159 lines
20 KiB
XML
159 lines
20 KiB
XML
<document id="9C71C832BD35D2529A537DC893A1EF1E" ID-DOI="10.5281/zenodo.157708" ID-GBIF-Dataset="aabec420-0763-4ca5-b4bb-e0528c17f814" ID-ISSN="11755326" ID-Zenodo-Dep="157708" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461018528122" checkinUser="plazi" docAuthor="Rafael, José Albertino & Ale-Rocha, Rosaly" docDate="2004" docId="C779878C9A46222EFECECCBEC87DFCD5" docLanguage="en" docName="zt00529.pdf" docOrigin="Zootaxa 529" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Elmohardyia nicaraguensis Rafael, n. sp." docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="3B40FFF49A45222BFFC6CF5FCC45FFC3" masterDocTitle="Nicaraguan Pipunculidae (Diptera): new records and description of new species" masterLastPageNumber="18" masterPageNumber="1" pageNumber="4" updateTime="1698569576216" updateUser="plazi">
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<mods:title id="4B2E029BCC842FAC54956D96D4441DB3">Nicaraguan Pipunculidae (Diptera): new records and description of new species</mods:title>
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<mods:namePart id="335CF0741C02A4F30D3DFF75EACC45B8">Rafael, José Albertino</mods:namePart>
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<mods:namePart id="EB417988E0DE7EE5EAB36FE5C94F31D4">Ale-Rocha, Rosaly</mods:namePart>
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<mods:title id="B8584C024DDD90478DB0E3836AC8F469">Zootaxa</mods:title>
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<mods:date id="C0430B8DFA341504CAD00F53A7A87AA1">2004</mods:date>
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<mods:number id="859512A5CFC0CC7F9B77A88399EF2340">529</mods:number>
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<treatment id="C779878C9A46222EFECECCBEC87DFCD5" ID-DOI="http://doi.org/10.5281/zenodo.6269496" ID-GBIF-Taxon="119602145" ID-Zenodo-Dep="6269496" LSID="urn:lsid:plazi:treatment:C779878C9A46222EFECECCBEC87DFCD5" httpUri="http://treatment.plazi.org/id/C779878C9A46222EFECECCBEC87DFCD5" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<subSubSection id="07CA65119A462228FECECCBECDC2FBDD" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="4F6F369A9A462228FECECCBECF45FC38" blockId="3.[264,768,993,1054]" box="[264,768,993,1019]" pageId="3" pageNumber="4">
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<heading id="142781F69A462228FECECCBECF45FC38" bold="true" box="[264,768,993,1019]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="7DA4EA889A462228FECECCBECF45FC38" bold="true" box="[264,768,993,1019]" pageId="3" pageNumber="4">
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<taxonomicName id="88D04D199A462228FECECCBECEF4FC38" ID-CoL="6F523" authority="Rafael" authorityName="Rafael" box="[264,689,993,1019]" class="Insecta" family="Pipunculidae" genus="Elmohardyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
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<emphasis id="7DA4EA889A462228FECECCBECE13FC38" bold="true" box="[264,598,993,1019]" italics="true" pageId="3" pageNumber="4">Elmohardyia nicaraguensis</emphasis>
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Rafael
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</taxonomicName>
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,
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<taxonomicNameLabel id="669757F39A462228FD78CCBECF45FC38" box="[702,768,993,1019]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4F6F369A9A462228FECECB5BCDC2FBDD" blockId="3.[264,768,993,1054]" box="[264,391,1028,1054]" pageId="3" pageNumber="4">
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(
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<figureCitation id="D7EB2A1F9A462228FED7CB5BCD3BFBDD" box="[273,382,1028,1054]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Figs. 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="07CA65119A46222EFECECB0BC87DFCD5" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="4F6F369A9A462228FECECB0BCEE4FACD" blockId="3.[264,1325,1108,1854]" pageId="3" pageNumber="4">Diagnosis. Scape and pedicel dark brown to black, flagellum yellow. Third costal section about as long as fourth. Abdomen gray pruinose on tergite 1, anterior and posterior border of tergite 2 and posterolaterally on tergites 3–5. Syntergosternite 8 with small membranous area. Inner surstylus strongly developed, longer than outer surstylus. Phallic guide with external side extremely developed.</paragraph>
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<paragraph id="4F6F369A9A462228FEFECA43CF0EF8FD" blockId="3.[264,1325,1108,1854]" pageId="3" pageNumber="4">
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Description. Male: body length 4.5 mm; wing 5.6 mm. Eyes holoptic for a distance of six facets. Frontal triangle and face with gray pruinosity. Scape and pedicel dark brown to black, flagellum yellow. Scutum and scutellum dark brown to black, brown pruinose. Notopleuron, postpronotal lobe and mesopleuron gray pruinose, except mesopleuron near mid and hindcoxae shining, without pruinosity in certain lights. Scutellum with inconspicuous bristles. Legs yellow, except coxae brown and distal tarsomeres brown. Wing with third costal section as long as fourth; vein rm placed near basal third of cell dm. Halter with stem yellow, knob brown to black. Abdomen (
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<figureCitation id="D7EB2A1F9A462228FCA9C96BCFF0F98D" box="[879,949,1588,1614]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
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) dark brown to black, subshining, gray pruinose on tergite 1, anterior and posterior border of tergite 2 and posterolaterally on tergites 3–5; tergite 1 with 1–2 black bristles laterally. Syntergosternite 8 black, longer than tergite 5, with small membranous area. Sternites 6 and 7 gray pruinose. Epandrium and surstyli (
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<figureCitation id="D7EB2A1F9A462228FE31C98BCE05F92D" box="[503,576,1748,1774]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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) yellow. Surstyli asymmetrical, the inner strongly developed, longer than outer surstylus; in lateral view as in
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<figureCitation id="D7EB2A1F9A462228FCFBC9A3CFEBF8D5" box="[829,942,1788,1814]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Figs. 3, 4</figureCitation>
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. Phallic guide as in
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<figureCitation id="D7EB2A1F9A462228FB5FC9A3C8A5F8D5" box="[1177,1248,1788,1814]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
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. Phallus as in
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<figureCitation id="D7EB2A1F9A462228FEA9C87BCDF2F8FD" box="[367,439,1828,1854]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
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. Ejaculatory apodeme as in
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<figureCitation id="D7EB2A1F9A462228FD39C87BCF03F8FD" box="[767,838,1828,1854]" captionStart="FIGURES 1 – 12. 17" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[275,1273,332,1524]" captionTargetId="figure@4.[264,1323,308,1568]" captionTargetPageId="4" captionText="FIGURES 1 – 12. 17 Elmohardyia nicaraguensis, n. sp., ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8 – 12 “ Eudorylas ” maesi, sp. n., pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme." httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
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.
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</paragraph>
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<caption id="1BAF66129A41222FFECEC963CDA8F92D" httpUri="https://zenodo.org/record/157709/files/figure.png" pageId="4" pageNumber="5" targetBox="[275,1273,332,1524]" targetPageId="4">
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<paragraph id="4F6F369A9A41222FFECEC963CDA8F92D" blockId="4.[264,1323,1594,1774]" pageId="4" pageNumber="5">
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<emphasis id="7DA4EA889A41222FFECEC963CD86F997" bold="true" box="[264,451,1596,1620]" pageId="4" pageNumber="5">FIGURES 1–12.</emphasis>
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17
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<taxonomicName id="88D04D199A41222FFE36C962CF5DF997" box="[496,792,1597,1620]" class="Insecta" family="Pipunculidae" genus="Elmohardyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
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<emphasis id="7DA4EA889A41222FFE36C962CF5DF997" box="[496,792,1597,1620]" italics="true" pageId="4" pageNumber="5">Elmohardyia nicaraguensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="669757F39A41222FFCEEC963CF20F990" box="[808,869,1596,1619]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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, ht ɗ: 1. abdomen, dv; 2. epandrium and surstyli, vv; 3. outer surstylus, lv; 4. inner surstylus, lv; 5. hypandrium, lv; 6. phallus, lv; 7. ejaculatory apodeme. 8–12 “
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<taxonomicName id="88D04D199A41222FFE35C9D5CE24F962" box="[499,609,1674,1697]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="7DA4EA889A41222FFE35C9D5CE24F962" box="[499,609,1674,1697]" italics="true" pageId="4" pageNumber="5">Eudorylas</emphasis>
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</taxonomicName>
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”
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<taxonomicName id="88D04D199A41222FFDB3C9D5CEF6F962" box="[629,691,1674,1697]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maesi" status="sp. nov.">
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<emphasis id="7DA4EA889A41222FFDB3C9D5CEF6F962" box="[629,691,1674,1697]" italics="true" pageId="4" pageNumber="5">maesi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="669757F39A41222FFD04C9D5CEBBF962" box="[706,766,1674,1697]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
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, pt ɗ: 8. abdomen, dv; 9. syntergosternite 8, epandrium and surstyli, vv; 10. inner surstylus, lv; 11. apex of phallic guide and phallus, lv; 12. ejaculatory apodeme.
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</paragraph>
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</caption>
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<paragraph id="4F6F369A9A41222EFEFEC87ACD85FEF5" blockId="4.[264,1323,1829,1895]" lastBlockId="5.[264,1325,284,790]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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One male specimen has very similar terminalia. It differs slightly from
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<typeStatus id="906B88389A41222FFBB0C87AC898F8FC" box="[1142,1245,1829,1855]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
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by the shape of the surstylus. Unfortunately the terminalia was lost and we prefer not to designate it as a
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<typeStatus id="906B88389A40222EFE92CE43CDF8FEF5" box="[340,445,284,310]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
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.
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</paragraph>
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<paragraph id="4F6F369A9A40222EFEFECE1BCFFEFE45" blockId="5.[264,1325,284,790]" pageId="5" pageNumber="6">
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Female. One female probably belongs here. It is not being designated
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<typeStatus id="906B88389A40222EFB00CE1BC96EFE9D" box="[1222,1323,324,350]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
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because we are not secure of the association with the male.
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</paragraph>
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<paragraph id="4F6F369A9A40222EFEFECECBC8ACFDE5" blockId="5.[264,1325,284,790]" pageId="5" pageNumber="6">
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Material.
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<typeStatus id="906B88389A40222EFE6BCECBCE5AFE6D" box="[429,543,404,430]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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: male (INPA). Verbatim text of three pin labels:
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<collectingCountry id="37C7760A9A40222EFBB3CECBC96CFE6D" box="[1141,1321,404,430]" name="Nicaragua" pageId="5" pageNumber="6">NICARAGUA</collectingCountry>
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,
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<collectingCountry id="37C7760A9A40222EFECECEE3CD2AFE15" box="[264,367,444,470]" name="Grenada" pageId="5" pageNumber="6">Granada</collectingCountry>
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, Volcán Mombacho, El Progreso,
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<quantity id="88289B7F9A40222EFCCACEE3CF1CFE15" box="[780,857,444,470]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="m" value="700.0">700 m</quantity>
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[white paper] /
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<date id="3B6E105A9A40222EFBD9CEE3C8DCFE15" box="[1055,1177,444,470]" pageId="5" pageNumber="6" value="1998-02-15">15.ii.1998</date>
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, J.M. Maes, Malaise trap [white paper] /
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<typeStatus id="906B88389A40222EFDA6CEBBCE88FE3D" box="[608,717,484,510]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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ɗ,
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<taxonomicName id="88D04D199A40222EFD3DCEBBC8D3FE3D" authority="Rafael" authorityName="Rafael" box="[763,1174,484,510]" class="Insecta" family="Pipunculidae" genus="Elmohardyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="7DA4EA889A40222EFD3DCEBBC805FE3E" box="[763,1088,484,509]" italics="true" pageId="5" pageNumber="6">Elmohardyia nicaraguensis</emphasis>
|
||
Rafael
|
||
</taxonomicName>
|
||
[red paper]. Terminalia placed in a microvial with glycerine and pinned along with the specimen.
|
||
</paragraph>
|
||
<paragraph id="4F6F369A9A40222EFEFECD6BCE4CFDB5" blockId="5.[264,1325,284,790]" pageId="5" pageNumber="6">
|
||
Other specimen: San Joaquin #1,
|
||
<date id="3B6E105A9A40222EFD06CD6BCE98FD8D" box="[704,733,564,590]" pageId="5" pageNumber="6" value="1998-03-31">31</date>
|
||
.iii.1998 (1ɗ MEL, terminalia lost); Santa Ana #3,
|
||
<date id="3B6E105A9A40222EFECECD03CD60FDB5" box="[264,293,604,630]" pageId="5" pageNumber="6" value="1999-02-18">18</date>
|
||
.ii.1999 (1Ψ MEL).
|
||
</paragraph>
|
||
<paragraph id="4F6F369A9A40222EFEFECDDBC85CFD5D" blockId="5.[264,1325,284,790]" box="[312,1049,644,670]" pageId="5" pageNumber="6">
|
||
Etymology. The specific name refers to the country
|
||
<collectingCountry id="37C7760A9A40222EFC5FCDDBC850FD5D" box="[921,1045,644,670]" name="Nicaragua" pageId="5" pageNumber="6">Nicaragua</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4F6F369A9A40222EFEFECDF3C87DFCD5" blockId="5.[264,1325,284,790]" pageId="5" pageNumber="6">
|
||
Remarks.
|
||
<taxonomicName id="88D04D199A40222EFE69CDF3CE30FD06" box="[431,629,684,709]" class="Insecta" family="Pipunculidae" genus="Elmohardyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nicaraguensis" status="sp. nov.">
|
||
<emphasis id="7DA4EA889A40222EFE69CDF3CE30FD06" box="[431,629,684,709]" italics="true" pageId="5" pageNumber="6">E. nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="669757F39A40222EFDBBCDF3CEF8FD05" box="[637,701,684,710]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
is near
|
||
<taxonomicName id="88D04D199A40222EFCDACDF3C84EFD05" authority="Hardy" authorityName="Hardy" box="[796,1035,684,710]" class="Insecta" family="Pipunculidae" genus="Elmohardyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="replicata">
|
||
<emphasis id="7DA4EA889A40222EFCDACDF3CFE3FD06" box="[796,934,684,709]" italics="true" pageId="5" pageNumber="6">E. replicata</emphasis>
|
||
(Hardy)
|
||
</taxonomicName>
|
||
in the similar pattern of abdominal pruinosity and surstyli shape. It is different by the shape of the hypandrium, notably by the extremely developed external side of the phallic guide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |