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<document id="9220FAAAB492850E10A134FBAAC2A394" ID-CLB-Dataset="29882" ID-DOI="10.11646/zootaxa.4434.1.5" ID-GBIF-Dataset="b1745231-e1cf-4233-9ac1-5e6b16290017" ID-ISSN="1175-5326" ID-Zenodo-Dep="1290988" ID-ZooBank="F513BADA-81B7-4DA5-9A23-17B0813268C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529082879981" checkinUser="plazi" docAuthor="Sibaja-Araya, Fabián &amp; Esquivel, Carlos" docDate="2018" docId="710E3237BA6CFFE0FF2D4C085E7CFD72" docLanguage="en" docName="zootaxa.4434.1.5.pdf" docOrigin="Zootaxa 4434 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Camelobaetidius guaycara Sibaja-Araya &amp; Esquivel 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="96" masterDocId="8D374A4FBA6DFFE7FFBA4B205A37FF96" masterDocTitle="Camelobaetidius guaycara, a new species of Baetidae (Ephemeroptera) from Costa Rica, Central America" masterLastPageNumber="98" masterPageNumber="89" pageNumber="90" updateTime="1698549619350" updateUser="ExternalLinkService">
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<mods:title id="F5BDA94E47A144AF0E9C9358D10FE23F">Camelobaetidius guaycara, a new species of Baetidae (Ephemeroptera) from Costa Rica, Central America</mods:title>
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<mods:namePart id="922595D5F88D16347D0B6D1C7045ADDF">Sibaja-Araya, Fabián</mods:namePart>
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<mods:namePart id="10943C664B12470DA4117EB1097F5FBA">Esquivel, Carlos</mods:namePart>
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<treatment id="710E3237BA6CFFE0FF2D4C085E7CFD72" ID-DOI="http://doi.org/10.5281/zenodo.6489064" ID-GBIF-Taxon="145966370" ID-Zenodo-Dep="6489064" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72" httpUri="http://treatment.plazi.org/id/710E3237BA6CFFE0FF2D4C085E7CFD72" lastPageId="7" lastPageNumber="96" pageId="1" pageNumber="90">
<subSubSection id="B1BDD0AABA6CFFE6FF2D4C085B14F8F2" pageId="1" pageNumber="90" type="nomenclature">
<paragraph id="F9188321BA6CFFE6FF2D4C0859B1F8D5" blockId="1.[151,902,1832,1892]" box="[151,902,1832,1859]" pageId="1" pageNumber="90">
<heading id="A250344DBA6CFFE6FF2D4C0859B1F8D5" bold="true" box="[151,902,1832,1859]" fontSize="11" level="1" pageId="1" pageNumber="90" reason="1">
<emphasis id="CBD35F33BA6CFFE6FF2D4C0859B1F8D5" bold="true" box="[151,902,1832,1859]" pageId="1" pageNumber="90">
<taxonomicName id="3EA7F8A2BA6CFFE6FF2D4C08592CF8D5" authority="Sibaja-Araya &amp; Esquivel" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[151,795,1832,1859]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA6CFFE6FF2D4C085BEFF8D4" bold="true" box="[151,472,1832,1858]" italics="true" pageId="1" pageNumber="90">Camelobaetidius guaycara</emphasis>
Sibaja-Araya &amp; Esquivel
</taxonomicName>
,
<taxonomicNameLabel id="D0E0E248BA6CFFE6FC924C0959B1F8D5" box="[808,902,1833,1859]" pageId="1" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F9188321BA6CFFE6FF2D4C6B5B14F8F2" blockId="1.[151,902,1832,1892]" box="[151,291,1867,1892]" pageId="1" pageNumber="90">
<figureCitation id="619C9FA4BA6CFFE6FF2D4C6B5ACAF8F2" box="[151,253,1867,1892]" captionStart="FIGURE 1" captionStartId="1.[151,250,1659,1681]" captionTargetBox="[151,1436,411,1635]" captionTargetId="figure@1.[151,1436,411,1635]" captionTargetPageId="1" captionText="FIGURE 1. Geographic distribution of Camelobaetidius guaycara sp. nov. in Costa Rica." httpUri="https://zenodo.org/record/1290990/files/figure.png" pageId="1" pageNumber="90">FigureS 1</figureCitation>
8.
</paragraph>
</subSubSection>
<subSubSection id="B1BDD0AABA6CFFE2FF2D4CB35EA6F87D" lastPageId="5" lastPageNumber="94" pageId="1" pageNumber="90" type="diagnosis">
<paragraph id="F9188321BA6CFFE5FF2D4CB35EB4FED6" blockId="1.[151,1437,1939,2036]" lastBlockId="2.[151,1437,151,752]" lastPageId="2" lastPageNumber="91" pageId="1" pageNumber="90">
<emphasis id="CBD35F33BA6CFFE6FF2D4CB35BE8F83A" bold="true" box="[151,479,1939,1964]" pageId="1" pageNumber="90">Diagnosis. Mature nymphs</emphasis>
. 1) labrum with anterior margin with about 13 Small, double, frayed Setae; arc of anterodorSal Setae of labrum with 1 + 6 Setae, the two inner oneS frayed; intermediate Setae preSent; Several Small Simple Setae near poSterior margin; ventral Surface with a denSe arc of Short, fine Setae near the anterior margin and with 4 Short Setae near lateral margin; a minute Seta near the anterior margin on both
<collectingRegion id="3B634DC3BA6FFFE5FBF54BB75EB0FF26" box="[1103,1159,151,176]" country="Haiti" name="Sud-Est" pageId="2" pageNumber="91">SideS</collectingRegion>
of the cleft (
<figureCitation id="619C9FA4BA6FFFE5FAAC4BB75F5FFF26" box="[1302,1384,151,176]" captionStart="FIGURE 4" captionStartId="4.[151,250,1964,1986]" captionTargetBox="[222,1365,204,1945]" captionTargetId="figure@4.[213,1377,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Camelobaetidius guaycara sp. nov., nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail." httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="2" pageNumber="91">Fig. 4a</figureCitation>
); 2); left mandible with 5 Short inciSorS, tip of the Slender proceSS eXtended at about the Same level aS the inciSorS (
<figureCitation id="619C9FA4BA6FFFE5FAD54B9C5A81FF6E" captionStart="FIGURE 4" captionStartId="4.[151,250,1964,1986]" captionTargetBox="[222,1365,204,1945]" captionTargetId="figure@4.[213,1377,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Camelobaetidius guaycara sp. nov., nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail." httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="2" pageNumber="91">Fig. 4b</figureCitation>
); 3) Segment II of labial palp with a Short triangular diStomedial projection (
<figureCitation id="619C9FA4BA6FFFE5FBBE4BFF5EA2FF6E" box="[1028,1173,223,248]" captionStart="FIGURE 5" captionStartId="5.[151,250,1617,1639]" captionTargetBox="[177,1397,202,1577]" captionTargetId="figure@5.[173,1417,193,1603]" captionTargetPageId="5" captionText="FIGURE 5. Camelobaetidius guaycara sp. nov., nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations." httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="2" pageNumber="91">FigS. 5c, d, e</figureCitation>
); 4); one Small thoracic gill at the baSe of foreleg; 5) tarSal clawS with 2732 denticleS (
<figureCitation id="619C9FA4BA6FFFE5FCDC4A24598DFE8B" box="[870,954,260,285]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="2" pageNumber="91">Fig. 6b</figureCitation>
); 6) poSterior margin of abdominal tergum IV with rounded SpineS of different SizeS (
<figureCitation id="619C9FA4BA6FFFE5FDD54A0758F6FED6" box="[623,705,295,320]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="2" pageNumber="91">Fig. 6c</figureCitation>
); 7) paraproct with 13 SpineS (
<figureCitation id="619C9FA4BA6FFFE5FB9E4A075E41FED6" box="[1060,1142,295,320]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="2" pageNumber="91">Fig. 6e</figureCitation>
).
</paragraph>
<paragraph id="F9188321BA6FFFE5FF7D4A6C5865FE11" blockId="2.[151,1437,151,752]" pageId="2" pageNumber="91">
Size: Mature nymphS. Body length:
<quantity id="3E5F2EC4BA6FFFE5FDD94A6C58D9FEF2" box="[611,750,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="2" pageNumber="91" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
, femaleS larger than maleS; antenna: 2.03.0 mm; cerci 3.5 4.0 mm; terminal filament 3.0
<quantity id="3E5F2EC4BA6FFFE5FE4C4A4F5879FE11" box="[502,590,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="91" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="F9188321BA6FFFE5FF7D4AB45B0FFD66" blockId="2.[151,1437,151,752]" pageId="2" pageNumber="91">
Body coloration: head with olive SpotS on fronS; pro and metanotum homogeneouS dark olive; meSonotum with elongate dark olive markS; femora with elongate olive markS on anterior face; abdominal terga coloration SeXually dimorphic, additionally both SeXeS from both SlopeS have diStinct patternS of abdominal coloration (
<figureCitation id="619C9FA4BA6FFFE5FAEB4AFC5FB9FE62" box="[1361,1422,476,501]" captionStart="FIGURE 2" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[169,1417,829,1715]" captionTargetId="figure@2.[151,1436,805,1737]" captionTargetPageId="2" captionText="FIGURE 2. Variation in body coloration of Camelobaetidius guaycara sp. nov from the Atlantic and Pacific slopes of Costa Rica (males a: Pacific slope, b: Atlantic slope; females c: Pacific slope, d: Atlantic slope)." httpUri="https://zenodo.org/record/1290992/files/figure.png" pageId="2" pageNumber="91">Fig 2</figureCitation>
), that of maleS being aS followS:
<emphasis id="CBD35F33BA6FFFE5FE4B4ADF5830FD8E" bold="true" box="[497,519,511,536]" pageId="2" pageNumber="91">a)</emphasis>
tergum VIII completely white while other terga have a dark/light olive pattern (14 individualS),
<emphasis id="CBD35F33BA6FFFE5FE9449045B7FFDAB" bold="true" box="[302,328,548,573]" pageId="2" pageNumber="91">b)</emphasis>
terga I, II, VIII completely white, other terga with dark/light olive pattern (21 individualS),
<emphasis id="CBD35F33BA6FFFE5FA3C49045FABFDAB" bold="true" box="[1414,1436,548,573]" pageId="2" pageNumber="91">c)</emphasis>
tergum VIII light gray with two Small SpotS and other terga with dark/light olive pattern (21 individualS), and
<emphasis id="CBD35F33BA6FFFE5FADA49675F4FFDF6" bold="true" box="[1376,1400,583,608]" pageId="2" pageNumber="91">d)</emphasis>
all terga with dark/light olive pattern (12 individualS); femaleS preSent two coloration patternS: one Similar to male pattern
<emphasis id="CBD35F33BA6FFFE5FF5149AF5B36FD3E" bold="true" box="[235,257,655,680]" pageId="2" pageNumber="91">c)</emphasis>
(15 individualS), and the other like male pattern
<emphasis id="CBD35F33BA6FFFE5FC9849AF590BFD3E" bold="true" box="[802,828,655,680]" pageId="2" pageNumber="91">d)</emphasis>
(31 individualS); femora with elongate olive markS on anterior face, femur-tibia articulation dark brown; tibiae and tarSi white; cerci dark grey and terminal filament white (
<figureCitation id="619C9FA4BA6FFFE5FF5E49F75B1EFD66" box="[228,297,727,752]" captionStart="FIGURE 3" captionStartId="3.[151,250,1193,1215]" captionTargetBox="[519,1068,265,1172]" captionTargetId="figure@3.[519,1068,265,1172]" captionTargetPageId="3" captionText="FIGURE 3. Coloration pattern of cerci and central filament of Camelobaetidius guaycara sp. nov. from the Atlantic and Pacific slopes of Costa Rica." httpUri="https://zenodo.org/record/1290994/files/figure.png" pageId="2" pageNumber="91">Fig. 3</figureCitation>
).
</paragraph>
<caption id="ADD8D3A9BA6FFFE5FF2D4DFF5E1EF885" httpUri="https://zenodo.org/record/1290992/files/figure.png" pageId="2" pageNumber="91" startId="2.[151,250,1759,1781]" targetBox="[169,1417,829,1715]" targetPageId="2">
<paragraph id="F9188321BA6FFFE5FF2D4DFF5E1EF885" blockId="2.[151,1436,1759,1811]" pageId="2" pageNumber="91">
<emphasis id="CBD35F33BA6FFFE5FF2D4DFF5B23F963" bold="true" box="[151,276,1759,1781]" pageId="2" pageNumber="91">FIGURE 2.</emphasis>
Variation in body coloration of
<taxonomicName id="3EA7F8A2BA6FFFE5FDE54DFF5945F963" authority="Sibaja-Araya &amp; Esquivel, 2018" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[607,882,1759,1781]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA6FFFE5FDE54DFF5945F963" box="[607,882,1759,1781]" italics="true" pageId="2" pageNumber="91">Camelobaetidius guaycara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0E0E248BA6FFFE5FCC34DFF59F1F963" box="[889,966,1759,1781]" pageId="2" pageNumber="91" rank="species">
<emphasis id="CBD35F33BA6FFFE5FCC34DFF59F1F963" bold="true" box="[889,966,1759,1781]" pageId="2" pageNumber="91">sp. nov</emphasis>
</taxonomicNameLabel>
from the Atlantic and Pacific slopes of Costa Rica (males
<emphasis id="CBD35F33BA6FFFE5FEAF4DDD5B15F885" bold="true" box="[277,290,1789,1811]" pageId="2" pageNumber="91">a</emphasis>
: Pacific slope,
<emphasis id="CBD35F33BA6FFFE5FE044DDD5BFBF885" bold="true" box="[446,460,1789,1811]" pageId="2" pageNumber="91">b</emphasis>
: Atlantic slope; females
<emphasis id="CBD35F33BA6FFFE5FD734DDD58E3F885" bold="true" box="[713,724,1789,1811]" pageId="2" pageNumber="91">c</emphasis>
: Pacific slope,
<emphasis id="CBD35F33BA6FFFE5FCCA4DDD5949F885" bold="true" box="[880,894,1789,1811]" pageId="2" pageNumber="91">d</emphasis>
: Atlantic slope).
</paragraph>
</caption>
<paragraph id="F9188321BA6FFFE5FF7D4C1B5836F8EE" blockId="2.[151,1437,1851,2020]" pageId="2" pageNumber="91">
<emphasis id="CBD35F33BA6FFFE5FF7D4C1B5BC7F8C2" bold="true" box="[199,496,1851,1876]" pageId="2" pageNumber="91">Body morphology. Head</emphasis>
: Antennae Scape and pedicel with minute, Simple Setae on lateral marginS; both dorSal and ventral SurfaceS bare.
</paragraph>
<paragraph id="F9188321BA6FFFE5FF7D4CA35FABF875" blockId="2.[151,1437,1851,2020]" pageId="2" pageNumber="91">
Labrum (
<figureCitation id="619C9FA4BA6FFFE5FE8B4CA35BB4F80D" box="[305,387,1923,1948]" captionStart="FIGURE 4" captionStartId="4.[151,250,1964,1986]" captionTargetBox="[222,1365,204,1945]" captionTargetId="figure@4.[213,1377,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Camelobaetidius guaycara sp. nov., nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail." httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="2" pageNumber="91">Fig. 4a</figureCitation>
) broader than long; anterior margin with about 13 double, frayed Setae, arc of anterodorSal Setae with 1 + 6 Setae, Second inner Seta longer than the firSt, both frayed, remainder of Setae Simple, all Setae of arc long, eXtending beyond margin of labrum; intermediate Seta abSent; lateral margin with 4 Simple Setae and 911
</paragraph>
<paragraph id="F9188321BA6EFFE4FF2D4BB75E5CFF43" blockId="3.[151,1436,151,213]" pageId="3" pageNumber="92">
frayed, anterolateral Setae. Ventral Surface with a denSe arc of Short, fine Setae near anterior margin, 4 Small Setae near the lateral margin and a minute Seta near the anterior margin on both
<collectingRegion id="3B634DC3BA6EFFE4FC6E4B9C5E3BFF43" box="[980,1036,188,213]" country="Haiti" name="Sud-Est" pageId="3" pageNumber="92">SideS</collectingRegion>
of cleft.
</paragraph>
<caption id="ADD8D3A9BA6EFFE4FF2D4F895B8DFB48" httpUri="https://zenodo.org/record/1290994/files/figure.png" pageId="3" pageNumber="92" startId="3.[151,250,1193,1215]" targetBox="[519,1068,265,1172]" targetPageId="3">
<paragraph id="F9188321BA6EFFE4FF2D4F895B8DFB48" blockId="3.[151,1435,1193,1246]" pageId="3" pageNumber="92">
<emphasis id="CBD35F33BA6EFFE4FF2D4F895B21FB29" bold="true" box="[151,278,1193,1215]" pageId="3" pageNumber="92">FIGURE 3.</emphasis>
Coloration pattern of cerci and central filament of
<taxonomicName id="3EA7F8A2BA6EFFE4FC814F895E78FB29" authority="Sibaja-Araya &amp; Esquivel, 2018" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[827,1103,1193,1215]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA6EFFE4FC814F895E78FB29" box="[827,1103,1193,1215]" italics="true" pageId="3" pageNumber="92">Camelobaetidius guaycara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0E0E248BA6EFFE4FBE04F895E99FB29" box="[1114,1198,1193,1215]" pageId="3" pageNumber="92" rank="species">
<emphasis id="CBD35F33BA6EFFE4FBE04F895E99FB29" bold="true" box="[1114,1198,1193,1215]" pageId="3" pageNumber="92">sp. nov.</emphasis>
</taxonomicNameLabel>
from the Atlantic and Pacific slopes of Costa Rica.
</paragraph>
</caption>
<paragraph id="F9188321BA6EFFE4FF7D4E2A58B2FA19" blockId="3.[151,1437,1290,2035]" pageId="3" pageNumber="92">
Left mandible (
<figureCitation id="619C9FA4BA6EFFE4FECC4E2A5BFBFAB5" box="[374,460,1290,1315]" captionStart="FIGURE 4" captionStartId="4.[151,250,1964,1986]" captionTargetBox="[222,1365,204,1945]" captionTargetId="figure@4.[213,1377,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Camelobaetidius guaycara sp. nov., nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail." httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="3" pageNumber="92">Fig. 4b</figureCitation>
): inciSorS with 6 denticleS Similar in Size; proStheca well developed; no Setae between proStheca and mola; area bearing the Slender proceSS (thumb) raiSed, the tip of the Slender proceSS eXtending to about Same level aS inciSorS; molar region longer than length between proStheca and Slender proceSS; and three Simple Setae on the mandibleS baSal region.
</paragraph>
<paragraph id="F9188321BA6EFFE4FF7D4EBA5AD1F989" blockId="3.[151,1437,1290,2035]" pageId="3" pageNumber="92">
Right mandible (
<figureCitation id="619C9FA4BA6EFFE4FE3D4EBA5BEBFA25" box="[391,476,1434,1459]" captionStart="FIGURE 4" captionStartId="4.[151,250,1964,1986]" captionTargetBox="[222,1365,204,1945]" captionTargetId="figure@4.[213,1377,193,1949]" captionTargetPageId="4" captionText="FIGURE 4. Camelobaetidius guaycara sp. nov., nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail." httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="3" pageNumber="92">Fig. 4c</figureCitation>
): inciSorS with 6 denticleS Similar in Size; proStheca not well developed; three Small Simple Setae between proStheca and mola; molar region with a well developed diSk-like proceSS (“molar diSk”) (fig. 4d); molar region Shorter than length between proStheca and molar diSk and four Simple Setae on mandibleS baSal region.
</paragraph>
<paragraph id="F9188321BA6EFFE4FF7D4D0A5F5AF9F1" blockId="3.[151,1437,1290,2035]" pageId="3" pageNumber="92">
HypopharynX (
<figureCitation id="619C9FA4BA6EFFE4FECD4D0A5BE7F9D5" box="[375,464,1578,1603]" captionStart="FIGURE 5" captionStartId="5.[151,250,1617,1639]" captionTargetBox="[177,1397,202,1577]" captionTargetId="figure@5.[173,1417,193,1603]" captionTargetPageId="5" captionText="FIGURE 5. Camelobaetidius guaycara sp. nov., nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations." httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="3" pageNumber="92">Fig. 5a</figureCitation>
): lingua Shorter than Superlinguae, apically hairy on dorSal and ventral SurfaceS; Superlinguae apically hairy on dorSal and ventral SurfaceS and with Serrate area at the middle of lateral margin.
</paragraph>
<paragraph id="F9188321BA6EFFE4FF7D4D5258B8F961" blockId="3.[151,1437,1290,2035]" pageId="3" pageNumber="92">
MaXillae (
<figureCitation id="619C9FA4BA6EFFE4FE814D525BA6F91D" box="[315,401,1650,1675]" captionStart="FIGURE 5" captionStartId="5.[151,250,1617,1639]" captionTargetBox="[177,1397,202,1577]" captionTargetId="figure@5.[173,1417,193,1603]" captionTargetPageId="5" captionText="FIGURE 5. Camelobaetidius guaycara sp. nov., nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations." httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="3" pageNumber="92">Fig. 5b</figureCitation>
): robuSt; galea with four apical canineS and two apical rowS of Setae, Some Setae of one of theSe rowS large and thick while otherS are thin, and Setae of other row Shorter and curved; three Setae on SubdiStal internal margin; maXillary palp not SurpaSSing apical canineS, Segment I Short; Segment II thick, bicepS-like; Segment III elongate, longer than Segment II.
</paragraph>
<paragraph id="F9188321BA6EFFE4FF7D4C225AE3F865" blockId="3.[151,1437,1290,2035]" pageId="3" pageNumber="92">
Labium (
<figureCitation id="619C9FA4BA6EFFE4FE954C225BB4F88D" box="[303,387,1794,1819]" captionStart="FIGURE 5" captionStartId="5.[151,250,1617,1639]" captionTargetBox="[177,1397,202,1577]" captionTargetId="figure@5.[173,1417,193,1603]" captionTargetPageId="5" captionText="FIGURE 5. Camelobaetidius guaycara sp. nov., nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations." httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="3" pageNumber="92">Fig. 5c</figureCitation>
): paragloSSa and gloSSa equal in length, both dorSally and ventrally with numerouS elongate and curved Setae on apeX; palp Segment I thick, aS long aS SegmentS II + III, dorSal Surface with Several microporeS along eXternal margin; Segment II with a Short triangular diStomedial projection, variable aS Shown in
<figureCitation id="619C9FA4BA6EFFE4FAA14C6A5A84F811" captionStart="FIGURE 5" captionStartId="5.[151,250,1617,1639]" captionTargetBox="[177,1397,202,1577]" captionTargetId="figure@5.[173,1417,193,1603]" captionTargetPageId="5" captionText="FIGURE 5. Camelobaetidius guaycara sp. nov., nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations." httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="3" pageNumber="92">Fig. 5d and 5e</figureCitation>
; diStomedial projection diStinctly larger on the right palp of Some individualS, dorSal Surface with Several microporeS and a row of 34 Simple Setae, 58 Simple marginal Setae on ventral Surface; Segment III rounded, broader than longer, with numerouS robuSt SpineS on ventral Surface, dorSal Surface with a row of robuSt Setae on apeX.
</paragraph>
<caption id="ADD8D3A9BA69FFE3FF2D4C8C5BC9F877" httpUri="https://zenodo.org/record/1290996/files/figure.png" pageId="4" pageNumber="93" startId="4.[151,250,1964,1986]" targetBox="[222,1365,204,1945]" targetPageId="4">
<paragraph id="F9188321BA69FFE3FF2D4C8C5BC9F877" blockId="4.[151,1435,1964,2017]" pageId="4" pageNumber="93">
<emphasis id="CBD35F33BA69FFE3FF2D4C8C5B25F854" bold="true" box="[151,274,1964,1986]" pageId="4" pageNumber="93">FIGURE 4.</emphasis>
<taxonomicName id="3EA7F8A2BA69FFE3FEA24C8C581DF855" authority="Sibaja-Araya &amp; Esquivel, 2018" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[280,554,1964,1987]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA69FFE3FEA24C8C581DF855" box="[280,554,1964,1987]" italics="true" pageId="4" pageNumber="93">Camelobaetidius guaycara</emphasis>
</taxonomicName>
<emphasis id="CBD35F33BA69FFE3FD954C8C58B1F854" bold="true" box="[559,646,1964,1986]" pageId="4" pageNumber="93">
<taxonomicNameLabel id="D0E0E248BA69FFE3FD954C8C58B6F854" box="[559,641,1964,1986]" pageId="4" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
nymph. a) labrum (left d. v., right v. v.), b) left mandible, c) right mandible, d) molar disk of right mandible detail.
</paragraph>
</caption>
<caption id="ADD8D3A9BA68FFE2FF2D4D71580CF913" httpUri="https://zenodo.org/record/1290998/files/figure.png" pageId="5" pageNumber="94" startId="5.[151,250,1617,1639]" targetBox="[177,1397,202,1577]" targetPageId="5">
<paragraph id="F9188321BA68FFE2FF2D4D71580CF913" blockId="5.[151,1436,1617,1669]" pageId="5" pageNumber="94">
<emphasis id="CBD35F33BA68FFE2FF2D4D715B25F9F1" bold="true" box="[151,274,1617,1639]" pageId="5" pageNumber="94">FIGURE 5.</emphasis>
<taxonomicName id="3EA7F8A2BA68FFE2FEA34D71581CF9F1" authority="Sibaja-Araya &amp; Esquivel, 2018" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[281,555,1617,1639]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA68FFE2FEA34D71581CF9F1" box="[281,555,1617,1639]" italics="true" pageId="5" pageNumber="94">Camelobaetidius guaycara</emphasis>
</taxonomicName>
<emphasis id="CBD35F33BA68FFE2FD8A4D7158BFF9F1" bold="true" box="[560,648,1617,1639]" pageId="5" pageNumber="94">
<taxonomicNameLabel id="D0E0E248BA68FFE2FD8A4D7158B4F9F1" box="[560,643,1617,1639]" pageId="5" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
nymph. a) hypopharynx (left v. v., right d. v.), b) maxilla, c) labium (left v. v., right d. v.), d), e) labial palp II variations.
</paragraph>
</caption>
<paragraph id="F9188321BA68FFE2FF7D4D925EA6F87D" blockId="5.[151,1436,1714,2027]" pageId="5" pageNumber="94">
<emphasis id="CBD35F33BA68FFE2FF7D4D925B1EF95D" bold="true" box="[199,297,1714,1739]" pageId="5" pageNumber="94">Thorax:</emphasis>
DorSally with dark olive pattern, pro and metanotum entirely dark olive, meSonotum with elongate dark olive markS; ventrally yellowiSh. Hind wing padS preSent. Small nipple-like thoracic gillS preSent at baSe of forecoXa; femora with elongate olive markS on anterior face, dorSal edge with one row of 4045 long Spine-like Setae, ventral edge with numerouS microporeS and Several Short Spine-like Setae, anterior face with Several fine and Short Setae, articulation femur-tibia dark brown; tibia light brown, patella-tibial Suture preSent, two rowS of Setae on dorSal edge, one of them with fine long Setae and the other with fine, Short Setae, ventral edge with Several microporeS near articulation with femur-tibia and 56 Spine-like Setae along itS margin, anterior face with Several fine and Short Setae; tarSi light brown, Several Short fine Setae on dorSal edge, ventral edge with 6 Setae along margin, 2 of them long and 1 Short on apeX (
<figureCitation id="619C9FA4BA68FFE2FD334CF258ECF87D" box="[649,731,2002,2027]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="5" pageNumber="94">Fig. 6a</figureCitation>
); clawS with 2732 denticleS (
<figureCitation id="619C9FA4BA68FFE2FB8A4CF25EB4F87D" box="[1072,1155,2002,2027]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="5" pageNumber="94">Fig. 6b</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="ADD8D3A9BA6BFFE1FF2D4C925AC9F871" httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="6" pageNumber="95" startId="6.[151,250,1970,1992]" targetBox="[199,1392,219,1933]" targetPageId="6">
<paragraph id="F9188321BA6BFFE1FF2D4C925AC9F871" blockId="6.[151,1436,1970,2023]" pageId="6" pageNumber="95">
<emphasis id="CBD35F33BA6BFFE1FF2D4C925B20F85E" bold="true" box="[151,279,1970,1992]" pageId="6" pageNumber="95">FIGURE 6.</emphasis>
<taxonomicName id="3EA7F8A2BA6BFFE1FE9B4C925800F85E" authority="Sibaja-Araya &amp; Esquivel, 2018" authorityName="Sibaja-Araya &amp; Esquivel" authorityYear="2018" box="[289,567,1970,1992]" class="Insecta" family="Baetidae" genus="Camelobaetidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="guaycara" status="sp. nov.">
<emphasis id="CBD35F33BA6BFFE1FE9B4C925800F85E" box="[289,567,1970,1992]" italics="true" pageId="6" pageNumber="95">Camelobaetidius guaycara</emphasis>
</taxonomicName>
<emphasis id="CBD35F33BA6BFFE1FDFB4C9258ABF85E" bold="true" box="[577,668,1970,1992]" pageId="6" pageNumber="95">
<taxonomicNameLabel id="D0E0E248BA6BFFE1FDFB4C9258AFF85E" box="[577,664,1970,1992]" pageId="6" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct.
</paragraph>
</caption>
<subSubSection id="B1BDD0AABA6AFFE0FF7D4BB75BADFEF6" pageId="7" pageNumber="96" type="description">
<paragraph id="F9188321BA6AFFE0FF7D4BB75F19FEAB" blockId="7.[151,1437,151,740]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FF7D4BB75B72FF26" bold="true" box="[199,325,151,176]" pageId="7" pageNumber="96">Abdomen:</emphasis>
poSterior margin of abdominal terga with rounded SpineS of different SizeS and Simple fine Setae not SurpaSSing length of larger SpineS (
<figureCitation id="619C9FA4BA6AFFE0FDE44B9B5883FF42" box="[606,692,187,212]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="7" pageNumber="96">Fig. 6c</figureCitation>
); gillS white, oval; reduced on SegmentS I and VII and with no pigmented tracheation; well-developed tracheation on otherS gillS, marginS with fine, Small Setae (
<figureCitation id="619C9FA4BA6AFFE0FA8E4BFE5FB9FF61" box="[1332,1422,222,247]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="7" pageNumber="96">Fig. 6d</figureCitation>
); paraproct with 13 Sclerotized SpineS, Several microporeS and Simple, fine Setae on dorSal Surface, (
<figureCitation id="619C9FA4BA6AFFE0FB474A215F67FE8C" box="[1277,1360,257,282]" captionStart="FIGURE 6" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[199,1392,219,1933]" captionTargetId="figure@6.[183,1403,193,1949]" captionTargetPageId="6" captionText="FIGURE 6. Camelobaetidius guaycara sp. nov., nymph. a) midleg, b) claws, c) posterior margin of terga IV, d) gill, e) paraproct." httpUri="https://zenodo.org/record/1291000/files/figure.png" pageId="7" pageNumber="96">Fig. 6e</figureCitation>
); cerci dark gray and terminal filament white, cerci approXimately 12% longer than the central filament (
<figureCitation id="619C9FA4BA6AFFE0FB664A045F17FEAB" box="[1244,1312,292,317]" captionStart="FIGURE 3" captionStartId="3.[151,250,1193,1215]" captionTargetBox="[519,1068,265,1172]" captionTargetId="figure@3.[519,1068,265,1172]" captionTargetPageId="3" captionText="FIGURE 3. Coloration pattern of cerci and central filament of Camelobaetidius guaycara sp. nov. from the Atlantic and Pacific slopes of Costa Rica." httpUri="https://zenodo.org/record/1290994/files/figure.png" pageId="7" pageNumber="96">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="F9188321BA6AFFE0FF7D4A675BADFEF6" blockId="7.[151,1437,151,740]" box="[199,410,327,352]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FF7D4A675B2AFEF6" bold="true" box="[199,285,327,352]" pageId="7" pageNumber="96">Adults.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B1BDD0AABA6AFFE0FF7D4A4B58BFFD0B" pageId="7" pageNumber="96" type="materials_examined">
<paragraph id="F9188321BA6AFFE0FF7D4A4B58BFFD0B" blockId="7.[151,1437,151,740]" pageId="7" pageNumber="96">
<materialsCitation id="49CF897CBA6AFFE0FF7D4A4B5E48FE5C" ID-GBIF-Occurrence="1849198423" collectingDate="2017-04-15" collectionCode="MNCR" collectorName="F. Sibaja-Araya" country="Costa Rica" county="Material" latitude="8.686958" location="Rio Claro" longLatPrecision="1" longitude="-83.04688" municipality="Rio Claro de Golfito" pageId="7" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="PuntarenaS province" typeStatus="holotype">
<emphasis id="CBD35F33BA6AFFE0FF7D4A4B5811FE12" bold="true" box="[199,550,363,388]" pageId="7" pageNumber="96">
<collectingCounty id="1079FBADBA6AFFE0FF7D4A4B5B18FE12" box="[199,303,363,388]" pageId="7" pageNumber="96">Material</collectingCounty>
examined:
<typeStatus id="261C3D83BA6AFFE0FE034A4B5811FE12" box="[441,550,363,388]" pageId="7" pageNumber="96" type="holotype">Holotype</typeStatus>
</emphasis>
, mature
<specimenCount id="EFA148A8BA6AFFE0FD3D4A4C58ABFE15" box="[647,668,364,387]" pageId="7" pageNumber="96" type="female"></specimenCount>
nymph on 95% alcohol (mouthpartS, legS, gillS, terga and paraproctS in microvial),
<collectingCountry id="81B0C3B1BA6AFFE0FE834AAF5B8FFE31" box="[313,440,398,423]" name="Costa Rica" pageId="7" pageNumber="96">CoSta Rica</collectingCountry>
,
<collectingRegion id="3B634DC3BA6AFFE0FE7F4AAF589AFE31" box="[453,685,398,423]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="96">PuntarenaS province</collectingRegion>
,
<collectingMunicipality id="197C195BBA6AFFE0FD034AAE599FFE31" box="[697,936,398,423]" pageId="7" pageNumber="96">Río Claro de Golfito</collectingMunicipality>
,
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,
<geoCoordinate id="9C93E5E6BA6AFFE0FB894AAE5EE5FE31" box="[1075,1234,398,423]" direction="north" orientation="latitude" pageId="7" pageNumber="96" precision="1" value="8.686958">8°4113.05N</geoCoordinate>
/
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, 70 meterS above
<location id="FC78D5FABA6AFFE0FEE54A925BB4FE5C" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72:FC78D5FABA6AFFE0FEE54A925BB4FE5C" box="[351,387,434,458]" country="Costa Rica" county="Material" latitude="8.686958" longLatPrecision="1" longitude="-83.04688" municipality="Rio Claro de Golfito" name="Sea" pageId="7" pageNumber="96" stateProvince="PuntarenaS province">Sea</location>
level,
<collectingDate id="9D5D5C09BA6AFFE0FE6C4A91586EFE5C" box="[470,601,433,458]" pageId="7" pageNumber="96" value="2017-04-15">
<date id="8D19A5E1BA6AFFE0FE6C4A91586EFE5C" box="[470,601,433,458]" pageId="7" pageNumber="96" value="2017-04-15">IV/15/2017</date>
</collectingDate>
,
<collectorName id="5452E6F7BA6AFFE0FDD24A92592CFE5C" box="[616,795,433,458]" pageId="7" pageNumber="96">F. Sibaja-Araya</collectorName>
coll., depoSited at the
<collectionCode id="9FB61BE4BA6AFFE0FB924A915E48FE5C" box="[1064,1151,433,458]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FB924A915E48FE5C" bold="true" box="[1064,1151,433,458]" pageId="7" pageNumber="96">MNCR</emphasis>
</collectionCode>
</materialsCitation>
.
<materialsCitation id="49CF897CBA6AFFE0FB2B4A9158BFFD0B" ID-GBIF-Occurrence="1849198422" collectingDate="2017-06-15" collectionCode="LEUNA, MZUCR, PERC, MNCR, MNRJ, FAMU" collectorName="F. Sibaja-Araya &amp; M. Guevara-Mora &amp; D. Romero-Serrano" country="Costa Rica" county="Same" latitude="10.301375" location="Same" longLatPrecision="1" longitude="-83.62352" municipality="Same" pageId="7" pageNumber="96" specimenCount="4" stateProvince="LimOn province" typeStatus="paratype">
<typeStatus id="261C3D83BA6AFFE0FB2B4A915F3EFE5C" box="[1169,1289,433,458]" pageId="7" pageNumber="96" type="paratype">
<emphasis id="CBD35F33BA6AFFE0FB2B4A915F3EFE5C" bold="true" box="[1169,1289,433,458]" pageId="7" pageNumber="96">Paratypes</emphasis>
</typeStatus>
, two mature nymphS (partS in microvial),
<collectingCounty id="1079FBADBA6AFFE0FE424AF55805FE7B" box="[504,562,469,493]" pageId="7" pageNumber="96">Same</collectingCounty>
data aS the
<typeStatus id="261C3D83BA6AFFE0FD6B4AF45901FE7B" box="[721,822,468,493]" pageId="7" pageNumber="96" type="holotype">holotype</typeStatus>
, houSed at
<collectionCode id="9FB61BE4BA6AFFE0FC7D4AF45E1EFE7B" box="[967,1065,468,493]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FC7D4AF45E1EFE7B" bold="true" box="[967,1065,468,493]" pageId="7" pageNumber="96">LEUNA</emphasis>
</collectionCode>
; two mature nymphS (partS in microvial),
<collectingMunicipality id="197C195BBA6AFFE0FEA04AD85B64FD86" box="[282,339,504,528]" pageId="7" pageNumber="96">Same</collectingMunicipality>
data aS the
<typeStatus id="261C3D83BA6AFFE0FE6D4AD7580BFD86" box="[471,572,503,528]" pageId="7" pageNumber="96" type="holotype">holotype</typeStatus>
, houSed at
<collectionCode id="9FB61BE4BA6AFFE0FD034AD75914FD86" box="[697,803,503,528]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FD034AD75914FD86" bold="true" box="[697,803,503,528]" pageId="7" pageNumber="96">MZUCR</emphasis>
</collectionCode>
; two mature nymphS,
<location id="FC78D5FABA6AFFE0FBA44AD85E6FFD86" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72:FC78D5FABA6AFFE0FBA44AD85E6FFD86" box="[1054,1112,504,528]" country="Costa Rica" county="Same" latitude="10.301375" longLatPrecision="1" longitude="-83.62352" municipality="Same" name="Same" pageId="7" pageNumber="96" stateProvince="LimOn province">Same</location>
data aS the
<typeStatus id="261C3D83BA6AFFE0FB674AD75F75FD86" box="[1245,1346,503,528]" pageId="7" pageNumber="96" type="holotype">holotype</typeStatus>
, houSed at
<collectionCode id="9FB61BE4BA6AFFE0FF0E493B5B37FDA2" box="[180,256,539,564]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FF0E493B5B37FDA2" bold="true" box="[180,256,539,564]" pageId="7" pageNumber="96">PERC</emphasis>
</collectionCode>
; eight mature nymphS,
<collectingCountry id="81B0C3B1BA6AFFE0FDAE493B58A3FDA2" box="[532,660,539,564]" name="Costa Rica" pageId="7" pageNumber="96">CoSta Rica</collectingCountry>
,
<collectingRegion id="3B634DC3BA6AFFE0FD1B493B596CFDA2" box="[673,859,539,564]" country="Costa Rica" name="Limon" pageId="7" pageNumber="96">LimÓn province</collectingRegion>
,
<location id="FC78D5FABA6AFFE0FCD2493B59E5FDA2" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72:FC78D5FABA6AFFE0FCD2493B59E5FDA2" box="[872,978,539,564]" country="Costa Rica" county="Same" latitude="10.301375" longLatPrecision="1" longitude="-83.62352" municipality="Same" name="Guacimo" pageId="7" pageNumber="96" stateProvince="LimOn province">Guácimo</location>
,
<location id="FC78D5FABA6AFFE0FC65493B5E0DFDA5" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72:FC78D5FABA6AFFE0FC65493B5E0DFDA5" box="[991,1082,539,563]" country="Costa Rica" county="Same" latitude="10.301375" longLatPrecision="1" longitude="-83.62352" municipality="Same" name="Duacari" pageId="7" pageNumber="96" stateProvince="LimOn province">Duacarí</location>
,
<location id="FC78D5FABA6AFFE0FBF2493B5EEEFDA2" LSID="urn:lsid:plazi:treatment:710E3237BA6CFFE0FF2D4C085E7CFD72:FC78D5FABA6AFFE0FBF2493B5EEEFDA2" box="[1096,1241,539,564]" country="Costa Rica" county="Same" latitude="10.301375" longLatPrecision="1" longitude="-83.62352" municipality="Same" name="Rio Jimenez" pageId="7" pageNumber="96" stateProvince="LimOn province">Rio Jiménez</location>
,
<geoCoordinate id="9C93E5E6BA6AFFE0FB5D493B5FA5FDA2" box="[1255,1426,539,564]" direction="north" orientation="latitude" pageId="7" pageNumber="96" precision="1" value="10.301375">10°1804.95N</geoCoordinate>
/
<geoCoordinate id="9C93E5E6BA6AFFE0FF2D491E5B7CFDC1" box="[151,331,574,599]" direction="west" orientation="longitude" pageId="7" pageNumber="96" precision="1" value="-83.62352">83°3724.67W</geoCoordinate>
, 45 meterS above Sea level,
<collectingDate id="9D5D5C09BA6AFFE0FD31491E5938FDC1" box="[651,783,574,599]" pageId="7" pageNumber="96" value="2017-06-15">
<date id="8D19A5E1BA6AFFE0FD31491E5938FDC1" box="[651,783,574,599]" pageId="7" pageNumber="96" value="2017-06-15">VI/15/2017</date>
</collectingDate>
,
<collectorName id="5452E6F7BA6AFFE0FCA6491F59F8FDC1" box="[796,975,574,599]" pageId="7" pageNumber="96">F. Sibaja-Araya</collectorName>
,
<collectorName id="5452E6F7BA6AFFE0FC61491F5E9CFDC1" box="[987,1195,575,599]" pageId="7" pageNumber="96">M. Guevara-Mora</collectorName>
,
<collectorName id="5452E6F7BA6AFFE0FB0D491F5FAEFDC1" box="[1207,1433,575,599]" pageId="7" pageNumber="96">D. Romero-Serrano</collectorName>
, reSpectively depoSited at
<collectionCode id="9FB61BE4BA6AFFE0FE784941582EFDEC" box="[450,537,609,634]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FE784941582EFDEC" bold="true" box="[450,537,609,634]" pageId="7" pageNumber="96">MNCR</emphasis>
</collectionCode>
(two mature nymphS),
<collectionCode id="9FB61BE4BA6AFFE0FC8E494159A1FDEC" box="[820,918,609,634]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FC8E494159A1FDEC" bold="true" box="[820,918,609,634]" pageId="7" pageNumber="96">LEUNA</emphasis>
</collectionCode>
(two mature nymphS),
<emphasis id="CBD35F33BA6AFFE0FB0B49415F20FDEC" bold="true" box="[1201,1303,609,634]" pageId="7" pageNumber="96">
<collectionCode id="9FB61BE4BA6AFFE0FB0B49415F35FDEC" box="[1201,1282,609,634]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="96">MNRJ</collectionCode>
(
</emphasis>
two mature nymphS), and
<emphasis id="CBD35F33BA6AFFE0FE8C49A45BA1FD0B" bold="true" box="[310,406,644,669]" pageId="7" pageNumber="96">
<collectionCode id="9FB61BE4BA6AFFE0FE8C49A45BBFFD0B" box="[310,392,644,669]" pageId="7" pageNumber="96">FAMU</collectionCode>
(
</emphasis>
two mature nymphS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1BDD0AABA6AFFE0FF7D49875E7CFD72" pageId="7" pageNumber="96" type="etymology">
<paragraph id="F9188321BA6AFFE0FF7D49875E7CFD72" blockId="7.[151,1437,151,740]" pageId="7" pageNumber="96">
<emphasis id="CBD35F33BA6AFFE0FF7D49875B7FFD56" bold="true" box="[199,328,679,704]" pageId="7" pageNumber="96">Etymology</emphasis>
: The name of thiS SpecieS honorS the indigenouS king Guaycara who Several centurieS ago ruled the Boruca people in Southern
<collectingCountry id="81B0C3B1BA6AFFE0FE5449EB585FFD72" box="[494,616,715,740]" name="Costa Rica" pageId="7" pageNumber="96">CoSta Rica</collectingCountry>
where thiS new SpecieS waS firSt collected.
</paragraph>
</subSubSection>
</treatment>
</document>