196 lines
18 KiB
XML
196 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.443.7880" ID-GBIF-Dataset="33ef48a3-7145-4103-9159-e11be2ba9adc" ID-PMC="PMC4205504" ID-Pensoft-Pub="1313-2970-443-61" ID-PubMed="25349497" ID-ZBK="E33841396A6E426C840D85BC32A12E78" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-443-61" ModsDocOrigin="ZooKeys 443" ModsDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" checkinTime="1451245273153" checkinUser="pensoft" docAuthor="Sokolov, Igor M. & Kavanaugh, David H." docDate="2014" docId="42047F2AF5AD8D98346134D0B09BB0AA" docLanguage="en" docName="ZooKeys 443: 61-118" docOrigin="ZooKeys 443" docSource="http://dx.doi.org/10.3897/zookeys.443.7880" docTitle="Geocharidius lencanus Sokolov & Kavanaugh, 2014, sp. n." docType="treatment" docUuid="B41C1418-E0FB-4896-BE0F-827FE08A84D1" docUuidSource="ZooBank" docVersion="4" lastPageNumber="86" masterDocId="3546FFA8CF78F064801706227F76A615" masterDocTitle="The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation" masterLastPageNumber="118" masterPageNumber="61" pageNumber="84" updateTime="1668159453511" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>443</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>61</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.443.7880</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.443.7880</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-443-61</mods:identifier>
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<mods:identifier type="ZBK">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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<mods:identifier type="ZooBank">E33841396A6E426C840D85BC32A12E78</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152055698" LSID="urn:lsid:zoobank.org:act:B41C1418-E0FB-4896-BE0F-827FE08A84D1" httpUri="http://treatment.plazi.org/id/42047F2AF5AD8D98346134D0B09BB0AA" lastPageId="25" lastPageNumber="86" pageId="23" pageNumber="84">
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<subSubSection pageId="23" pageNumber="84" type="multiple">
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<paragraph pageId="23" pageNumber="84">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="84" type="nomenclature">
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<paragraph pageId="23" pageNumber="84">
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<taxonomicName LSID="http://zoobank.org/B41C1418-E0FB-4896-BE0F-827FE08A84D1" class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="23" pageNumber="84" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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<taxonomicNameLabel pageId="23" pageNumber="84">sp. n.</taxonomicNameLabel>
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Figs 5C, 6G, 18B, 19
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<normalizedToken originalValue="H–J">H-J</normalizedToken>
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, 20C, 21C, 22, 23
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="85" type="type material">
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<paragraph pageId="24" pageNumber="85">
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<pageBreakToken pageId="24" pageNumber="85" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="24" pageNumber="85">
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HOLOTYPE, a male, in KUNHM, point-mounted, dissected, labeled: \ HONDURAS: Lempira Dept., P.N. Celaque, nr. Gracias, Campamiento Naranjo,
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="14.545">14°32.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-88.66167">88°39.7'W</geoCoordinate>
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, 2500m, 12-13-V-2002, cloud forest litter R. Anderson, RSA2002-020 \ SM0
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<normalizedToken originalValue="…">...</normalizedToken>
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KUNHM-ENT \ HOLOTYPE
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="24" pageNumber="85" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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Sokolov and Kavanaugh 2014 [red label] \. PARATYPES: A total of 6 specimens (3 males and 2 females were dissected), deposited in CAS, CMNC and KUNHM; 4 specimens labeled same as holotype; 2 specimens labeled: \ HONDURAS: Lempira Dept., P.N. Celaque, nr. Gracias, Campamiento Naranjo, 2500m, 14°32.7', N
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-88.66167">88°39.7'W</geoCoordinate>
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, 12-13.V.2002, cloud forest litter R. Anderson, 2002-020E \ CMNC \.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="85" type="type locality">
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<paragraph pageId="24" pageNumber="85">Type locality.</paragraph>
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<paragraph pageId="24" pageNumber="85">Honduras, Lempira Department, Celaque National Park.</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="85" type="etymology">
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<paragraph pageId="24" pageNumber="85">Etymology.</paragraph>
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<paragraph pageId="24" pageNumber="85">
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The specific epithet is a Latinized adjective in the masculine form based on the name of the indigenous people, the
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<taxonomicName genus="Lenca" lsidName="Lenca" pageId="24" pageNumber="85" rank="genus">Lenca</taxonomicName>
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, living in the territory of Celaque National Park during historic times.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="86" type="recognition">
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<paragraph pageId="25" pageNumber="86">
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<pageBreakToken pageId="25" pageNumber="86" start="start">Recognition</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="25" pageNumber="86">
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Externally, members of this species represent a larger version of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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adults, described above. Adults of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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are distinguished from those of other members of the integripennis species group by the following combination of external characters: medium to large size, rather wide habitus and fully microsculptured dorsal body surface. Males and females of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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are distinguished from those of the other members of the integripennis species group by the structure of the median lobe and the shape of spermatheca, respectively.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="86" type="description">
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<paragraph pageId="25" pageNumber="86">Description.</paragraph>
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<paragraph pageId="25" pageNumber="86">
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Size. Medium to large for genus (SBL range 1.30-1.47 mm, mean 1.39
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.091mm, n=4).
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</paragraph>
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<paragraph pageId="25" pageNumber="86">
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Habitus. Body form (Fig. 18B) moderately convex, ovoid, general proportions (WE/SBL 0.40
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.006), proportions of head (WH/WPm 0.73
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.008) and pronotum (WPm/WE 0.77
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.010) average for group.
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</paragraph>
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<paragraph pageId="25" pageNumber="86">Color. Body rufotestaceous, appendages testaceous.</paragraph>
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<paragraph pageId="25" pageNumber="86">Microsculpture. Mesh pattern of irregularly isodiametric sculpticells present over all dorsal surfaces of head, pronotum and elytra. Proepisternum also with evident microsculpture.</paragraph>
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<paragraph pageId="25" pageNumber="86">Mouthparts. Maxillae and labium (Fig. 5C).</paragraph>
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<paragraph pageId="25" pageNumber="86">
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Prothorax. Pronotum moderately transverse (WPm/LP 1.28
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.009), with lateral margins moderately constricted posteriorly (WPm/WPp 1.34
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.019). Posterior angles slightly obtuse (100-110°). Width between posterior angles nearly equal to the width between anterior angles (WPa/WPp 1.01
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.012).
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</paragraph>
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<paragraph pageId="25" pageNumber="86">Legs. Protibia (Fig. 6G).</paragraph>
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<paragraph pageId="25" pageNumber="86">
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Elytra. Moderately convex, slightly depressed along suture, markedly wide (WE/LE 0.69
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.013), without traces of striae. Humeri rounded, in outline forming right angle with longitudinal axis of body. Lateral margins convex, evenly divergent in basal fourth, evenly rounded to apex in apical third.
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</paragraph>
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<paragraph pageId="25" pageNumber="86">Male genitalia. Median lobe (Fig. 19H) with shaft subparallel with a long attenuated preapical part, apex small and narrowly rounded. Ventral margin almost straight. Dorsal sclerites of internal sac in form of a long narrow fig, flagellum-like in apical two-thirds, and slightly dilated and curved dorsally in basal third. Right paramere with long and narrow apical constriction (Fig. 19J). Left paramere with long and narrow apical constriction (Fig. 19I). Ring sclerite with handle rectangularly rounded (Fig. 20C).</paragraph>
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<paragraph pageId="25" pageNumber="86">Female internal genitalia. Spermatheca sclerotized, fusiform, slightly dilated and rectangularly bent apically, with short cornu and long nodulus (Fig. 21C). Length of spermathecal gland less than length of spermatheca. Spermathecal duct not coiled.</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="86" type="geographical distribution">
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<paragraph pageId="25" pageNumber="86">Geographical distribution.</paragraph>
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<paragraph pageId="25" pageNumber="86">This species is known only from Celaque National Park, in the Cerro las Minas range of Honduras (Fig. 22, yellow diamond).</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="86" type="way of life">
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<paragraph pageId="25" pageNumber="86">Way of life.</paragraph>
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<paragraph pageId="25" pageNumber="86">Specimens were collected by sifting cloud forest litter at an elevation of 2500 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="86" type="relationships">
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<paragraph pageId="25" pageNumber="86">Relationships.</paragraph>
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<paragraph pageId="25" pageNumber="86">
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The rectangular shape of the handle of the ring sclerite (Fig. 20C) in males and the shape of spermatheca (Fig. 21C) in females suggest a close relationship with
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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(Figs 20B, 21B), described above.
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</paragraph>
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<caption pageId="25" pageNumber="86">
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<paragraph pageId="25" pageNumber="86">
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Figure 22. Map of southern Mexico, Guatemala and adjacent part of Honduras, showing positions of locality records for the species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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: white diamond,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius andersoni" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="andersoni">Geocharidius andersoni</taxonomicName>
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; white circles (black point in a circle shows "terra typica" for the species),
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius zullinii" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="zullinii">Geocharidius zullinii</taxonomicName>
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; white squares,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius vignatagliantii" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="vignatagliantii">Geocharidius vignatagliantii</taxonomicName>
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; white triangle,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius gimlii" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="gimlii">Geocharidius gimlii</taxonomicName>
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; black and white triangle,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius integripennis" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="integripennis">Geocharidius integripennis</taxonomicName>
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; green squares,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius longinoi" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="longinoi">Geocharidius longinoi</taxonomicName>
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; green circles,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius erwini" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="erwini">Geocharidius erwini</taxonomicName>
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; green flowers,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius minimus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="minimus">Geocharidius minimus</taxonomicName>
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; green diamonds,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius balini" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="balini">Geocharidius balini</taxonomicName>
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; green triangle,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius jalapensis" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="jalapensis">Geocharidius jalapensis</taxonomicName>
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; yellow quadrangle,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius antigua" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="antigua">Geocharidius antigua</taxonomicName>
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; yellow diamond,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius lencanus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="lencanus">Geocharidius lencanus</taxonomicName>
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; yellow flower,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius celaquensis" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="celaquensis">Geocharidius celaquensis</taxonomicName>
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; yellow triangles,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius disjunctus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="disjunctus">Geocharidius disjunctus</taxonomicName>
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; yellow circles,
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius comayaguanus" order="Coleoptera" pageId="25" pageNumber="86" phylum="Arthropoda" rank="species" species="comayaguanus">Geocharidius comayaguanus</taxonomicName>
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. Physiographic features: CH, Chiapas Highlands; GC, Guatemalan Cordillera; HIH, Honduran Interior Highlands; MTZ, Motagua Fault Zone; SC, Sierra de los Cuchumatanes; SMC, Sierra Madre de Chiapas. Elevation scale bar is given in meters.
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</paragraph>
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</caption>
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<caption pageId="25" pageNumber="86">
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<paragraph pageId="25" pageNumber="86">Figures 23. Diagrams illustrating size variation for sympatric pairs of closely and remotely related species of the integripennis group at different geographical localities of Nuclear Central America. Closely related pairs (for shared characters see subchapters on Relationships in the text for corresponding species): A Chiapas Highlands, Mexico B Mataquescuintla, Guatemala C Celaque National Park, Honduras. Remotely related species: D Volcano Agua, Guatemala E Volcano Atitlan, Guatemala F Pico Pijol National Park, Honduras. Legend: black dot - median; box - 25-75% range of values; whiskers - non-outlier range of values; b - coefficient of regression.</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |