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<mods:title id="756404A60D07B36BF9A1EDEED1B579A5">A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island</mods:title>
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<treatment id="312287F0B7250E388909E6C0FAFA4B8A" ID-DOI="http://doi.org/10.5281/zenodo.6309085" ID-GBIF-Taxon="193333176" ID-Zenodo-Dep="6309085" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A" httpUri="http://treatment.plazi.org/id/312287F0B7250E388909E6C0FAFA4B8A" lastPageNumber="37" pageId="36" pageNumber="37">
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<taxonomicName id="7E8B4D65B7250E388909E6C0FB904978" ID-CoL="65SV9" authority="(Budde-Lund, 1885)" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[513,1074,1278,1306]" class="Malacostraca" family="Agnaridae" genus="Agnara" kingdom="Animalia" order="Isopoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="8BFFEAF4B7250E388909E6C0FC98497B" bold="true" box="[513,826,1278,1305]" italics="true" pageId="36" pageNumber="37">Agnara madagascariensis</emphasis>
(
<bibRefCitation id="DD1A4B17B7250E388842E6C0FB884978" author="Budde-Lund G." box="[842,1066,1279,1306]" pageId="36" pageNumber="37" refId="ref21297" refString="Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen &amp; Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885</bibRefCitation>
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<paragraph id="B93436E6B7250E3889D4E718FCF54923" blockId="36.[513,1074,1278,1345]" box="[732,855,1319,1345]" pageId="36" pageNumber="37">
<figureCitation id="21B02A63B7250E3889D4E718FC874923" box="[732,805,1319,1345]" captionStart="Fig" captionStartId="3.[189,232,1142,1168]" captionTargetBox="[198,1394,269,1093]" captionTargetId="figure-182@3.[189,1399,265,1102]" captionTargetPageId="3" captionText="Fig. 1. Map of the study areas and distribution of species of terrestrial isopods from Isla Grande. 1. Ligia baudiniana Milne Edwards, 1840. 2. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov. 3. Tylos marcuzzii Giordani Soika, 1954. 4. Tylos niveus Budde-Lund, 1885. 5. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. 6. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989. 7. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. 8. Armadilloniscus ninae Schultz, 1984. 9. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. 10. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. 11. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). 12. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986.13. Trichorhina heterophthalma Lemos de Castro, 1964.14. Trichorhina bermudezae Carpio-Díaz, López-Orozco &amp; Campos-Filho, 2018. 15. Porcellionides pruinosus (Brandt, 1833). 16. Agnara madagascariensis (Budde-Lund, 1885). 17. Ctenorillo tuberosus (Budde-Lund, 1904)." figureDoi="http://doi.org/10.5281/zenodo.6037894" httpUri="https://zenodo.org/record/6037894/files/figure.png" pageId="36" pageNumber="37">Figs 1</figureCitation>
,
<figureCitation id="21B02A63B7250E38883BE718FCF54923" box="[819,855,1319,1345]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[296,1291,264,1755]" captionTargetId="figure-113@9.[296,1291,264,1755]" captionTargetPageId="9" captionText="Fig. 2. A. Tylos marcuzzii Giordani Soika, 1954.B. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos- Filho sp. nov. C. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. D. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989.E. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. F. Armadilloniscus ninae Schultz, 1984. G. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. H. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. I. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). J. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986. K. Agnara madagascariensis (Budde-Lund, 1885). L. Ctenorillo tuberosus (Budde-Lund, 1904). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6037898" httpUri="https://zenodo.org/record/6037898/files/figure.png" pageId="36" pageNumber="37">2K</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F191656DB7250E388BB5E751FC8D4A61" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="B93436E6B7250E388BB5E751FCD549EA" blockId="36.[189,887,1390,1417]" box="[189,887,1390,1417]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="999B11C8B7250E388BB5E751FCD549EA" box="[189,887,1390,1417]" pageId="36" pageNumber="37">
<treatmentCitation id="382A10F7B7250E388BB5E751FCD049EA" author="Budde-Lund G." box="[189,882,1390,1417]" page="189" pageId="36" pageNumber="37" year="1885">
<taxonomicName id="7E8B4D65B7250E388BB5E751FCD049EA" ID-CoL="42HJJ" authority="Budde-Lund, 1885: 189" authorityName="Budde-Lund" authorityPageNumber="189" authorityYear="1885" box="[189,882,1390,1417]" class="Malacostraca" family="Agnaridae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="8BFFEAF4B7250E388BB5E751FDED49EA" box="[189,591,1390,1416]" italics="true" pageId="36" pageNumber="37">Metoponorthus Madagascariensis</emphasis>
<bibRefCitation id="DD1A4B17B7250E38895EE751FCD049EA" author="Budde-Lund G." box="[598,882,1390,1417]" pageId="36" pageNumber="37" refId="ref21297" refString="Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen &amp; Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885: 189</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B93436E6B7250E388BB5E78BFE6849AC" blockId="36.[189,458,1460,1486]" box="[189,458,1460,1486]" pageId="36" pageNumber="37">
<heading id="E27C818AB7250E388BB5E78BFE6849AC" bold="true" box="[189,458,1460,1486]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="8BFFEAF4B7250E388BB5E78BFE6849AC" bold="true" box="[189,458,1460,1486]" pageId="36" pageNumber="37">Additional references</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7250E388BB5E7D7FC8D4A61" blockId="36.[189,815,1512,1539]" box="[189,815,1512,1539]" pageId="36" pageNumber="37">
<bibRefCitation id="DD1A4B17B7250E388BB5E7D7FDB54A60" author="Schmalfuss H. &amp; Ferrara F." box="[189,535,1512,1539]" pageId="36" pageNumber="37" pagination="15 - 9" refId="ref25187" refString="Schmalfuss H. &amp; Ferrara F. 1978. Terrestrial isopods from West Africa, Part 2: families Tylidae, Ligiidae, Trichoniscidae, Styloniscidae, Rhyscotidae, Halophilosciidae, Philosciidae, Platyarthridae, Rhyscotidae, Trachelipidae, Porcellionidae, Armadillidiidae. Monitore zoologico italiano, nuova serie, Supplemento 11: 15 - 9. https: // doi. org / 10.1080 / 03749444.1978.10736575" type="journal article" year="1978">Schmalfuss &amp; Ferrara (1978)</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7250E38892BE7D6FC8E4A61" author="Ferrara F. &amp; Taiti S." box="[547,812,1512,1539]" pageId="36" pageNumber="37" pagination="93 - 121" refId="ref22101" refString="Ferrara F. &amp; Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121." type="journal article" year="1986">Ferrara &amp; Taiti (1986)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F191656DB7250E388BB5E411FDC04B24" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="B93436E6B7250E388BB5E411FE084A2A" blockId="36.[189,426,1582,1608]" box="[189,426,1582,1608]" pageId="36" pageNumber="37">
<heading id="E27C818AB7250E388BB5E411FE084A2A" bold="true" box="[189,426,1582,1608]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="8BFFEAF4B7250E388BB5E411FE084A2A" bold="true" box="[189,426,1582,1608]" pageId="36" pageNumber="37">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7250E388BB5E45DFDC04B24" blockId="36.[189,1399,1633,1862]" pageId="36" pageNumber="37">
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–
<emphasis id="8BFFEAF4B7250E388A73E45EFBD14A1E" bold="true" box="[379,1139,1633,1660]" pageId="36" pageNumber="37">
<collectingRegion id="7B4FF804B7250E388A73E45EFE754A1E" box="[379,471,1633,1660]" pageId="36" pageNumber="37">Bolívar</collectingRegion>
,
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,
<location id="BC54603DB7250E3889F1E45DFC6C4A1E" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A:BC54603DB7250E3889F1E45DFC6C4A1E" box="[761,974,1634,1660]" country="Colombia" latitude="10.177517" longLatPrecision="1" longitude="-75.74733" municipality="Cano Raton" name="Islas del Rosario" pageId="36" pageNumber="37" stateProvince="Bolivar">Islas del Rosario</location>
,
<location id="BC54603DB7250E3888D4E45DFBD14A1E" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A:BC54603DB7250E3888D4E45DFBD14A1E" box="[988,1139,1634,1660]" country="Colombia" latitude="10.177517" longLatPrecision="1" longitude="-75.74733" municipality="Cano Raton" name="Isla Grande" pageId="36" pageNumber="37" stateProvince="Bolivar">Isla Grande</location>
</emphasis>
•
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,
<specimenCount id="AF8DFD6FB7250E388FEBE45DFA8A4A1F" box="[1251,1320,1634,1661]" pageId="36" pageNumber="37" type="female">6 ♀♀</specimenCount>
;
<collectingMunicipality id="5950AC9CB7250E388E31E45DFEA44AC7" pageId="36" pageNumber="37">Caño Ratón</collectingMunicipality>
,
<location id="BC54603DB7250E388A1CE4B4FE644AC6" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A:BC54603DB7250E388A1CE4B4FE644AC6" box="[276,454,1674,1701]" country="Colombia" latitude="10.177517" longLatPrecision="1" longitude="-75.74733" municipality="Cano Raton" name="Casa Comunal" pageId="36" pageNumber="37" stateProvince="Bolivar">Casa Comunal</location>
;
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;
<date id="CD351026B7250E38886BE4B4FC554AC7" box="[867,1015,1675,1701]" pageId="36" pageNumber="37" value="2017-09-08">
<collectingDate id="DD71E9CEB7250E38886BE4B4FC554AC7" box="[867,1015,1675,1701]" pageId="36" pageNumber="37" value="2017-09-08">8 Sep. 2017</collectingDate>
</date>
;
<collectorName id="147E5330B7250E388F0EE4B4FB5A4AC7" box="[1030,1272,1675,1701]" pageId="36" pageNumber="37">C.M. López-Orozco</collectorName>
,
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.
<collectorName id="147E5330B7250E388E24E4B5FEB14AAF" pageId="36" pageNumber="37">Borja- Arrieta</collectorName>
,
<collectorName id="147E5330B7250E388A2BE48CFE594AAF" box="[291,507,1715,1741]" pageId="36" pageNumber="37">Y.M. Carpio-Díaz</collectorName>
and
<collectorName id="147E5330B7250E388934E48CFD6F4AAF" box="[572,717,1715,1741]" pageId="36" pageNumber="37">A. Figueroa</collectorName>
leg.;
<specimenCode id="E92D9E9DB7250E388812E48CFC484AAF" box="[794,1002,1715,1741]" collectionCode="CUDC-CRU" pageId="36" pageNumber="37">CUDC-CRU 138</specimenCode>
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•
<materialsCitation id="09E33CBBB7250E388F06E48CFC1B4A97" ID-GBIF-Occurrence="3464759317" collectingDate="2017-09-08" collectionCode="CUDC-CRU" collectorName="A. Figueroa" country="Colombia" latitude="10.177" location="Orika" longLatPrecision="1" longitude="-75.741" pageId="36" pageNumber="37" specimenCode="CUDC-CRU 139" specimenCount="2" specimenCount-female="2" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">
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<location id="BC54603DB7250E388F62E48CFB134AAF" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A:BC54603DB7250E388F62E48CFB134AAF" box="[1130,1201,1715,1741]" country="Colombia" latitude="10.177" longLatPrecision="1" longitude="-75.741" name="Orika" pageId="36" pageNumber="37" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Orika</location>
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;
<date id="CD351026B7250E388A80E4E4FDB54A97" box="[392,535,1755,1781]" pageId="36" pageNumber="37" value="2017-09-08">
<collectingDate id="DD71E9CEB7250E388A80E4E4FDB54A97" box="[392,535,1755,1781]" pageId="36" pageNumber="37" value="2017-09-08">8 Sep. 2017</collectingDate>
</date>
;
<collectorName id="147E5330B7250E388929E4E4FD0F4A97" box="[545,685,1755,1781]" pageId="36" pageNumber="37">A. Figueroa</collectorName>
leg.;
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•
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<location id="BC54603DB7250E388E00E4E4FEBB4B7F" LSID="urn:lsid:plazi:treatment:312287F0B7250E388909E6C0FAFA4B8A:BC54603DB7250E388E00E4E4FEBB4B7F" country="Colombia" latitude="10.179698" longLatPrecision="1" longitude="-75.729645" name="Casa Sra. Miriam" pageId="36" pageNumber="37" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Casa Sra. Miriam</location>
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,
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;
<date id="CD351026B7250E3889BBE53BFCE24B7C" box="[691,832,1796,1822]" pageId="36" pageNumber="37" value="2017-09-08">
<collectingDate id="DD71E9CEB7250E3889BBE53BFCE24B7C" box="[691,832,1796,1822]" pageId="36" pageNumber="37" value="2017-09-08">8 Sep. 2017</collectingDate>
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;
<collectorName id="147E5330B7250E388844E53BFB994B7C" box="[844,1083,1796,1822]" pageId="36" pageNumber="37">C.M. López-Orozco</collectorName>
,
<collectorName id="147E5330B7250E388F4EE53BFAA44B7F" box="[1094,1286,1795,1822]" pageId="36" pageNumber="37">R. Borja-Arrieta</collectorName>
and
<collectorName id="147E5330B7250E388E34E53BFEEC4B24" pageId="36" pageNumber="37">Y.M. Carpio-Díaz</collectorName>
leg.;
<specimenCode id="E92D9E9DB7250E388A98E513FDFC4B24" box="[400,606,1836,1862]" collectionCode="CUDC-CRU" pageId="36" pageNumber="37">CUDC-CRU 140</specimenCode>
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.
</paragraph>
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<subSubSection id="F191656DB7250E388BB5E54DFAFA4B8A" pageId="36" pageNumber="37" type="distribution">
<paragraph id="B93436E6B7250E388BB5E54DFEF44BEE" blockId="36.[189,342,1906,1932]" box="[189,342,1906,1932]" pageId="36" pageNumber="37">
<heading id="E27C818AB7250E388BB5E54DFEF44BEE" bold="true" box="[189,342,1906,1932]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="8BFFEAF4B7250E388BB5E54DFEF44BEE" bold="true" box="[189,342,1906,1932]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7250E388BB5E599FAFA4B8A" blockId="36.[188,1398,1958,2024]" pageId="36" pageNumber="37">
<collectingCountry id="C19C7676B7250E388BB5E599FE2A4BA2" box="[189,392,1958,1984]" name="Ascension Island" pageId="36" pageNumber="37">Ascension Island</collectingCountry>
,
<collectingCountry id="C19C7676B7250E388A9BE599FE514BA2" box="[403,499,1958,1984]" name="Senegal" pageId="36" pageNumber="37">Senegal</collectingCountry>
,
<collectingCountry id="C19C7676B7250E388AF5E599FDF04BA2" box="[509,594,1958,1984]" name="Guinea-Bissau" pageId="36" pageNumber="37">Guinea</collectingCountry>
<collectingRegion id="7B4FF804B7250E388950E599FD0B4BA2" box="[600,681,1958,1984]" country="Guinea-Bissau" name="Bissau" pageId="36" pageNumber="37">Bissau</collectingRegion>
, Arabian Peninsula,
<collectingCountry id="C19C7676B7250E38889FE599FB854BA2" box="[919,1063,1958,1984]" name="Madagascar" pageId="36" pageNumber="37">Madagascar</collectingCountry>
, southern
<collectingCountry id="C19C7676B7250E388F95E599FB444BA2" box="[1181,1254,1958,1984]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
,
<collectingCountry id="C19C7676B7250E388FF8E599FAE44BA2" box="[1264,1350,1958,1984]" name="Taiwan" pageId="36" pageNumber="37">Taiwan</collectingCountry>
and
<collectingCountry id="C19C7676B7250E388BB4E5F1FE974B8A" box="[188,309,1998,2024]" name="Venezuela" pageId="36" pageNumber="37">Venezuela</collectingCountry>
(
<bibRefCitation id="DD1A4B17B7250E388A4CE5F1FD214B8A" author="Ashmole P. &amp; Ashmole M." box="[324,643,1998,2024]" pageId="36" pageNumber="37" refId="ref21176" refString="Ashmole P. &amp; Ashmole M. 2000. St Helena and Ascension Island: a Natural History. Anthony Nelson, Oswestry, England." type="book" year="2000">Ashmole &amp; Ashmole 2000</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7250E388987E5F1FC944B8A" author="Schmidt C." box="[655,822,1998,2024]" pageId="36" pageNumber="37" pagination="1 - 12" refId="ref25296" refString="Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12." type="journal article" year="2001">Schmidt 2001</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7250E38884AE5F1FBB34B8A" author="Schmalfuss H." box="[834,1041,1998,2024]" pageId="36" pageNumber="37" pagination="1 - 341" refId="ref25160" refString="Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341." type="journal article" year="2003">Schmalfuss 2003</bibRefCitation>
). First record for
<collectingCountry id="C19C7676B7250E388FD5E5F1FAF74B8A" box="[1245,1365,1998,2024]" name="Colombia" pageId="36" pageNumber="37">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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