD6DFEB0" box="[559,685,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="2" pageNumber="251" unit="cm" value="0.6" valueMax="0.9" valueMin="0.3">0.3–0.9 cm</quantity>
long. Capitula 20–ca. 100, homogamous, discoid, sessile, fused 3/4 of length; involucre 
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tall, 
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diam., obconic, 4–5-seriate, subequal; phyllaries weakly imbricate, narrow elliptic or narrow oblanceolate, light greenish, upper half darker, apex obtuse, densely lanulose, margins scarious, persistent, outer phyllaries 2.5–3 × 
<quantity id="AC559B5BFF9B905FFD97FE77FCC6FE04" box="[637,774,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="251" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
, inner phyllaries 3.5–4.6 × 
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long; receptacle flat, naked. Florets 1 per capitulum, bisexual, fertile; corolla actinomorphic, deeply 5-lobed, lilac, corolla tube 2.5–3.2 × 
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, villose towards base, corolla lobes 3–3.2 × 
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, glandular-punctate, pilose towards apices, apex acute; anther whitish or lilac, apical anther appendages lanceolate, acute, anther base sagittate, acute; style shaft 
<quantity id="AC559B5BFF9B905FFF62FD87FED1FDB4" box="[136,273,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="2" pageNumber="251" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
long, white or pale lilac, glabrous throughout except for pubescent upper ca. 
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beneath style arms, style arms 
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long, apex acute, pubescent outside, hairs acute, style base glabrous, lacking basal node. Cypsela turbinate, 2.9–3.1 × 
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, obscurely-ribbed, light brown, sericeous, densely reddish glandular-punctate at base (when fully developed); carpopodium inconspicuous; pappus setae 3–4-seriate, stramineous, paleaceous, persistent, ± serrulate, outermost series setae 
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, innermost series setae 
<quantity id="AC559B5BFF9B905FFC85FD17FC24FD24" box="[879,996,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.1" metricValueMin="4.0" pageId="2" pageNumber="251" unit="mm" value="4.55" valueMax="5.1" valueMin="4.0">4–5.1 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="3FD26636FF9B905FFF62F80AFDBFF7FC" ID-DOI="http://doi.org/10.5281/zenodo.5886340" ID-Zenodo-Dep="5886340" httpUri="https://zenodo.org/record/5886340/files/figure.png" pageId="2" pageNumber="251" startId="2.[136,229,1987,2009]" targetBox="[269,1311,804,1933]" targetPageId="2">
<paragraph id="6B1236BEFF9B905FFF62F80AFDBFF7FC" blockId="2.[136,1452,1987,2081]" pageId="2" pageNumber="251">
<emphasis id="59D9EAACFF9B905FFF62F80AFF3EF805" bold="true" box="[136,254,1987,2009]" pageId="2" pageNumber="251">FIGURE 1.</emphasis>
<taxonomicName id="ACAD4D3DFF9B905FFEEEF80AFE29F805" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[260,489,1987,2008]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="251" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9B905FFEEEF80AFE29F805" box="[260,489,1987,2008]" italics="true" pageId="2" pageNumber="251">Eremanthus tomentosus</emphasis>
</taxonomicName>
. 
<emphasis id="59D9EAACFF9B905FFE1FF80AFDC6F805" bold="true" box="[501,518,1987,2008]" pageId="2" pageNumber="251">A</emphasis>
. Flowering branch. 
<emphasis id="59D9EAACFF9B905FFD23F80AFD19F805" bold="true" box="[713,729,1987,2008]" pageId="2" pageNumber="251">B</emphasis>
. Leaf adaxial surface. 
<emphasis id="59D9EAACFF9B905FFC5EF80AFC05F804" bold="true" box="[948,965,1987,2009]" pageId="2" pageNumber="251">C</emphasis>
. Leaf abaxial surface. 
<emphasis id="59D9EAACFF9B905FFB4BF80AFB72F805" bold="true" box="[1185,1202,1987,2008]" pageId="2" pageNumber="251">D</emphasis>
. Cyme of syncephalia. 
<emphasis id="59D9EAACFF9B905FFA7FF80AFA65F805" bold="true" box="[1429,1445,1987,2008]" pageId="2" pageNumber="251">E</emphasis>
. Tertiary order inflorescence. 
<emphasis id="59D9EAACFF9B905FFE4BF82EFE70F821" bold="true" box="[417,432,2023,2044]" pageId="2" pageNumber="251">F</emphasis>
. Longitudinal section of syncephalium. 
<emphasis id="59D9EAACFF9B905FFCD2F82EFC8BF820" bold="true" box="[824,843,2023,2045]" pageId="2" pageNumber="251">G</emphasis>
. Corolla, androecium and upper part of style. 
<emphasis id="59D9EAACFF9B905FFAE7F82EFAE0F821" bold="true" box="[1293,1312,2023,2044]" pageId="2" pageNumber="251">H</emphasis>
. Cypsela with pappus. 
<emphasis id="59D9EAACFF9B905FFF32F7C2FECBF7FD" bold="true" box="[216,267,2059,2080]" pageId="2" pageNumber="251">A.–I.</emphasis>
from 
<emphasis id="59D9EAACFF9B905FFEAFF7C2FDF6F7FD" box="[325,566,2059,2080]" italics="true" pageId="2" pageNumber="251">Bringel Jr. &amp; Faria 1713</emphasis>
(UFP).
</paragraph>
</caption>
<caption id="3FD26636FF9A905EFF62F8FBFC1FF8B6" ID-DOI="http://doi.org/10.5281/zenodo.5886344" ID-Zenodo-Dep="5886344" httpUri="https://zenodo.org/record/5886344/files/figure.png" pageId="3" pageNumber="252" startId="3.[136,229,1842,1864]" targetBox="[152,1435,191,1808]" targetPageId="3">
<paragraph id="6B1236BEFF9A905EFF62F8FBFC1FF8B6" blockId="3.[136,1452,1842,1900]" pageId="3" pageNumber="252">
<emphasis id="59D9EAACFF9A905EFF62F8FBFF3EF89A" bold="true" box="[136,254,1842,1864]" pageId="3" pageNumber="252">FIGURE 2.</emphasis>
<taxonomicName id="ACAD4D3DFF9A905EFEEFF8FBFE2BF895" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[261,491,1842,1864]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="252" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9A905EFEEFF8FBFE2BF895" box="[261,491,1842,1864]" italics="true" pageId="3" pageNumber="252">Eremanthus tomentosus</emphasis>
</taxonomicName>
. 
<emphasis id="59D9EAACFF9A905EFE12F8FBFDC9F89A" bold="true" box="[504,521,1842,1863]" pageId="3" pageNumber="252">A</emphasis>
. Habitat. 
<emphasis id="59D9EAACFF9A905EFD80F8FBFDBAF89A" bold="true" box="[618,634,1842,1863]" pageId="3" pageNumber="252">B</emphasis>
. Habit. 
<emphasis id="59D9EAACFF9A905EFD23F8FBFD1AF895" bold="true" box="[713,730,1842,1864]" pageId="3" pageNumber="252">C</emphasis>
. Stem. 
<emphasis id="59D9EAACFF9A905EFCCFF8FBFCF6F89A" bold="true" box="[805,822,1842,1863]" pageId="3" pageNumber="252">D</emphasis>
. Leaf abaxial surface. 
<emphasis id="59D9EAACFF9A905EFBFFF8FBFBE5F89A" bold="true" box="[1045,1061,1842,1863]" pageId="3" pageNumber="252">E</emphasis>
. Cyme of syncephalia. 
<emphasis id="59D9EAACFF9A905EFAE1F8FBFADAF89A" bold="true" box="[1291,1306,1842,1863]" pageId="3" pageNumber="252">F</emphasis>
. Tertiary order inflorescence. 
<emphasis id="59D9EAACFF9A905EFEFEF89FFEE7F8B1" bold="true" box="[276,295,1878,1900]" pageId="3" pageNumber="252">G</emphasis>
. Fruiting syncephalium. 
<emphasis id="59D9EAACFF9A905EFDFDF89FFD91F8B6" bold="true" box="[535,593,1878,1899]" pageId="3" pageNumber="252">A.–E.</emphasis>
, 
<emphasis id="59D9EAACFF9A905EFDB7F89FFDB0F8B1" bold="true" box="[605,624,1878,1900]" pageId="3" pageNumber="252">G</emphasis>
. by J.E.Q. Faria. 
<emphasis id="59D9EAACFF9A905EFCF2F89FFCEAF8B6" bold="true" box="[792,810,1878,1899]" pageId="3" pageNumber="252">F.</emphasis>
by J.B. Bringel Jr.
</paragraph>
</caption>
<subSubSection id="23B76535FF9A9059FF57F853FBECFF00" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252" type="distribution">
<paragraph id="6B1236BEFF9A9059FF57F853FBECFF00" blockId="3.[136,1451,1946,2081]" lastBlockId="4.[136,1452,158,221]" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">
<emphasis id="59D9EAACFF9A905EFF57F853FD1CF869" bold="true" box="[189,732,1946,1972]" pageId="3" pageNumber="252">Distribution, habitat, and conservation status:</emphasis>
— 
<taxonomicName id="ACAD4D3DFF9A905EFD12F853FBC2F869" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[760,1026,1946,1972]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="252" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9A905EFD12F853FBC2F869" box="[760,1026,1946,1972]" italics="true" pageId="3" pageNumber="252">Eremanthus tomentosus</emphasis>
</taxonomicName>
grows in hillsides in 
<emphasis id="59D9EAACFF9A905EFB02F852FAA7F869" box="[1256,1383,1947,1972]" italics="true" pageId="3" pageNumber="252">campo sujo</emphasis>
(open grassland with few, scattered shrubs or treelets) on cambisol with small quartzitic rocks. It is currently known from three populations in southeastern 
<collectingRegion id="A969F85CFF9A905EFE14F82BFDFFF821" box="[510,575,2018,2044]" country="Brazil" name="Goias" pageId="3" pageNumber="252">Goiás</collectingRegion>
and surrounding areas in 
<collectingRegion id="A969F85CFF9A905EFCB4F82BFC36F821" box="[862,1014,2018,2044]" country="Brazil" name="Minas Gerais" pageId="3" pageNumber="252">Minas Gerais</collectingRegion>
, near the São Marcos river. In addition, a fourth population was found about 
<quantity id="AC559B5BFF9A905EFDD8F7CFFD49F7FD" box="[562,649,2054,2080]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="252" unit="km" value="160.0">160 km</quantity>
away, in the Brasília Botanical Garden reserve in the 
<collectingRegion id="A969F85CFF9A905EFB13F7CFFA6BF7FD" box="[1273,1451,2054,2080]" country="Brazil" name="Distrito Federal" pageId="3" pageNumber="252">Distrito Federal</collectingRegion>
(
<figureCitation id="F3962A3BFF9D9059FF7AFF57FF15FF65" box="[144,213,158,184]" captionStart="FIGURE 3" captionStartId="4.[136,229,1143,1165]" captionTargetBox="[161,1429,255,1110]" captionTargetId="figure-58@4.[151,1436,245,1119]" captionTargetPageId="4" captionText="FIGURE 3. Distribution of the three new species of Lychnophorinae (Vernonieae – Asteraceae)." figureDoi="http://doi.org/10.5281/zenodo.5886346" httpUri="https://zenodo.org/record/5886346/files/figure.png" pageId="4" pageNumber="253">Fig. 3</figureCitation>
). According to the 
<bibRefCitation id="0F3C4B4FFF9D9059FE46FF57FD82FF65" author="IUCN" box="[428,578,158,184]" pageId="4" pageNumber="253" refId="ref8773" refString="IUCN (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 25 June 2021)" type="url" year="2019">IUCN (2019)</bibRefCitation>
criterion of geographic range, this species is recognized as Endangered, since its extent of occurrence is estimated in 
<quantity id="AC559B5BFF9D9059FDD4FF0BFD68FF01" box="[574,680,194,220]" metricMagnitude="6" metricUnit="m" metricValue="2.327" pageId="4" pageNumber="253" unit="km" value="2327.0">2,327 km</quantity>
<superScript id="9CD89BF6FF9D9059FD42FF0BFD70FF0D" attach="left" box="[680,688,194,208]" fontSize="6" pageId="4" pageNumber="253">2</superScript>
and area of occupancy in 
<quantity id="AC559B5BFF9D9059FC3CFF0BFBDDFF01" box="[982,1053,194,220]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="253" unit="km" value="16.0">16 km</quantity>
<superScript id="9CD89BF6FF9D9059FBF6FF0BFBE4FF0D" attach="left" box="[1052,1060,194,208]" fontSize="6" pageId="4" pageNumber="253">2</superScript>
.
</paragraph>
</subSubSection>
<caption id="3FD26636FF9D9059FF62FBBEFBE9FB53" ID-DOI="http://doi.org/10.5281/zenodo.5886346" ID-Zenodo-Dep="5886346" box="[136,1065,1143,1166]" httpUri="https://zenodo.org/record/5886346/files/figure.png" pageId="4" pageNumber="253" startId="4.[136,229,1143,1165]" targetBox="[161,1429,255,1110]" targetPageId="4">
<paragraph id="6B1236BEFF9D9059FF62FBBEFBE9FB53" blockId="4.[136,1065,1143,1166]" box="[136,1065,1143,1166]" pageId="4" pageNumber="253">
<emphasis id="59D9EAACFF9D9059FF62FBBEFF3EFB51" bold="true" box="[136,254,1143,1165]" pageId="4" pageNumber="253">FIGURE 3.</emphasis>
Distribution of the three new species of 
<taxonomicName id="ACAD4D3DFF9D9059FD6EFBB1FCE0FB53" box="[644,800,1144,1166]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="264" phylum="Tracheophyta" rank="subTribe" subTribe="Lychnophorinae">Lychnophorinae</taxonomicName>
(
<taxonomicName id="ACAD4D3DFF9D9059FCC6FBB1FC5AFB53" authorityName="Cassini &amp; J.Phys.Chim.Hist.Nat" authorityYear="1819" box="[812,922,1144,1166]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="250" phylum="Tracheophyta" rank="tribe" tribe="Vernonieae">Vernonieae</taxonomicName>
– 
<taxonomicName id="ACAD4D3DFF9D9059FC5BFBB1FBDFFB53" box="[945,1055,1144,1166]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</caption>
<subSubSection id="23B76535FF9D9059FF57FB75FC30F9BE" pageId="4" pageNumber="253" type="etymology">
<paragraph id="6B1236BEFF9D9059FF57FB75FF21FB27" blockId="4.[136,1452,1212,2067]" pageId="4" pageNumber="253">
<emphasis id="59D9EAACFF9D9059FF57FB75FE88FB0B" bold="true" box="[189,328,1212,1238]" pageId="4" pageNumber="253">Etymology:</emphasis>
—The species epithet refers to the tomentose indument covering the adaxial leaf surface of the new species.
</paragraph>
<paragraph id="6B1236BEFF9D9059FF57FACDFC30F9BE" blockId="4.[136,1452,1212,2067]" pageId="4" pageNumber="253">
<materialsCitation id="DBC53CE3FF9D9059FF57FACDFBBEFA9F" ID-GBIF-Occurrence="3450195329" collectionCode="J" country="Brazil" elevation="1125" latitude="-15.916111" location="Morro do Urubu" longLatPrecision="21" longitude="-47.891113" municipality="Brasilia. Estacao Ecologica Jardim Botanico de Brasilia" pageId="4" pageNumber="253" specimenCount="2" stateProvince="Distrito Federal" typeStatus="paratype">
<emphasis id="59D9EAACFF9D9059FF57FACDFD0AFAC3" bold="true" box="[189,714,1284,1310]" pageId="4" pageNumber="253">
Additional specimens examined (
<typeStatus id="B416881CFF9D9059FDA3FACDFD03FAC3" box="[585,707,1284,1310]" pageId="4" pageNumber="253" type="paratype">paratypes</typeStatus>
)
</emphasis>
:— 
<collectingCountry id="13BA762EFF9D9059FD06FACDFC97FAC3" box="[748,855,1284,1310]" name="Brazil" pageId="4" pageNumber="253">BRAZIL</collectingCountry>
. 
<collectingRegion id="A969F85CFF9D9059FC88FACDFBD7FAC3" box="[866,1047,1284,1310]" country="Brazil" name="Distrito Federal" pageId="4" pageNumber="253">Distrito Federal</collectingRegion>
. 
<collectingMunicipality id="8B76ACC4FF9D9059FBC8FACDFEB2FA9F" pageId="4" pageNumber="253">Brasília. Estação Ecológica Jardim Botânico de Brasília</collectingMunicipality>
, área do 
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, 
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, 
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, elev. 
<quantity id="AC559B5BFF9D9059FBFAFAE1FBA1FA9E" box="[1040,1121,1320,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="4" pageNumber="253" unit="m" value="1125.0">
<elevation id="E080D18DFF9D9059FBFAFAE1FBA1FA9E" box="[1040,1121,1320,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="4" pageNumber="253" unit="m" value="1125.0">1125m</elevation>
</quantity>
. 
<collectionCode id="0DBCAE7BFF9D9059FB87FAE1FBBEFA9F" box="[1133,1150,1320,1346]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">
<emphasis id="59D9EAACFF9D9059FB87FAE1FBBEFA9F" box="[1133,1150,1320,1346]" italics="true" pageId="4" pageNumber="253">J</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="59D9EAACFF9D9059FB97FAE1FB74FA9F" box="[1149,1204,1320,1346]" italics="true" pageId="4" pageNumber="253">
. 
<materialsCitation id="DBC53CE3FF9D9059FB6BFAE1FB56FA9F" ID-GBIF-Occurrence="3450195309" box="[1153,1174,1320,1346]" collectionCode="E" country="Brazil" elevation="1125" isTailEnumeration="true" latitude="-15.916111" location="Morro do Urubu" longLatPrecision="21" longitude="-47.891113" municipality="Brasilia. Estacao Ecologica Jardim Botanico de Brasilia" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Distrito Federal">
<collectionCode id="0DBCAE7BFF9D9059FB6BFAE1FB56FA9F" box="[1153,1174,1320,1346]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
.Q.
</emphasis>
<materialsCitation id="DBC53CE3FF9D9059FB56FAE0FF3DFABA" ID-GBIF-Occurrence="3450195304" collectionCode="HEPH, UB, UFP" country="Brazil" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<emphasis id="59D9EAACFF9D9059FB56FAE0FA8AFA9F" box="[1212,1354,1320,1346]" italics="true" pageId="4" pageNumber="253">
<location id="6E726065FF9D9059FB56FAE0FB3CFA9F" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FB56FAE0FB3CFA9F" box="[1212,1276,1321,1346]" country="Brazil" name="Faria" pageId="4" pageNumber="253">Faria</location>
10684
</emphasis>
(
<collectionCode id="0DBCAE7BFF9D9059FAB0FAE1FA68FA9F" box="[1370,1448,1320,1346]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15477" lsid="urn:lsid:biocol.org:col:15477" name="Jardim Botânico de Brasília" pageId="4" pageNumber="253" type="Herbarium">HEPH</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FF62FA84FF71FABA" box="[136,177,1357,1383]" country="France" name="Laboratoire de Biostratigraphie" pageId="4" pageNumber="253">UB</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FF57FA84FF35FABA" box="[189,245,1357,1383]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15350" lsid="urn:lsid:biocol.org:col:15350" name="Universidade Federal de Pernambuco" pageId="4" pageNumber="253" type="Herbarium">UFP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FEE2FA85FDDAFABB" ID-GBIF-Occurrence="3450195343" box="[264,538,1356,1382]" country="Brazil" location="Campo Alegre" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Goias">
<collectingRegion id="A969F85CFF9D9059FEE2FA85FE8DFABB" box="[264,333,1356,1382]" country="Brazil" name="Goias" pageId="4" pageNumber="253">Goiás</collectingRegion>
. 
<location id="6E726065FF9D9059FEBDFA85FE39FABB" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FEBDFA85FE39FABB" box="[343,505,1356,1382]" country="Brazil" name="Campo Alegre" pageId="4" pageNumber="253" stateProvince="Goias">Campo Alegre</location>
de
</materialsCitation>
<materialsCitation id="DBC53CE3FF9D9059FDC8FA85FDB1FA57" ID-GBIF-Occurrence="3450195324" collectingDate="2020-06-06" collectionCode="NE, J" country="Brazil" elevation="917" latitude="-17.541945" location="21 km NE de Campo Alegre" longLatPrecision="21" longitude="-47.656113" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Goias">
<collectingRegion id="A969F85CFF9D9059FDC8FA85FDA6FABB" box="[546,614,1356,1382]" country="Brazil" name="Goias" pageId="4" pageNumber="253">Goiás</collectingRegion>
, estrada à direita da GO-213 ca. 
<location id="6E726065FF9D9059FC3EFA85FAD5FABB" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FC3EFA85FAD5FABB" box="[980,1301,1356,1383]" country="Brazil" latitude="-17.541945" longLatPrecision="21" longitude="-47.656113" name="21 km NE de Campo Alegre" pageId="4" pageNumber="253" stateProvince="Goias">
<quantity id="AC559B5BFF9D9059FC3EFA85FBDBFABB" box="[980,1051,1356,1382]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="253" unit="km" value="21.0">21 km</quantity>
<collectionCode id="0DBCAE7BFF9D9059FBC8FA84FB87FABA" box="[1058,1095,1357,1383]" country="Australia" lsid="urn:lsid:biocol.org:col:14575" name="University of New England" pageId="4" pageNumber="253" type="Herbarium">NE</collectionCode>
de Campo Alegre
</location>
, 
<geoCoordinate id="0E995079FF9D9059FACAFA85FA67FABB" box="[1312,1447,1356,1382]" degrees="17" direction="south" minutes="32" orientation="latitude" pageId="4" pageNumber="253" precision="15" seconds="31" value="-17.541945">17°32’31”S</geoCoordinate>
, 
<geoCoordinate id="0E995079FF9D9059FF62FAB9FED8FA57" box="[136,280,1392,1418]" degrees="47" direction="west" minutes="39" orientation="longitude" pageId="4" pageNumber="253" precision="15" seconds="22" value="-47.656113">47°39’22”W</geoCoordinate>
, elev. 
<quantity id="AC559B5BFF9D9059FE8BFAB9FE6BFA56" box="[353,427,1392,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.17" pageId="4" pageNumber="253" unit="m" value="917.0">
<elevation id="E080D18DFF9D9059FE8BFAB9FE6BFA56" box="[353,427,1392,1419]" metricMagnitude="2" metricUnit="m" metricValue="9.17" pageId="4" pageNumber="253" unit="m" value="917.0">917 m</elevation>
</quantity>
, 
<date id="1F13107EFF9D9059FE53FAB9FD94FA57" box="[441,596,1392,1419]" pageId="4" pageNumber="253" value="2020-06-06">
<collectingDate id="0F57E996FF9D9059FE53FAB9FD94FA57" box="[441,596,1392,1419]" pageId="4" pageNumber="253" value="2020-06-06">06 June 2020</collectingDate>
</date>
, 
<collectionCode id="0DBCAE7BFF9D9059FD8BFAB9FDB1FA57" box="[609,625,1392,1418]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">
<emphasis id="59D9EAACFF9D9059FD8BFAB9FDB1FA57" box="[609,625,1392,1418]" italics="true" pageId="4" pageNumber="253">J</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="59D9EAACFF9D9059FD9BFAB9FC53FA57" box="[625,915,1392,1418]" italics="true" pageId="4" pageNumber="253">
. 
<materialsCitation id="DBC53CE3FF9D9059FD9FFAB9FD49FA57" ID-GBIF-Occurrence="3450195313" box="[629,649,1392,1418]" collectingDate="2020-06-06" collectionCode="B" country="Brazil" elevation="917" isTailEnumeration="true" latitude="-17.541945" location="21 km NE de Campo Alegre" longLatPrecision="21" longitude="-47.656113" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Goias">
<collectionCode id="0DBCAE7BFF9D9059FD9FFAB9FD49FA57" box="[629,649,1392,1418]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="253" type="Herbarium">B</collectionCode>
</materialsCitation>
.A. 
<materialsCitation id="DBC53CE3FF9D9059FD47FAB9FC9DFA57" ID-GBIF-Occurrence="3450195305" box="[685,861,1392,1418]" collectionCode="J" country="Brazil" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
<location id="6E726065FF9D9059FD47FAB9FCE5FA57" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FD47FAB9FCE5FA57" box="[685,805,1392,1418]" country="Brazil" name="Bringel Jr." pageId="4" pageNumber="253">Bringel Jr.</location>
&amp; 
<collectionCode id="0DBCAE7BFF9D9059FCA6FAB9FC9DFA57" box="[844,861,1392,1418]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">J</collectionCode>
</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FC8AFAB9FCB5FA57" ID-GBIF-Occurrence="3450195301" box="[864,885,1392,1418]" collectionCode="E" country="Brazil" isTailEnumeration="true" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FC8AFAB9FCB5FA57" box="[864,885,1392,1418]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
.Q.
</emphasis>
<materialsCitation id="DBC53CE3FF9D9059FC71FAB8FDD0FA73" ID-GBIF-Occurrence="3450195341" collectionCode="CEN, UFP, GO" country="Brazil" location="Campo Alegre" municipality="Faria" pageId="4" pageNumber="253" specimenCount="1">
<emphasis id="59D9EAACFF9D9059FC71FAB8FBDBFA57" box="[923,1051,1392,1418]" italics="true" pageId="4" pageNumber="253">
<collectingMunicipality id="8B76ACC4FF9D9059FC71FAB8FC1BFA57" box="[923,987,1393,1418]" pageId="4" pageNumber="253">Faria</collectingMunicipality>
1700
</emphasis>
(
<collectionCode id="0DBCAE7BFF9D9059FBC6FAB9FBA8FA57" box="[1068,1128,1392,1418]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="4" pageNumber="253" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FB9FFAB8FB6DFA56" box="[1141,1197,1393,1419]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15350" lsid="urn:lsid:biocol.org:col:15350" name="Universidade Federal de Pernambuco" pageId="4" pageNumber="253" type="Herbarium">UFP</collectionCode>
); margem da GO-213, ca. 
<quantity id="AC559B5BFF9D9059FF5AFA5DFF38FA73" box="[176,248,1428,1454]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="253" unit="km" value="13.0">13 km</quantity>
de 
<location id="6E726065FF9D9059FEC9FA5DFE06FA73" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FEC9FA5DFE06FA73" box="[291,454,1428,1454]" country="Brazil" municipality="Faria" name="Campo Alegre" pageId="4" pageNumber="253">Campo Alegre</location>
– 
<collectionCode id="0DBCAE7BFF9D9059FE0FFA5DFDD0FA73" box="[485,528,1428,1454]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13862" name="Philosophical Society" pageId="4" pageNumber="253" type="Herbarium">GO</collectionCode>
</materialsCitation>
, 
<materialsCitation id="DBC53CE3FF9D9059FDF7FA5DFBC3FA73" ID-GBIF-Occurrence="3450195330" box="[541,1027,1428,1455]" collectingDate="2020-06-06" collectionCode="J" country="Brazil" latitude="-17.54861" longLatPrecision="21" longitude="-47.7" pageId="4" pageNumber="253" specimenCount="1">
<geoCoordinate id="0E995079FF9D9059FDF7FA5DFD64FA72" box="[541,676,1428,1455]" degrees="17" direction="south" minutes="32" orientation="latitude" pageId="4" pageNumber="253" precision="15" seconds="55" value="-17.54861">17°32’55”S</geoCoordinate>
, 
<geoCoordinate id="0E995079FF9D9059FD5AFA5DFCFFFA73" box="[688,831,1428,1454]" degrees="47" direction="west" minutes="42" orientation="longitude" pageId="4" pageNumber="253" precision="15" seconds="00" value="-47.7">47°42’00”W</geoCoordinate>
, 
<date id="1F13107EFF9D9059FCA6FA5DFC27FA73" box="[844,999,1428,1455]" pageId="4" pageNumber="253" value="2020-06-06">
<collectingDate id="0F57E996FF9D9059FCA6FA5DFC27FA73" box="[844,999,1428,1455]" pageId="4" pageNumber="253" value="2020-06-06">06 June 2020</collectingDate>
</date>
, 
<collectionCode id="0DBCAE7BFF9D9059FC19FA5DFBC3FA73" box="[1011,1027,1428,1454]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">
<emphasis id="59D9EAACFF9D9059FC19FA5DFBC3FA73" box="[1011,1027,1428,1454]" italics="true" pageId="4" pageNumber="253">J</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="59D9EAACFF9D9059FBEEFA5DFAE3FA73" box="[1028,1315,1428,1454]" italics="true" pageId="4" pageNumber="253">
. 
<materialsCitation id="DBC53CE3FF9D9059FBEDFA5DFBDBFA73" ID-GBIF-Occurrence="3450195331" box="[1031,1051,1428,1454]" collectingDate="2020-06-06" collectionCode="B" country="Brazil" isTailEnumeration="true" latitude="-17.54861" longLatPrecision="21" longitude="-47.7" pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FBEDFA5DFBDBFA73" box="[1031,1051,1428,1454]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="253" type="Herbarium">B</collectionCode>
</materialsCitation>
.A. 
<materialsCitation id="DBC53CE3FF9D9059FBD5FA5DFB2DFA73" ID-GBIF-Occurrence="3450195303" box="[1087,1261,1428,1454]" collectionCode="J" country="Brazil" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
<location id="6E726065FF9D9059FBD5FA5DFB77FA73" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FBD5FA5DFB77FA73" box="[1087,1207,1428,1454]" country="Brazil" name="Bringel Jr." pageId="4" pageNumber="253">Bringel Jr.</location>
&amp; 
<collectionCode id="0DBCAE7BFF9D9059FB36FA5DFB2DFA73" box="[1244,1261,1428,1454]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">J</collectionCode>
</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FB1BFA5DFAC6FA73" ID-GBIF-Occurrence="3450195314" box="[1265,1286,1428,1454]" collectionCode="E" country="Brazil" isTailEnumeration="true" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FB1BFA5DFAC6FA73" box="[1265,1286,1428,1454]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
.Q.
</emphasis>
<materialsCitation id="DBC53CE3FF9D9059FAC1FA5CFEB4FA0F" ID-GBIF-Occurrence="3450195318" collectionCode="CEN, UFP, J" country="Brazil" isTailEnumeration="true" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<emphasis id="59D9EAACFF9D9059FAC1FA5CFA6BFA73" box="[1323,1451,1428,1454]" italics="true" pageId="4" pageNumber="253">
<location id="6E726065FF9D9059FAC1FA5CFAABFA73" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FAC1FA5CFAABFA73" box="[1323,1387,1429,1454]" country="Brazil" name="Faria" pageId="4" pageNumber="253">Faria</location>
1684
</emphasis>
(
<collectionCode id="0DBCAE7BFF9D9059FF7AFA71FF0CFA0F" box="[144,204,1464,1490]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="4" pageNumber="253" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FF3DFA70FECFFA0E" box="[215,271,1465,1491]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15350" lsid="urn:lsid:biocol.org:col:15350" name="Universidade Federal de Pernambuco" pageId="4" pageNumber="253" type="Herbarium">UFP</collectionCode>
); ibid., 
<collectionCode id="0DBCAE7BFF9D9059FE89FA71FEB4FA0F" box="[355,372,1464,1490]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">
<emphasis id="59D9EAACFF9D9059FE89FA71FEB4FA0F" box="[355,372,1464,1490]" italics="true" pageId="4" pageNumber="253">J</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="59D9EAACFF9D9059FE9EFA71FD98FA0F" box="[372,600,1464,1490]" italics="true" pageId="4" pageNumber="253">
. 
<materialsCitation id="DBC53CE3FF9D9059FE92FA71FE4DFA0F" ID-GBIF-Occurrence="3450195350" box="[376,397,1464,1490]" collectionCode="E" country="Brazil" isTailEnumeration="true" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FE92FA71FE4DFA0F" box="[376,397,1464,1490]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
.Q. 
<materialsCitation id="DBC53CE3FF9D9059FE5BFA70FDE4FA0F" ID-GBIF-Occurrence="3450195354" box="[433,548,1464,1490]" collectionCode="J" country="Brazil" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<location id="6E726065FF9D9059FE5BFA70FE31FA0F" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FE5BFA70FE31FA0F" box="[433,497,1465,1490]" country="Brazil" name="Faria" pageId="4" pageNumber="253">Faria</location>
&amp; 
<collectionCode id="0DBCAE7BFF9D9059FDFEFA71FDE4FA0F" box="[532,548,1464,1490]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="253" type="Herbarium">J</collectionCode>
</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FDC3FA71FDFDFA0F" ID-GBIF-Occurrence="3450195351" box="[553,573,1464,1490]" collectionCode="B" country="Brazil" isTailEnumeration="true" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FDC3FA71FDFDFA0F" box="[553,573,1464,1490]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="253" type="Herbarium">B</collectionCode>
</materialsCitation>
.A.
</emphasis>
<materialsCitation id="DBC53CE3FF9D9059FDB5FA71FE4FFA2A" ID-GBIF-Occurrence="3450195334" collectionCode="HDJF, HUEG, RB, UB, UFP, E" country="Brazil" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
<emphasis id="59D9EAACFF9D9059FDB5FA71FCE2FA0F" box="[607,802,1464,1490]" italics="true" pageId="4" pageNumber="253">
<location id="6E726065FF9D9059FDB5FA71FD16FA0F" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FDB5FA71FD16FA0F" box="[607,726,1464,1490]" country="Brazil" name="Bringel Jr." pageId="4" pageNumber="253">Bringel Jr.</location>
10651
</emphasis>
(
<collectionCode id="0DBCAE7BFF9D9059FCDBFA71FCB6FA0F" box="[817,886,1464,1490]" country="0" httpUri="http://grbio.org/cool/fmh6-brwg" name="Universidade Federal dos Vales do Jequitinhonha e Mucuri" pageId="4" pageNumber="253" type="Herbarium">HDJF</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FC6BFA71FC12FA0F" box="[897,978,1464,1490]" country="Brazil" httpUri="http://grbio.org/cool/5x9w-vti9" name="Universidade Estadual de Goiás" pageId="4" pageNumber="253" type="Herbarium">HUEG</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FC37FA70FBC5FA0E" box="[989,1029,1465,1491]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="4" pageNumber="253" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FBFAFA70FBF9FA0E" box="[1040,1081,1465,1491]" country="France" name="Laboratoire de Biostratigraphie" pageId="4" pageNumber="253">UB</collectionCode>
, 
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); rodovia não pavimentada GO-213, ca. 
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do entroncamento com a BR-050, 
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, 
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, 
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</date>
, 
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<emphasis id="59D9EAACFF9D9059FB10FA15FF2DF9C7" italics="true" pageId="4" pageNumber="253">
. 
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.A. 
<materialsCitation id="DBC53CE3FF9D9059FADFFA15FF77F9C7" ID-GBIF-Occurrence="3450195348" collectionCode="J" country="Brazil" location="Bringel Jr." pageId="4" pageNumber="253" specimenCount="1">
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&amp; 
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</materialsCitation>
. 
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</materialsCitation>
.Q.
</emphasis>
<materialsCitation id="DBC53CE3FF9D9059FF1FF9C8FDCFF9C6" ID-GBIF-Occurrence="3450195326" box="[245,527,1536,1563]" collectionCode="CEN, UFP" country="Brazil" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<emphasis id="59D9EAACFF9D9059FF1FF9C8FEB5F9C7" box="[245,373,1536,1562]" italics="true" pageId="4" pageNumber="253">
<location id="6E726065FF9D9059FF1FF9C8FEF5F9C7" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FF1FF9C8FEF5F9C7" box="[245,309,1537,1562]" country="Brazil" name="Faria" pageId="4" pageNumber="253">Faria</location>
1368
</emphasis>
(
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, 
<collectionCode id="0DBCAE7BFF9D9059FE24F9C8FDC6F9C6" box="[462,518,1537,1563]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15350" lsid="urn:lsid:biocol.org:col:15350" name="Universidade Federal de Pernambuco" pageId="4" pageNumber="253" type="Herbarium">UFP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FDF0F9C9FEE9F9E2" ID-GBIF-Occurrence="3450195352" country="Brazil" location="rio Sao Marcos" municipality="Paracatu" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Minas Gerais">
<collectingRegion id="A969F85CFF9D9059FDF0F9C9FD76F9C7" box="[538,694,1536,1562]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="253">Minas Gerais</collectingRegion>
. 
<collectingMunicipality id="8B76ACC4FF9D9059FD2BF9C8FCE4F9C6" box="[705,804,1537,1563]" pageId="4" pageNumber="253">Paracatu</collectingMunicipality>
, seguindo pela GO-020 atravessando o 
<location id="6E726065FF9D9059FB03F9C9FA57F9C6" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FB03F9C9FA57F9C6" box="[1257,1431,1536,1563]" country="Brazil" municipality="Paracatu" name="rio Sao Marcos" pageId="4" pageNumber="253" stateProvince="Minas Gerais">rio São Marcos</location>
e adentrando no
</materialsCitation>
<materialsCitation id="DBC53CE3FF9D9059FED8F9EDFD3AF9E2" ID-GBIF-Occurrence="3450195308" box="[306,762,1572,1599]" collectionCode="E" country="Brazil" location="estado de Minas Gerais" pageId="4" pageNumber="253" specimenCount="1" stateProvince="Minas Gerais">
<collectingRegion id="A969F85CFF9D9059FED8F9EDFD80F9E3" box="[306,576,1572,1598]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="253">estado de Minas Gerais</collectingRegion>
, ca. de 
<quantity id="AC559B5BFF9D9059FD72F9EDFD21F9E3" box="[664,737,1572,1598]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="253" unit="km" value="38.0">38 km</quantity>
<collectionCode id="0DBCAE7BFF9D9059FD03F9ECFD3AF9E2" box="[745,762,1573,1599]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
<materialsCitation id="DBC53CE3FF9D9059FCE9F9EDFF1FF9BF" ID-GBIF-Occurrence="3450195342" collectingDate="2020-06-07" collectionCode="J" country="Brazil" elevation="886" latitude="-17.3775" longLatPrecision="21" longitude="-47.483055" pageId="4" pageNumber="253" specimenCount="1">
da BR-050, 
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, 
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, elev. 
<quantity id="AC559B5BFF9D9059FB11F9EDFA86F9E2" box="[1275,1350,1572,1599]" metricMagnitude="2" metricUnit="m" metricValue="8.86" pageId="4" pageNumber="253" unit="m" value="886.0">
<elevation id="E080D18DFF9D9059FB11F9EDFA86F9E2" box="[1275,1350,1572,1599]" metricMagnitude="2" metricUnit="m" metricValue="8.86" pageId="4" pageNumber="253" unit="m" value="886.0">886 m</elevation>
</quantity>
, 
<date id="1F13107EFF9D9059FAB9F9EDFF03F9BF" pageId="4" pageNumber="253" value="2020-06-07">
<collectingDate id="0F57E996FF9D9059FAB9F9EDFF03F9BF" pageId="4" pageNumber="253" value="2020-06-07">07 June 2020</collectingDate>
</date>
, 
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<emphasis id="59D9EAACFF9D9059FF24F981FF1FF9BF" box="[206,223,1608,1634]" italics="true" pageId="4" pageNumber="253">J</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="59D9EAACFF9D9059FF35F981FE03F9BF" box="[223,451,1608,1634]" italics="true" pageId="4" pageNumber="253">
. 
<materialsCitation id="DBC53CE3FF9D9059FF08F981FF37F9BF" ID-GBIF-Occurrence="3450195323" box="[226,247,1608,1634]" collectingDate="2020-06-07" collectionCode="E" country="Brazil" elevation="886" isTailEnumeration="true" latitude="-17.3775" longLatPrecision="21" longitude="-47.483055" pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FF08F981FF37F9BF" box="[226,247,1608,1634]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="253" type="Herbarium">E</collectionCode>
</materialsCitation>
.Q. 
<materialsCitation id="DBC53CE3FF9D9059FEF6F980FE4FF9BF" ID-GBIF-Occurrence="3450195337" box="[284,399,1608,1634]" collectionCode="J" country="Brazil" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<location id="6E726065FF9D9059FEF6F980FE9CF9BF" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FEF6F980FE9CF9BF" box="[284,348,1609,1634]" country="Brazil" name="Faria" pageId="4" pageNumber="253">Faria</location>
&amp; 
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</materialsCitation>
. 
<materialsCitation id="DBC53CE3FF9D9059FE7EF981FE68F9BF" ID-GBIF-Occurrence="3450195312" box="[404,424,1608,1634]" collectionCode="B" country="Brazil" isTailEnumeration="true" location="Faria" pageId="4" pageNumber="253" specimenCount="1">
<collectionCode id="0DBCAE7BFF9D9059FE7EF981FE68F9BF" box="[404,424,1608,1634]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="253" type="Herbarium">B</collectionCode>
</materialsCitation>
.A.
</emphasis>
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<emphasis id="59D9EAACFF9D9059FE20F981FD4DF9BF" box="[458,653,1608,1634]" italics="true" pageId="4" pageNumber="253">
<location id="6E726065FF9D9059FE20F981FD81F9BF" LSID="urn:lsid:plazi:treatment:E30487A8FF98905BFF62F9BDFEF7FE69:6E726065FF9D9059FE20F981FD81F9BF" box="[458,577,1608,1634]" country="Brazil" name="Bringel Jr." pageId="4" pageNumber="253">Bringel Jr.</location>
10679
</emphasis>
(
<collectionCode id="0DBCAE7BFF9D9059FD76F981FD21F9BF" box="[668,737,1608,1634]" country="0" httpUri="http://grbio.org/cool/fmh6-brwg" name="Universidade Federal dos Vales do Jequitinhonha e Mucuri" pageId="4" pageNumber="253" type="Herbarium">HDJF</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FD06F980FCF8F9BE" box="[748,824,1609,1635]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15477" lsid="urn:lsid:biocol.org:col:15477" name="Jardim Botânico de Brasília" pageId="4" pageNumber="253" type="Herbarium">HEPH</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FCA9F980FCABF9BE" box="[835,875,1609,1635]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="4" pageNumber="253" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FC9DF980FC60F9BE" box="[887,928,1609,1635]" country="France" name="Laboratoire de Biostratigraphie" pageId="4" pageNumber="253">UB</collectionCode>
, 
<collectionCode id="0DBCAE7BFF9D9059FC46F980FC24F9BE" box="[940,996,1609,1635]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15350" lsid="urn:lsid:biocol.org:col:15350" name="Universidade Federal de Pernambuco" pageId="4" pageNumber="253" type="Herbarium">UFP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="23B76535FF9D905BFF57F9A5FEF7FE69" lastPageId="6" lastPageNumber="255" pageId="4" pageNumber="253" type="discussion">
<paragraph id="6B1236BEFF9D905BFF57F9A5FEF7FE69" blockId="4.[136,1452,1212,2067]" lastBlockId="6.[136,1452,158,436]" lastPageId="6" lastPageNumber="255" pageId="4" pageNumber="253">
<emphasis id="59D9EAACFF9D9059FF57F9A5FE85F95B" bold="true" box="[189,325,1644,1670]" pageId="4" pageNumber="253">Discussion:</emphasis>
— 
<taxonomicName id="ACAD4D3DFF9D9059FE8BF9A5FDAFF95B" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[353,623,1644,1670]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9D9059FE8BF9A5FDAFF95B" box="[353,623,1644,1670]" italics="true" pageId="4" pageNumber="253">Eremanthus tomentosus</emphasis>
</taxonomicName>
belongs to 
<emphasis id="59D9EAACFF9D9059FD16F9A4FCD4F95B" box="[764,788,1645,1670]" italics="true" pageId="4" pageNumber="253">E.</emphasis>
subgen. 
<taxonomicName id="ACAD4D3DFF9D9059FC95F9A5FF7BF976" authority="MacLeish (1987: 275)" authorityName="MacLeish" authorityPageNumber="275" authorityYear="1987" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="section" section="Synglomerus">
<emphasis id="59D9EAACFF9D9059FC95F9A5FBC5F95B" box="[895,1029,1644,1670]" italics="true" pageId="4" pageNumber="253">Eremanthus</emphasis>
sect. 
<emphasis id="59D9EAACFF9D9059FBA7F9A5FB1EF95B" box="[1101,1246,1644,1670]" italics="true" pageId="4" pageNumber="253">Synglomerus</emphasis>
<bibRefCitation id="0F3C4B4FFF9D9059FB02F9A5FF7BF976" author="MacLeish, N. F. F." pageId="4" pageNumber="253" pagination="265 - 290" refId="ref9021" refString="MacLeish, N. F. F. (1987) Revision of Eremanthus (Compositae: Vernonieae). Annals of Missouri Botanical Garden 74: 265 - 290. https: // doi. org / 10.2307 / 2399398" type="journal article" year="1987">MacLeish (1987: 275)</bibRefCitation>
</taxonomicName>
because of the heads organized in a cyme of syncephalia (not a compound cyme of syncephalia like in sect. 
<taxonomicName id="ACAD4D3DFF9D9059FF62F97DFECEF913" authorityName="Lessing" authorityYear="1829" box="[136,270,1716,1742]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="genus">
<emphasis id="59D9EAACFF9D9059FF62F97DFECEF913" box="[136,270,1716,1742]" italics="true" pageId="4" pageNumber="253">Eremanthus</emphasis>
</taxonomicName>
), obconic heads and cypsela with a paleaceous, persistent pappus (
<bibRefCitation id="0F3C4B4FFF9D9059FC13F97DFB6EF913" author="MacLeish, N. F. F." box="[1017,1198,1716,1742]" pageId="4" pageNumber="253" pagination="265 - 290" refId="ref9021" refString="MacLeish, N. F. F. (1987) Revision of Eremanthus (Compositae: Vernonieae). Annals of Missouri Botanical Garden 74: 265 - 290. https: // doi. org / 10.2307 / 2399398" type="journal article" year="1987">MacLeish 1987</bibRefCitation>
). It is morphologically related to 
<taxonomicName id="ACAD4D3DFF9D9059FEE9F910FD9BF92E" authority="Baker (1873: 162)" authorityName="Baker" authorityPageNumber="162" authorityYear="1873" box="[259,603,1752,1779]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="cinctus">
<emphasis id="59D9EAACFF9D9059FEE9F910FEB6F92F" box="[259,374,1753,1778]" italics="true" pageId="4" pageNumber="253">E. cinctus</emphasis>
<bibRefCitation id="0F3C4B4FFF9D9059FE69F911FD9BF92E" author="Baker, J. G." box="[387,603,1752,1779]" pageId="4" pageNumber="253" pagination="5 - 180" refId="ref7707" refString="Baker, J. G. (1873) Compositae I. Vernoniaceae. In: Martius †, C. F. P. von &amp; Eichler, A. W. (Eds.) Flora brasiliensis, vol. 6 (2). Fried. Fleischer, Munich, Vienna, Leipzig, pp. 5 - 180." type="book chapter" year="1873">Baker (1873: 162)</bibRefCitation>
</taxonomicName>
by its reddish-brown tomentose stems and hemispherical syncephalium surrounded by conspicuous secondary bracts. However, the new species differs from 
<taxonomicName id="ACAD4D3DFF9D9059FBA9F934FB72F8CB" authorityName="Baker" authorityYear="1873" box="[1091,1202,1789,1814]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="cinctus">
<emphasis id="59D9EAACFF9D9059FBA9F934FB72F8CB" box="[1091,1202,1789,1814]" italics="true" pageId="4" pageNumber="253">E. cinctus</emphasis>
</taxonomicName>
by its petiolate leaves (vs. sessile), the adaxial leaf surface tomentum (tomentose vs. glabrate), secondary bracts with constricted obtuse bases (vs. prominently enlarged bases), heads fused in 3/4 of length (vs. fused in the entire length) and cypsela with reddish glands at the base (vs. without reddish glands at the base). The other 
<taxonomicName id="ACAD4D3DFF9D9059FC4FF8A1FB45F85F" box="[933,1157,1896,1922]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="59D9EAACFF9D9059FC4FF8A1FBEBF85F" box="[933,1067,1896,1922]" italics="true" pageId="4" pageNumber="253">Eremanthus</emphasis>
species
</taxonomicName>
with adaxially tomentose leaves is 
<taxonomicName id="ACAD4D3DFF9D9059FF1BF844FD05F87A" authority="Schultz-Bipontinus (1861: 166)" authorityName="Schultz-Bipontinus" authorityPageNumber="166" authorityYear="1861" box="[241,709,1932,1959]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="59D9EAACFF9D9059FF1BF844FE95F87B" box="[241,341,1932,1958]" italics="true" pageId="4" pageNumber="253">E. mollis</emphasis>
<bibRefCitation id="0F3C4B4FFF9D9059FEB7F845FD05F87A" author="Schultz-Bipontinus, C. H." box="[349,709,1932,1959]" pageId="4" pageNumber="253" pagination="157 - 190" refId="ref10563" refString="Schultz-Bipontinus, C. H. (1861) Cassiniaceae uniflorae, oder Verzeichniss der Cassiniaceen mit 1 - bluthigen Kopfchen. Jahresbericht der Pollichia, eines Naturwissenschaftlichen Vereins der bayerischen Pfalz 18 / 19: 157 - 190." type="journal article" year="1861">Schultz-Bipontinus (1861: 166)</bibRefCitation>
</taxonomicName>
, which also shares with 
<taxonomicName id="ACAD4D3DFF9D9059FC30F844FBB8F87B" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[986,1144,1933,1958]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9D9059FC30F844FBB8F87B" box="[986,1144,1933,1958]" italics="true" pageId="4" pageNumber="253">E. tomentosus</emphasis>
</taxonomicName>
the presence of third order syncephalia in some individuals. The new species differs by its habit (treelet vs. subshrub to shrub), inflorescence 
<typeStatus id="B416881CFF9D9059FF62F81CFF78F832" box="[136,184,2005,2031]" pageId="4" pageNumber="253">type</typeStatus>
(cyme of syncephalia vs. corymb of syncephalia), syncephalium shape (hemispherical vs. spherical), number of pappus series (3–4-seriate vs. biseriate). 
<taxonomicName id="ACAD4D3DFF9D9059FDB6F831FCAAF7CF" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[604,874,2040,2066]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="253" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9D9059FDB6F831FCAAF7CF" box="[604,874,2040,2066]" italics="true" pageId="4" pageNumber="253">Eremanthus tomentosus</emphasis>
</taxonomicName>
occurs in sympatry with four other species of the genus: 
<taxonomicName id="ACAD4D3DFF9F905BFF32FF56FEFCFF65" authorityName="Schultz-Bipontinus" authorityYear="1861" box="[216,316,158,184]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="59D9EAACFF9F905BFF32FF56FEFCFF65" box="[216,316,158,184]" italics="true" pageId="6" pageNumber="255">E. mollis</emphasis>
</taxonomicName>
, 
<taxonomicName id="ACAD4D3DFF9F905BFEA0FF56FD29FF65" authority="Lessing (1829: 317)" authorityName="Lessing" authorityPageNumber="317" authorityYear="1829" box="[330,745,158,184]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="glomerulatus">
<emphasis id="59D9EAACFF9F905BFEA0FF56FE3EFF65" box="[330,510,158,184]" italics="true" pageId="6" pageNumber="255">E. glomerulatus</emphasis>
<bibRefCitation id="0F3C4B4FFF9F905BFDEFFF57FD29FF65" author="Lessing, C. F." box="[517,745,158,184]" pageId="6" pageNumber="255" pagination="240 - 356" refId="ref8842" refString="Lessing, C. F. (1829) De synanthereis herbarii regii berolinensis dissertatio prima. Vernonieae. Linnaea 4: 240 - 356." type="journal article" year="1829">Lessing (1829: 317)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="ACAD4D3DFF9F905BFD1FFF56FF7EFF00" authority="(Gardner 1847: 425) Schultz-Bipontinus (1861: 165)" authorityName="Schultz-Bipontinus" authorityPageNumber="165" authorityYear="1861" baseAuthorityName="Gardner" baseAuthorityPageNumber="425" baseAuthorityYear="1847" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="goyazensis">
<emphasis id="59D9EAACFF9F905BFD1FFF56FC4DFF65" box="[757,909,159,184]" italics="true" pageId="6" pageNumber="255">E. goyazensis</emphasis>
(
<bibRefCitation id="0F3C4B4FFF9F905BFC76FF57FBB4FF64" author="Gardner, G." box="[924,1140,158,185]" pageId="6" pageNumber="255" pagination="417 - 449" refId="ref8362" refString="Gardner, G. (1847) Contributions towards a Flora of Brazil, being the characters of several new species of Compositae, belonging to the tribes Vernoniaceae and Eupatoriaceae, from the Province of Goyaz. The London Journal of Botany 6: 417 - 449." type="journal article" year="1847">Gardner 1847: 425</bibRefCitation>
) 
<bibRefCitation id="0F3C4B4FFF9F905BFB69FF57FF7EFF00" author="Schultz-Bipontinus, C. H." pageId="6" pageNumber="255" pagination="157 - 190" refId="ref10563" refString="Schultz-Bipontinus, C. H. (1861) Cassiniaceae uniflorae, oder Verzeichniss der Cassiniaceen mit 1 - bluthigen Kopfchen. Jahresbericht der Pollichia, eines Naturwissenschaftlichen Vereins der bayerischen Pfalz 18 / 19: 157 - 190." type="journal article" year="1861">Schultz-Bipontinus (1861: 165)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="ACAD4D3DFF9F905BFF13FF0AFD71FF00" authority="Kuntze (1898: 145)" authorityName="Kuntze" authorityPageNumber="145" authorityYear="1898" box="[249,689,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="mattogrossensis">
<emphasis id="59D9EAACFF9F905BFF13FF0AFE0BFF01" box="[249,459,195,220]" italics="true" pageId="6" pageNumber="255">E. mattogrossensis</emphasis>
<bibRefCitation id="0F3C4B4FFF9F905BFE39FF0AFD71FF00" author="Kuntze, C. E. O." box="[467,689,194,221]" pageId="6" pageNumber="255" refId="ref8812" refString="Kuntze, C. E. O. (1898) Revisio Generum Plantarum, vol. 3, pars 3. Arthur Felix, Leipzig, 576 pp." type="book" year="1898">Kuntze (1898: 145)</bibRefCitation>
</taxonomicName>
, but these species have compound cymes of syncephalia with heads fused in less than 1/2 of the length (
<taxonomicName id="ACAD4D3DFF9F905BFDCAFF2EFD14FEDD" authorityName="Lessing" authorityYear="1829" box="[544,724,230,256]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="glomerulatus">
<emphasis id="59D9EAACFF9F905BFDCAFF2EFD14FEDD" box="[544,724,230,256]" italics="true" pageId="6" pageNumber="255">E. glomerulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="ACAD4D3DFF9F905BFCE1FF2EFC1EFEDD" authorityName="Kuntze" authorityYear="1898" box="[779,990,231,256]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="mattogrossensis">
<emphasis id="59D9EAACFF9F905BFCE1FF2EFC1EFEDD" box="[779,990,231,256]" italics="true" pageId="6" pageNumber="255">E. mattogrossensis</emphasis>
</taxonomicName>
) or the entire length (
<taxonomicName id="ACAD4D3DFF9F905BFB32FF2EFAB1FEDD" authorityName="Schultz-Bipontinus" authorityYear="1861" baseAuthorityName="Gardner" baseAuthorityYear="1847" box="[1240,1393,231,256]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="goyazensis">
<emphasis id="59D9EAACFF9F905BFB32FF2EFAB1FEDD" box="[1240,1393,231,256]" italics="true" pageId="6" pageNumber="255">E. goyazensis</emphasis>
</taxonomicName>
) (vs. not compound cyme and heads fused in 3/4 of the length) (
<bibRefCitation id="0F3C4B4FFF9F905BFCC0FEC3FC1FFEF9" author="MacLeish, N. F. F." box="[810,991,266,292]" pageId="6" pageNumber="255" pagination="133 - 136" refId="ref8981" refString="MacLeish, N. F. F. (1984) Eight new combinations in Vernonia (Compositae: Vernonieae). Systematic Botany 9: 133 - 136. https: // doi. org / 10.2307 / 2418817" type="journal article" year="1984">MacLeish 1984</bibRefCitation>
, 
<bibRefCitation id="0F3C4B4FFF9F905BFC01FEC3FBE6FEF9" author="MacLeish, N. F. F." box="[1003,1062,266,292]" pageId="6" pageNumber="255" pagination="265 - 290" refId="ref9021" refString="MacLeish, N. F. F. (1987) Revision of Eremanthus (Compositae: Vernonieae). Annals of Missouri Botanical Garden 74: 265 - 290. https: // doi. org / 10.2307 / 2399398" type="journal article" year="1987">1987</bibRefCitation>
, 
<bibRefCitation id="0F3C4B4FFF9F905BFBD9FEC3FAD4FEF9" author="Loeuille, B. &amp; Semir, J. &amp; Pirani, J. R." box="[1075,1300,266,292]" pageId="6" pageNumber="255" pagination="1 - 139" refId="ref8931" refString="Loeuille, B., Semir, J. &amp; Pirani, J. R. (2019) A synopsis of the subtribe Lychnophorinae (Asteraceae: Vernonieae). Phytotaxa 398: 1 - 139. https: // doi. org / 10.11646 / phytotaxa. 398.1.1" type="journal article" year="2019">
Loeuille 
<emphasis id="59D9EAACFF9F905BFB72FEC2FB12FEF9" box="[1176,1234,266,292]" italics="true" pageId="6" pageNumber="255">et al.</emphasis>
2019
</bibRefCitation>
) (
<tableCitation id="262F0305FF9F905BFAC1FEC3FA40FEF9" box="[1323,1408,266,292]" captionStart="TABLE 1" captionStartId="5.[136,224,737,763]" captionTargetPageId="5" captionText="TABLE 1. Diagnostic morphological characters of Eremanthus tomentosus and related species. Source material: E. cinctus – Aparecida da Silva et al. 3875 (IBGE, SP, US), Loeuille et al. 306 (HUFU, SPF), Schumacher 5005 (MB, NY); E. glomerulatus – Loeuille et al. 296 (K, MBM, NY, SPF, UB, UFG, US), Loeuille et al. 831 (SPF, US), Souza et al. 7890 (ESA, HUFU, K, SPF, UEC); E. mattogrossensis – Loeuille et al. 453 (SPF), Nave et al. 1258 (ESA, K, UFMT), Ratter et al. 1926 (NY, UB); E. mollis – Irwin et al. (NY, SP, US), Heringer et al. 6125 (IBGE, NY, US), Loeuille et al. 305 (K, MBM, SPF, UFG); as well as data from MacLeish (1987) and Loeuille et al. (2019)." httpUri="http://table.plazi.org/id/3FD26636FF9C9058FF62FD28F95BFCBA" pageId="6" pageNumber="255" tableUuid="3FD26636FF9C9058FF62FD28F95BFCBA">Table 1</tableCitation>
). A hybrid origin of the new species cannot be excluded since 
<taxonomicName id="ACAD4D3DFF9F905BFCFAFEE6FC68FE95" authorityName="Schultz-Bipontinus" authorityYear="1861" baseAuthorityName="Gardner" baseAuthorityYear="1847" box="[784,936,303,328]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="goyazensis">
<emphasis id="59D9EAACFF9F905BFCFAFEE6FC68FE95" box="[784,936,303,328]" italics="true" pageId="6" pageNumber="255">E. goyazensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="ACAD4D3DFF9F905BFC37FEE6FB80FE95" authorityName="Schultz-Bipontinus" authorityYear="1861" box="[989,1088,302,328]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="59D9EAACFF9F905BFC37FEE6FB80FE95" box="[989,1088,302,328]" italics="true" pageId="6" pageNumber="255">E. mollis</emphasis>
</taxonomicName>
share some morphological traits with the new species and the three of them were always found sympatrically. Nonetheless, only 
<taxonomicName id="ACAD4D3DFF9F905BFB47FE9AFA89FEB1" authorityName="Loeuille &amp; Bringel" authorityYear="2022" box="[1197,1353,339,364]" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="255" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="59D9EAACFF9F905BFB47FE9AFA89FEB1" box="[1197,1353,339,364]" italics="true" pageId="6" pageNumber="255">E. tomentosus</emphasis>
</taxonomicName>
displays corolla villose towards the base and cypsela with reddish glands at the base. We encourage future efforts to investigate this hypothesis.
</paragraph>
</subSubSection>
</treatment>
</document>