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<document id="E0C3654AE590A2A35DB7643505960710" ID-CLB-Dataset="51166" ID-DOI="10.11646/zootaxa.4066.5.4" ID-GBIF-Dataset="d36cd9ed-466d-4218-83b3-650be11cfc64" ID-ISSN="1175-5326" ID-Zenodo-Dep="255984" ID-ZooBank="F847C440-E9F4-4793-9959-8927336142E6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459691682967" checkinUser="plazi" docAuthor="Chainthong, Damrong &amp; Boonsoong, Boonsatien" docDate="2016" docId="CF2C87C0AD6702798181FF5CFBCFCFB2" docLanguage="en" docName="zootaxa.4066.5.4.pdf" docOrigin="Zootaxa 4066 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onychogomphus duaricus Fraser 1924" docType="treatment" docVersion="9" lastPageNumber="569" masterDocId="3315FFB8AD6202718116FFC4FF9ECE62" masterDocTitle="Description of two final stadium Onychogomphus larvae from Thailand (Odonata: Gomphidae)" masterLastPageNumber="570" masterPageNumber="561" pageNumber="566" updateTime="1698209179711" updateUser="ExternalLinkService">
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<mods:title id="37FA45BC31FF341FA1FEB3CD9E0525A8">Description of two final stadium Onychogomphus larvae from Thailand (Odonata: Gomphidae)</mods:title>
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<mods:namePart id="0E6E63A4C34C64E736E95342691B1A6E">Chainthong, Damrong</mods:namePart>
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<mods:namePart id="4E447D6F663D10B38E1FBAE620ECB943">Boonsoong, Boonsatien</mods:namePart>
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<treatment id="CF2C87C0AD6702798181FF5CFBCFCFB2" ID-DOI="http://doi.org/10.5281/zenodo.5660085" ID-GBIF-Taxon="119333458" ID-Zenodo-Dep="5660085" LSID="urn:lsid:plazi:treatment:CF2C87C0AD6702798181FF5CFBCFCFB2" httpUri="http://treatment.plazi.org/id/CF2C87C0AD6702798181FF5CFBCFCFB2" lastPageId="8" lastPageNumber="569" pageId="5" pageNumber="566">
<subSubSection id="0F9F655DAD6702748181FF5CFD0ACED0" box="[151,660,151,178]" pageId="5" pageNumber="566" type="nomenclature">
<paragraph id="473A36D6AD6702748181FF5CFD0ACED0" blockId="5.[151,660,151,211]" box="[151,660,151,178]" pageId="5" pageNumber="566">
<heading id="1C7281BAAD6702748181FF5CFD0ACED0" bold="true" box="[151,660,151,178]" fontSize="11" level="1" pageId="5" pageNumber="566" reason="1">
<emphasis id="75F1EAC4AD6702748181FF5CFD0ACED0" bold="true" box="[151,660,151,178]" pageId="5" pageNumber="566">
2.
<taxonomicName id="80854D55AD67027481A2FF53FD0ACED0" ID-CoL="49R99" authority="Fraser, 1924" authorityName="Fraser" authorityYear="1924" box="[180,660,151,178]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="duaricus">
<emphasis id="75F1EAC4AD67027481A2FF53FE6DCED3" bold="true" box="[180,499,151,177]" italics="true" pageId="5" pageNumber="566">Onychogomphus duaricus</emphasis>
Fraser, 1924
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0F9F655DAD6702748181FF7EFED4CEB1" box="[151,330,186,211]" pageId="5" pageNumber="566" type="description">
<paragraph id="473A36D6AD6702748181FF7EFED4CEB1" blockId="5.[151,660,151,211]" box="[151,330,186,211]" pageId="5" pageNumber="566">
(
<figureCitation id="DFBE2A53AD6702748189FF7EFF70CEB1" box="[159,238,186,211]" captionStart="FIGURES 1 2" captionStartId="1.[151,264,1584,1607]" captionTargetBox="[340,1246,339,1561]" captionTargetId="figure@1.[340,1247,339,1563]" captionTargetPageId="1" captionText="FIGURES 1 2. Onychogomphus castor Lieftinck and O. duaricus Fraser, respectively: habitus of larvae, dorsal view." httpUri="https://zenodo.org/record/255985/files/figure.png" pageId="5" pageNumber="566">Figs. 2</figureCitation>
,
<figureCitation id="DFBE2A53AD67027481ECFF7FFEDCCEB1" box="[250,322,187,211]" captionStart-0="FIGURE 13" captionStart-1="FIGURES 14 19" captionStart-2="FIGURES 20 22" captionStartId-0="5.[151,250,1981,2003]" captionStartId-1="6.[151,264,1374,1397]" captionStartId-2="7.[151,264,1529,1552]" captionTargetBox-1="[167,1427,207,1346]" captionTargetBox-2="[165,1419,201,1501]" captionTargetId-1="figure@6.[151,1436,193,1353]" captionTargetId-2="figure@7.[151,1435,193,1508]" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 13. Onychogomphus duaricus Fraser: exuvia, dorsal view. Scale = 1 mm." captionText-1="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." captionText-2="FIGURES 20 22. Onychogomphus duaricus Fraser: (20); right side of abdomen of exuvia, dorsal view; (21) lateral view of abdomen in exuvia; (22) ventral view of abdomen. Scale = 1 mm." httpUri-1="https://zenodo.org/record/255988/files/figure.png" httpUri-2="https://zenodo.org/record/255989/files/figure.png" pageId="5" pageNumber="566">1322</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0F9F655DAD6702748181FEC6FD90CC75" pageId="5" pageNumber="566" type="materials_examined">
<paragraph id="473A36D6AD6702748181FEC6FB4BCFAD" blockId="5.[151,1436,258,643]" pageId="5" pageNumber="566">
<emphasis id="75F1EAC4AD6702748181FEC6FE45CF79" bold="true" box="[151,475,258,283]" pageId="5" pageNumber="566">Material examined (4 F0, 2</emphasis>
♀,
<specimenCount id="5183FD5FAD6702748313FEC6FDB1CF78" box="[517,559,258,283]" pageId="5" pageNumber="566" type="generic">
<emphasis id="75F1EAC4AD6702748313FEC6FD8DCF79" bold="true" box="[517,531,258,283]" pageId="5" pageNumber="566">2</emphasis>
</specimenCount>
<emphasis id="75F1EAC4AD6702748339FEC6FDDCCF79" bold="true" box="[559,578,258,283]" pageId="5" pageNumber="566">):</emphasis>
1 F0 larva,
<date id="333B1016AD67027483DCFEC7FCD4CF79" box="[714,842,259,283]" pageId="5" pageNumber="566" value="2013-03-30">30.III.2013</date>
, Kang Som Meaw (13°2428″
<geoCoordinate id="22B15011AD670274858DFEC6FB61CF79" box="[1179,1279,258,283]" direction="north" orientation="latitude" pageId="5" pageNumber="566" precision="925" value="99.26667">N 99°16</geoCoordinate>
52″E, altitude
<quantity id="807D9B33AD6702748181FEE3FF7FCF5D" box="[151,225,295,319]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="566" unit="m" value="200.0">200 m</quantity>
), Suanpung district, Ratchaburi province, D. Chainthong leg.; 1 ♀ and its exuvia, emerged on
<date id="333B1016AD6702748412FEE2FA1ACF5D" box="[1284,1412,294,319]" pageId="5" pageNumber="566" value="2013-04-26">26.IV.2013</date>
. 2 F0 larvae,
<date id="333B1016AD6702748009FE8FFE17CF01" box="[287,393,331,355]" pageId="5" pageNumber="566" value="2014-02-04">4.II.2014</date>
, Huai Khayeng (14°3620″
<geoCoordinate id="22B15011AD67027483C7FE8EFCDECF01" box="[721,832,330,355]" direction="north" orientation="latitude" pageId="5" pageNumber="566" precision="925" value="98.566666">N 98°34</geoCoordinate>
38″E, altitude
<quantity id="807D9B33AD67027482E5FE8EFBD8CF01" box="[1011,1094,330,355]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="5" pageNumber="566" unit="m" value="206.0">206 m</quantity>
), Thong Pha Phum district, Kanchanaburi province, D. Chainthong leg.; 1 ♀ and its exuvia, emerged on
<date id="333B1016AD6702748553FEABFB24CFE5" box="[1093,1210,367,391]" pageId="5" pageNumber="566" value="2014-05-18">18.V.2014</date>
;
<specimenCount id="5183FD5FAD67027485DBFEABFB61CFE4" box="[1229,1279,367,391]" pageId="5" pageNumber="566" type="generic">1 ♂</specimenCount>
, emerged on
<date id="333B1016AD6702748181FE57FE93CFC9" box="[151,269,403,427]" pageId="5" pageNumber="566" value="2014-05-20">20.V.2014</date>
. 1 F0 larva,
<date id="333B1016AD67027480B7FE57FDB3CFC9" box="[417,557,403,427]" pageId="5" pageNumber="566" value="2014-12-14">14.XII.2014</date>
, Huai Pak Kok (14°3934″
<geoCoordinate id="22B15011AD6702748277FE56FC57CFC9" box="[865,969,402,427]" direction="north" orientation="latitude" pageId="5" pageNumber="566" precision="925" value="98.53333">N 98°32</geoCoordinate>
02″E, altitude
<quantity id="807D9B33AD6702748567FE56FB20CFC9" box="[1137,1214,402,427]" metricMagnitude="2" metricUnit="m" metricValue="1.76" pageId="5" pageNumber="566" unit="m" value="176.0">176 m</quantity>
), Thong Pha Phum district, Kanchanaburi province, D. Chainthong leg.;
<specimenCount id="5183FD5FAD67027483FDFE73FC8BCFAC" box="[747,789,439,463]" pageId="5" pageNumber="566" type="generic">1 ♂</specimenCount>
and its exuvia, emerged on
<date id="333B1016AD6702748559FE72FB4ECFAD" box="[1103,1232,438,463]" pageId="5" pageNumber="566" value="2015-03-23">23.III.2015</date>
.
</paragraph>
<paragraph id="473A36D6AD67027481D1FE1EFD90CC75" blockId="5.[151,1436,258,643]" pageId="5" pageNumber="566">
<emphasis id="75F1EAC4AD67027481D1FE1EFEDCCF91" box="[199,322,474,499]" italics="true" pageId="5" pageNumber="566">Coloration</emphasis>
: Larvae uniformly dark brown, body stout and bulky, integument finely granulose, body covered with short hair-like setae (
<figureCitation id="DFBE2A53AD67027480ADFE3AFD9ECC75" box="[443,512,510,535]" captionStart="FIGURES 1 2" captionStartId="1.[151,264,1584,1607]" captionTargetBox="[340,1246,339,1561]" captionTargetId="figure@1.[340,1247,339,1563]" captionTargetPageId="1" captionText="FIGURES 1 2. Onychogomphus castor Lieftinck and O. duaricus Fraser, respectively: habitus of larvae, dorsal view." httpUri="https://zenodo.org/record/255985/files/figure.png" pageId="5" pageNumber="566">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0F9F655DAD67027981D1FDE0FBCFCFB2" lastPageId="8" lastPageNumber="569" pageId="5" pageNumber="566" type="description">
<paragraph id="473A36D6AD67027481D1FDE0FDE2CCE0" blockId="5.[151,1436,258,643]" pageId="5" pageNumber="566">
<emphasis id="75F1EAC4AD67027481D1FDE0FE2ACC59" box="[199,436,547,571]" italics="true" pageId="5" pageNumber="566">Measurements (n=4)</emphasis>
: Total body length
<quantity id="807D9B33AD6702748381FDE7FD61CC58" box="[663,767,547,571]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="5" pageNumber="566" unit="mm" value="23.9">23.9 mm</quantity>
, length of abdomen
<quantity id="807D9B33AD67027482FAFDE7FBCACC58" box="[1004,1108,547,571]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="5" pageNumber="566" unit="mm" value="13.4">13.4 mm</quantity>
, abdominal maximum width
<quantity id="807D9B33AD6702748181FD82FF6DCC3C" box="[151,243,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="566" unit="mm" value="5.7">5.7 mm</quantity>
, head maximum width
<quantity id="807D9B33AD6702748318FD82FDF5CC3C" box="[526,619,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="566" unit="mm" value="4.6">4.6 mm</quantity>
, length of hind femur
<quantity id="807D9B33AD670274826AFD83FC47CC3C" box="[892,985,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="566" unit="mm" value="3.3">3.3 mm</quantity>
, length of epiproct
<quantity id="807D9B33AD67027485D0FD82FABDCC3C" box="[1222,1315,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="566" unit="mm" value="1.5">1.5 mm</quantity>
, length of paraprocts
<quantity id="807D9B33AD6702748004FDAEFEF5CCE0" box="[274,363,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="566" unit="mm" value="1.6">1.6 mm</quantity>
, length of cerci
<quantity id="807D9B33AD6702748309FDAFFDE6CCE0" box="[543,632,619,643]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="566" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<caption id="13FA665EAD6702748181F879FC7BC9B1" box="[151,997,1981,2003]" pageId="5" pageNumber="566">
<paragraph id="473A36D6AD6702748181F879FC7BC9B1" blockId="5.[151,997,1981,2003]" box="[151,997,1981,2003]" pageId="5" pageNumber="566">
<emphasis id="75F1EAC4AD6702748181F879FE81C9B0" bold="true" box="[151,287,1981,2003]" pageId="5" pageNumber="566">FIGURE 13.</emphasis>
<taxonomicName id="80854D55AD6702748033F879FDE6C9B1" authority="Fraser" authorityName="Fraser" box="[293,632,1981,2003]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="duaricus">
<emphasis id="75F1EAC4AD6702748033F879FDB1C9B1" box="[293,559,1981,2003]" italics="true" pageId="5" pageNumber="566">Onychogomphus duaricus</emphasis>
Fraser
</taxonomicName>
: exuvia, dorsal view. Scale = 1 mm.
</paragraph>
</caption>
<caption id="13FA665EAD6402778181FA9AFC1CCBD0" httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="6" pageNumber="567" targetBox="[167,1427,207,1346]" targetPageId="6">
<paragraph id="473A36D6AD6402778181FA9AFC1CCBD0" blockId="6.[151,1436,1374,1458]" pageId="6" pageNumber="567">
<emphasis id="75F1EAC4AD6402778181FA9AFECDCB16" bold="true" box="[151,339,1374,1397]" pageId="6" pageNumber="567">FIGURES 1419.</emphasis>
<taxonomicName id="80854D55AD640277804FFA9AFD34CB16" authority="Fraser" authorityName="Fraser" box="[345,682,1374,1396]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="duaricus">
<emphasis id="75F1EAC4AD640277804FFA9AFDFCCB16" box="[345,610,1374,1396]" italics="true" pageId="6" pageNumber="567">Onychogomphus duaricus</emphasis>
Fraser
</taxonomicName>
: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17).
</paragraph>
</caption>
<paragraph id="473A36D6AD64027781D1FA1DFECDC9A4" blockId="6.[151,1437,1497,1990]" pageId="6" pageNumber="567">
<emphasis id="75F1EAC4AD64027781D1FA1DFE9ACB90" box="[199,260,1497,1522]" italics="true" pageId="6" pageNumber="567">Head</emphasis>
: wider than long, wider than prothorax but narrower than synthorax, frontal part stouter, posterior lobe of head shorter than eye length, eyes large and broadest across, with three large ocelli. Antennae four-segmented, first two segments short and rather circular, third segment largest, twice as long as first two segments and weakly swollen externally (length
<quantity id="807D9B33AD64027780D0F982FDBCC83C" box="[454,546,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="567" unit="mm" value="1.4">1.4 mm</quantity>
), fourth segment tiny (length
<quantity id="807D9B33AD6402778262F982FC51C83C" box="[884,975,1606,1630]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="567" unit="mm" value="0.1">0.1 mm</quantity>
); all segments conspicuously spinulated, without setae (
<figureCitation id="DFBE2A53AD640277802DF9ADFE10C8E0" box="[315,398,1641,1666]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="6" pageNumber="567">Fig. 14</figureCitation>
). Mandibles as in
<figureCitation id="DFBE2A53AD640277834CF9ADFD41C8E0" box="[602,735,1641,1666]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="6" pageNumber="567">Figs. 1516</figureCitation>
with formula: L 1234 0 a(m18 or m110)b / R 1234 y a(m1 6 or m18)b with a&gt;b in both mandibles. Maxilla: galeolacinia with seven moderately incurved teeth, three dorsal teeth nearly equal in length and robustness, three ventral teeth of same sizes, apical one largest; stipes and palp very strongly setose (
<figureCitation id="DFBE2A53AD640277804BF911FE29C88C" box="[349,439,1749,1774]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="6" pageNumber="567">Fig. 17</figureCitation>
). Labium: flat and not protruding when at rest (
<figureCitation id="DFBE2A53AD6402778506F911FBF5C88C" box="[1040,1131,1749,1774]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="6" pageNumber="567">Fig. 18</figureCitation>
). Prementum-postmentum articulation reaching posterior margin of procoxae. Prementum subrectangular, longer than wide (width 2.3, length 3.4), sides convex, convergent basally, with small teeth and minute setae at lateral margins; apical margin convex, with ventral row of 2931 short, subquadrate reddish brown teeth and dorsal rows of whitish piliform setae on apical border; labial palp with uniformly inflexed inner edge, yellowish brown, apical lobe reddish, rounded, its internal margin with row of 1516 equidistant teeth. Movable hook reddish brown, sharp and moderately incurved (length
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).
</paragraph>
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<paragraph id="473A36D6AD6502768181FA3DFCAFC84C" blockId="7.[151,1436,1529,1582]" pageId="7" pageNumber="568">
<emphasis id="75F1EAC4AD6502768181FA3DFECBC86C" bold="true" box="[151,341,1529,1552]" pageId="7" pageNumber="568">FIGURES 2022.</emphasis>
<taxonomicName id="80854D55AD650276804AFA3DFD2CC86D" authority="Fraser" authorityName="Fraser" box="[348,690,1529,1551]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="568" phylum="Arthropoda" rank="species" species="duaricus">
<emphasis id="75F1EAC4AD650276804AFA3DFDF6C86D" box="[348,616,1529,1551]" italics="true" pageId="7" pageNumber="568">Onychogomphus duaricus</emphasis>
Fraser
</taxonomicName>
: (20); right side of abdomen of exuvia, dorsal view; (21) lateral view of abdomen in exuvia; (22) ventral view of abdomen. Scale = 1 mm.
</paragraph>
</caption>
<paragraph id="473A36D6AD65027681D1F99EFD75C945" blockId="7.[151,1436,1626,2011]" pageId="7" pageNumber="568">
<emphasis id="75F1EAC4AD65027681D1F99EFE86C811" box="[199,280,1626,1651]" italics="true" pageId="7" pageNumber="568">Thorax</emphasis>
: Small, suboval, prothorax narrower than head, dorsal portion raised at sides forming two U-shaped ridges. Wing sheaths strongly divergent, reaching middle of S4 (inner wing pads length
<quantity id="807D9B33AD6502768596F9BAFB44C8F5" box="[1152,1242,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="7" pageNumber="568" unit="mm" value="5.3">5.3 mm</quantity>
, outer wing pads length
<quantity id="807D9B33AD65027681F3F966FEDFC8D9" box="[229,321,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="568" unit="mm" value="4.5">4.5 mm</quantity>
). Legs short and stout; fore and middle legs strongly curved; protibiae (length
<quantity id="807D9B33AD65027685D0F967FA81C8D9" box="[1222,1311,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="568" unit="mm" value="1.9">1.9 mm</quantity>
) decidedly longer than profemora (length
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). Mesotibiae (length
<quantity id="807D9B33AD6502768257F903FC04C8BD" box="[833,922,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="568" unit="mm" value="1.7">1.7 mm</quantity>
) slightly longer than mesofemora (length
<quantity id="807D9B33AD650276846FF902FF59C961" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="568" unit="mm" value="1.6">1.6 mm</quantity>
); metafemur slightly longer than metatibia. Tarsal formula 2-2-3, tarsi yellowish. Minute short rows of setae scattered along femur, tibia, and tarsus of all six legs.
</paragraph>
<paragraph id="473A36D6AD65027981D1F8F6FE2ACE9A" blockId="7.[151,1436,1626,2011]" lastBlockId="8.[151,1437,151,464]" lastPageId="8" lastPageNumber="569" pageId="7" pageNumber="568">
<emphasis id="75F1EAC4AD65027681D1F8F6FEAFC929" box="[199,305,1842,1867]" italics="true" pageId="7" pageNumber="568">Abdomen</emphasis>
: less convex dorsally, dark brown, without markings (
<figureCitation id="DFBE2A53AD650276828AF8F6FC6EC929" box="[924,1008,1842,1867]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="7" pageNumber="568">Fig. 19</figureCitation>
). Mid-dorsal spines absent on S1 and short, blunt spines present on S29 (
<figureCitation id="DFBE2A53AD650276832EF892FD10C90D" box="[568,654,1878,1903]" captionStart="FIGURES 20 22" captionStartId="7.[151,264,1529,1552]" captionTargetBox="[165,1419,201,1501]" captionTargetId="figure@7.[151,1435,193,1508]" captionTargetPageId="7" captionText="FIGURES 20 22. Onychogomphus duaricus Fraser: (20); right side of abdomen of exuvia, dorsal view; (21) lateral view of abdomen in exuvia; (22) ventral view of abdomen. Scale = 1 mm." httpUri="https://zenodo.org/record/255989/files/figure.png" pageId="7" pageNumber="568">Fig. 20</figureCitation>
). Lacking lateral spines on S26. Lateral edges of abdomen serrated with projecting spines on S79 (
<figureCitation id="DFBE2A53AD6502768307F8BEFD00C9F1" box="[529,670,1914,1939]" captionStart="FIGURES 20 22" captionStartId="7.[151,264,1529,1552]" captionTargetBox="[165,1419,201,1501]" captionTargetId="figure@7.[151,1435,193,1508]" captionTargetPageId="7" captionText="FIGURES 20 22. Onychogomphus duaricus Fraser: (20); right side of abdomen of exuvia, dorsal view; (21) lateral view of abdomen in exuvia; (22) ventral view of abdomen. Scale = 1 mm." httpUri="https://zenodo.org/record/255989/files/figure.png" pageId="7" pageNumber="568">Figs. 2021</figureCitation>
). Sternites yellow brown, with few scattered long setae, sternite 8 divided in 5 plates, remainder in 3 plates, median sutures of sternites slightly convergent apically on 2, parallel on 37 and gradually divergent on 89, posterior margin of sternites 19 straight, wavy on 10 (
<figureCitation id="DFBE2A53AD65027685A8F806FA8BC9B9" box="[1214,1301,1986,2011]" captionStart="FIGURES 20 22" captionStartId="7.[151,264,1529,1552]" captionTargetBox="[165,1419,201,1501]" captionTargetId="figure@7.[151,1435,193,1508]" captionTargetPageId="7" captionText="FIGURES 20 22. Onychogomphus duaricus Fraser: (20); right side of abdomen of exuvia, dorsal view; (21) lateral view of abdomen in exuvia; (22) ventral view of abdomen. Scale = 1 mm." httpUri="https://zenodo.org/record/255989/files/figure.png" pageId="7" pageNumber="568">Fig. 22</figureCitation>
). No lateral spine and no dorsal hook on S10. Anal appendages elongated. Epiproct same length as paraproct, cerci slightly shorter. Epiproct narrowly triangular with two black spots close to the tip, conical. Paraprocts and cerci with tips sharply pointed (
<figureCitation id="DFBE2A53AD6A02798042FF1BFE39CE9A" box="[340,423,223,248]" captionStart="FIGURES 14 19" captionStartId="6.[151,264,1374,1397]" captionTargetBox="[167,1427,207,1346]" captionTargetId="figure@6.[151,1436,193,1353]" captionTargetPageId="6" captionText="FIGURES 14 19. Onychogomphus duaricus Fraser: (14) antenna, dorsal view; (15) right mandible, ventral and internal views; (16) left mandible, ventral and internal views; (17) right maxilla, ventral view; (18) ventral view of prementum; (19) dorsal view of anal appendages. Scale = 1 mm (14, 18, 19), 0.5 mm (15, 16, 17)." httpUri="https://zenodo.org/record/255988/files/figure.png" pageId="8" pageNumber="569">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="473A36D6AD6A027981D1FEC0FBCFCFB2" blockId="8.[151,1437,151,464]" pageId="8" pageNumber="569">
<emphasis id="75F1EAC4AD6A027981D1FEC0FE14CF7F" bold="true" box="[199,394,260,285]" pageId="8" pageNumber="569">Biological notes.</emphasis>
These two larvae of
<taxonomicName id="80854D55AD6A02798361FEC0FCADCF7F" box="[631,819,260,285]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="75F1EAC4AD6A02798361FEC0FCADCF7F" box="[631,819,260,285]" italics="true" pageId="8" pageNumber="569">Onychogomphus</emphasis>
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occur in headwater streams, where they were found in the sandy gravel substrate. On each sampling date, physico-chemical parameters were measured. During the odonate sampling, physico-chemical parameters varied according to time. The streams ranged from 4.0
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wide and
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in depth. The water temperature ranged from 21.024.0 °C,
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from 7.17.5, dissolved oxygen from
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/L, total dissolved solids from 6.320.0 mg/L, and conductivity from 90.3297.0 µS/cm. These variations correlated of water chemistry generally followed the seasonal cycle.
</paragraph>
</subSubSection>
</treatment>
</document>