treatments-xml/data/38/F0/06/38F0069FA0FE542C9F4015B5339F2B44.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="38F0069FA0FE542C9F4015B5339F2B44" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila cavicola Pall-Gergely &amp; Dumrongrojwattana 2023, sp. nov." docType="treatment" docUuid="4EEC485D-638D-4136-95BF-340F508B04E8" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1147</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:4EEC485D-638D-4136-95BF-340F508B04E8" httpUri="http://treatment.plazi.org/id/38F0069FA0FE542C9F4015B5339F2B44" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/4EEC485D-638D-4136-95BF-340F508B04E8" authority="Páll-Gergely &amp; Dumrongrojwattana" authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila cavicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cavicola" status="sp. nov.">
Angustopila cavicola 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Dumrongrojwattana
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Angustopila cavicola Pall-Gergely &amp; Dumrongrojwattana, sp. nov. (holotype, CUMZ 7441). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; microstructure of the protoconch showing protoconch-teleoconch boundary (E), eroded ventral (F) and frontal (G) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure30" httpUri="https://binary.pensoft.net/fig/808839" pageId="0" pageNumber="1">Figs 30</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Variability of Angustopila cavicola Pall-Gergely &amp; Dumrongrojwattana, sp. nov. A holotype (CUMZ 7441) B, C paratypes (HNHM 100181)." figureDoi="10.3897/zookeys.1147.93824.figure32" httpUri="https://binary.pensoft.net/fig/808841" pageId="0" pageNumber="1">, 32</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Angustopila cavicola Pall-Gergely &amp; Dumrongrojwattana, sp. nov. from Laos (2019 / 118). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; aperture (E), microstructure of the protoconch (F), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure33" httpUri="https://binary.pensoft.net/fig/808842" pageId="0" pageNumber="1">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Thailand • 1 empty shell (H: 1.18 mm, D: 0.92 mm); Loei Province, Phu Tham Pha Tang; approx. GPS coordinates: 
<geoCoordinate degrees="17" direction="north" minutes="56.91" orientation="latitude" precision="9" value="17.9485">17°56.91'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="55.35" orientation="longitude" precision="9" value="101.9225">101°55.35'E</geoCoordinate>
; 29 Jun. 2016; P. Dumrongrojwattana, P. Juangsantad, K. Khwantong, N. Namisa &amp; P. Panthong leg.; CUMZ 7441.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Thailand • 2 figured shells, same data as for holotype; HNHM 100181 • 8 shells; same data as for holotype; coll. PD • 8 shells; same data as for holotype; coll. PGB • 2 specimens in ethanol; same data as for holotype; coll. PGB.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Thailand" specimenCount="1">
<collectingCountry name="Thailand">Thailand</collectingCountry>
• 11 j/b shells; same data as for holotype; coll. PD
</materialsCitation>
. 
<materialsCitation collectingDate="2019-10-07" collectorName="A. Hunyadi" country="Laos" county="Na Mor" elevation="635" latitude="20.872833" location="Tham Nathong" longLatPrecision="12" longitude="101.783" municipality="Ban Nathong" specimenCount="1" stateProvince="Udomxai Province">
<collectingCountry name="Laos">Laos</collectingCountry>
• 1 shell; 
<collectingRegion country="Laos" name="Oudomxai">Udomxai Province</collectingRegion>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.5" unit="km" value="6.5">6.5 km</quantity>
southeast from centre of 
<collectingCounty>Na Mor</collectingCounty>
towards 
<collectingRegion country="Laos" name="Oudomxai">Udomxai</collectingRegion>
, 
<collectingMunicipality>Ban Nathong</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:38F0069FA0FE542C9F4015B5339F2B44:480ACBA5A0B9D3E5A4EB0A3B27E83495" country="Laos" county="Na Mor" latitude="20.872833" longLatPrecision="12" longitude="101.783" municipality="Ban Nathong" name="Tham Nathong" stateProvince="Udomxai Province">Tham Nathong</location>
, below cave spring (locality code: 2019/118); 
<geoCoordinate degrees="20" direction="north" minutes="52.37" orientation="latitude" precision="9" value="20.872833">20°52.37'N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="46.98" orientation="longitude" precision="9" value="101.783">101°46.98'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.35" unit="m" value="635.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.35" unit="m" value="635.0">635 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2019-10-07">7 Oct. 2019</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized to large, conical 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with few whorls, a narrow umbilicus, a conspicuously large aperture, a strongly expanded peristome and a short but strong parietal tooth that points slightly towards the palatal wall.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description (of the type sample).</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell medium to large-sized for the genus, higher to much higher than wide; off-white, conical, apex broad, body whorl widest from apertural view; protoconch consists of 1.25 whorls, without spiral striation; teleoconch finely ornamented with irregular radial growth lines crossed by weak rows of irregularly spaced spiral threads (ca. 12-14 on body whorl from apertural view); sculpture weaker than in most other 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species; whorls 3.5-4 (conspicuously few compared to large shell size), rounded; aperture slightly oblique to shell axis from lateral view; umbilicus very narrow; aperture subcircular, parietal callus convex; peristome strongly expanded especially at parietal and palatal region, not reflected; parietal callus overlapping and not smeared onto penultimate whorl; parietal tooth elevated, strong, normally long, slightly bent towards palatal side, does not reach parietal callus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 1.02-1.18, D = 0.87-0.97, H/D*100 = 109.3-130.3 (shells from the type locality; 
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 6), RUD = 17.6-19.8 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3); H = 1.27, D = 0.99, H/D*100 = 128.3, RUD = 20.2 (2019/118; 
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila fabella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fabella">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila fabella</emphasis>
</taxonomicName>
is smaller, has a more pointed apex, a comparatively smaller aperture, a weaker sculpture, and a less expanded peristome. All other conical species with a single tooth are much smaller than 
<taxonomicName lsidName="A. cavicola" pageId="0" pageNumber="1" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">A. cavicola</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This new species was collected inside a cave, in the dark zone (
<taxonomicName authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila cavicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila cavicola</emphasis>
</taxonomicName>
means cave-dweller in Latin).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
The type series is known from Loei Province, Thailand. An additional, single shell, reminiscent of the typical 
<taxonomicName lsidName="A. cavicola" pageId="0" pageNumber="1" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">A. cavicola</emphasis>
</taxonomicName>
sp. nov. shells, was collected in Northern Laos ca. 325 km north of the type series (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure31" httpUri="https://binary.pensoft.net/fig/808840" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure30" httpUri="https://binary.pensoft.net/fig/808839" pageId="0" pageNumber="1" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 30.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila cavicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila cavicola</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Dumrongrojwattana, sp. nov. (holotype, CUMZ 7441). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and apical (D) sides of the shell; microstructure of the protoconch showing protoconch-teleoconch boundary (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), eroded ventral (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure31" httpUri="https://binary.pensoft.net/fig/808840" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
Distribution of 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure32" httpUri="https://binary.pensoft.net/fig/808841" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
Variability of 
<taxonomicName authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila cavicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila cavicola</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Dumrongrojwattana, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype (CUMZ 7441) 
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
paratypes (HNHM 100181).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure33" httpUri="https://binary.pensoft.net/fig/808842" pageId="0" pageNumber="1" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 33.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Dumrongrojwattana" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila cavicola" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cavicola">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila cavicola</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Dumrongrojwattana, sp. nov. from Laos (2019/118). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; aperture (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), microstructure of the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">The single shell from Laos has an even weaker sculpture (practically smooth) and a lower parietal tooth than the Thai ones. Otherwise, it is similar to the Thai population in terms of shell size, and shell and aperture shapes.</paragraph>
</subSubSection>
</treatment>
</document>