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López-Orozco, Carpio-Díaz &amp; Campos-Filho
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<figureCitation id="21B02A63B70B0E1689C8E370FCAB4D08" box="[704,777,335,362]" captionStart="Fig" captionStartId="3.[189,232,1142,1168]" captionTargetBox="[198,1394,269,1093]" captionTargetId="figure-182@3.[189,1399,265,1102]" captionTargetPageId="3" captionText="Fig. 1. Map of the study areas and distribution of species of terrestrial isopods from Isla Grande. 1. Ligia baudiniana Milne Edwards, 1840. 2. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov. 3. Tylos marcuzzii Giordani Soika, 1954. 4. Tylos niveus Budde-Lund, 1885. 5. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. 6. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989. 7. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. 8. Armadilloniscus ninae Schultz, 1984. 9. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. 10. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. 11. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). 12. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986.13. Trichorhina heterophthalma Lemos de Castro, 1964.14. Trichorhina bermudezae Carpio-Díaz, López-Orozco &amp; Campos-Filho, 2018. 15. Porcellionides pruinosus (Brandt, 1833). 16. Agnara madagascariensis (Budde-Lund, 1885). 17. Ctenorillo tuberosus (Budde-Lund, 1904)." figureDoi="http://doi.org/10.5281/zenodo.6037894" httpUri="https://zenodo.org/record/6037894/files/figure.png" pageId="10" pageNumber="11">Figs 1</figureCitation>
,
<figureCitation id="21B02A63B70B0E16881FE36FFC9E4D08" box="[791,828,336,362]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[296,1291,264,1755]" captionTargetId="figure-113@9.[296,1291,264,1755]" captionTargetPageId="9" captionText="Fig. 2. A. Tylos marcuzzii Giordani Soika, 1954.B. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos- Filho sp. nov. C. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. D. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989.E. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. F. Armadilloniscus ninae Schultz, 1984. G. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. H. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. I. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). J. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986. K. Agnara madagascariensis (Budde-Lund, 1885). L. Ctenorillo tuberosus (Budde-Lund, 1904). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6037898" httpUri="https://zenodo.org/record/6037898/files/figure.png" pageId="10" pageNumber="11">2B</figureCitation>
,
<figureCitation id="21B02A63B70B0E168840E370FCD64D08" box="[840,884,335,362]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[189,232,1891,1917]" captionStartId-1="12.[189,232,1961,1987]" captionStartId-2="13.[189,232,1227,1253]" captionTargetBox-0="[194,1373,756,1837]" captionTargetBox-1="[202,1389,904,1903]" captionTargetBox-2="[201,1386,280,1164]" captionTargetId-0="figure-192@11.[189,1399,742,1850]" captionTargetId-1="figure-203@12.[189,1399,888,1914]" captionTargetId-2="figure-122@13.[189,1399,265,1186]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." captionText-1="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." captionText-2="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi-0="http://doi.org/10.5281/zenodo.6037900" figureDoi-1="http://doi.org/10.5281/zenodo.6037904" figureDoi-2="http://doi.org/10.5281/zenodo.6037906" httpUri-0="https://zenodo.org/record/6037900/files/figure.png" httpUri-1="https://zenodo.org/record/6037904/files/figure.png" httpUri-2="https://zenodo.org/record/6037906/files/figure.png" pageId="10" pageNumber="11">3–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F191656DB70B0E168BB5E3AEFCB14D9F" pageId="10" pageNumber="11" type="etymology">
<paragraph id="B93436E6B70B0E168BB5E3AEFEE74DC9" blockId="10.[189,325,401,427]" box="[189,325,401,427]" pageId="10" pageNumber="11">
<heading id="E27C818AB70B0E168BB5E3AEFEE74DC9" bold="true" box="[189,325,401,427]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="8BFFEAF4B70B0E168BB5E3AEFEE74DC9" bold="true" box="[189,325,401,427]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E380FCB14D9F" blockId="10.[189,1398,447,509]" pageId="10" pageNumber="11">The new species is named after Mr Hernando Gómez Molina, also known as Mr Negro, for his efforts to conserve the biodiversity of the Rosario Islands.</paragraph>
</subSubSection>
<subSubSection id="F191656DB70B0E168BB5E01BFA954818" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="B93436E6B70B0E168BB5E01BFE0F4E5C" blockId="10.[189,429,548,574]" box="[189,429,548,574]" pageId="10" pageNumber="11">
<heading id="E27C818AB70B0E168BB5E01BFE0F4E5C" bold="true" box="[189,429,548,574]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="8BFFEAF4B70B0E168BB5E01BFE0F4E5C" bold="true" box="[189,429,548,574]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B70B0E168BE2E06CFDAB4EB5" blockId="10.[189,1399,595,727]" pageId="10" pageNumber="11">
<materialsCitation id="09E33CBBB70B0E168BE2E06CFDAB4EB5" ID-GBIF-Occurrence="3464759411" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; R. Borja-Arrieta &amp; Y. M. Carpio- Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Bolivar" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 127" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="8BFFEAF4B70B0E168BE2E06CFEFF4E0F" bold="true" box="[234,349,595,621]" pageId="10" pageNumber="11">
<typeStatus id="66308844B70B0E168BE2E06CFEFF4E0F" box="[234,349,595,621]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="C19C7676B70B0E168BB5E049FEFA4EF2" box="[189,344,630,656]" name="Colombia" pageId="10" pageNumber="11">COLOMBIA</collectingCountry>
–
<emphasis id="8BFFEAF4B70B0E168A76E04AFB374EF2" bold="true" box="[382,1173,629,656]" pageId="10" pageNumber="11">
<location id="BC54603DB70B0E168A76E04AFE794EF2" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168A76E04AFE794EF2" box="[382,475,629,656]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Bolivar" pageId="10" pageNumber="11">Bolívar</location>
,
<location id="BC54603DB70B0E168AE5E049FD584EF2" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168AE5E049FD584EF2" box="[493,762,630,656]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Cartagena de Indias" pageId="10" pageNumber="11">Cartagena de Indias</location>
,
<location id="BC54603DB70B0E168803E049FC4A4EF2" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168803E049FC4A4EF2" box="[779,1000,630,656]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Islas del Rosario" pageId="10" pageNumber="11">Islas del Rosario</location>
,
<collectingMunicipality id="5950AC9CB70B0E1688F2E049FB374EF2" box="[1018,1173,630,656]" pageId="10" pageNumber="11">Isla Grande</collectingMunicipality>
</emphasis>
•
<specimenCount id="AF8DFD6FB70B0E168FBFE048FB724EF3" box="[1207,1232,631,657]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
;
<location id="BC54603DB70B0E168FEAE049FAD04EF2" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168FEAE049FAD04EF2" box="[1250,1394,630,656]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Playa Libre" pageId="10" pageNumber="11">Playa Libre</location>
;
<geoCoordinate id="DCBF5021B70B0E168BB5E0A7FEC94ED6" box="[189,363,664,692]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="52.3" value="10.181194">10°10′52.3″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B70B0E168A71E0A7FD8E4ED6" box="[377,556,664,692]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="54.8" value="-75.73189">75°43′54.8″ W</geoCoordinate>
;
<date id="CD351026B70B0E168933E0A6FD6E4ED6" box="[571,716,665,692]" pageId="10" pageNumber="11" value="2017-10-05">
<collectingDate id="DD71E9CEB70B0E168933E0A6FD6E4ED6" box="[571,716,665,692]" pageId="10" pageNumber="11" value="2017-10-05">5 Oct. 2017</collectingDate>
</date>
;
<collectorName id="147E5330B70B0E1689D2E0A5FC6E4ED6" box="[730,972,666,692]" pageId="10" pageNumber="11">C.M. López-Orozco</collectorName>
,
<collectorName id="147E5330B70B0E1688D2E0A5FB3F4ED1" box="[986,1181,665,692]" pageId="10" pageNumber="11">R. Borja-Arrieta</collectorName>
and
<collectorName id="147E5330B70B0E168FD1E0A5FF564EB5" pageId="10" pageNumber="11">Y.M. Carpio- Díaz</collectorName>
leg.;
<specimenCode id="E92D9E9DB70B0E168A3EE082FDA64EB5" box="[310,516,701,727]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 127</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B93436E6B70B0E168BE2E0C1FA954818" blockId="10.[189,1399,766,1146]" pageId="10" pageNumber="11">
<materialsCitation id="09E33CBBB70B0E168BE2E0C1FB2C4F3D" ID-GBIF-Occurrence="3464759302" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; Borja-Arrieta &amp; Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Cartagena de Indias" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 128" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar" typeStatus="paratype">
<emphasis id="8BFFEAF4B70B0E168BE2E0C1FECB4F7A" bold="true" box="[234,361,766,792]" pageId="10" pageNumber="11">
<typeStatus id="66308844B70B0E168BE2E0C1FECB4F7A" box="[234,361,766,792]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="C19C7676B70B0E168BB5E11DFEFA4F5E" box="[189,344,802,828]" name="Colombia" pageId="10" pageNumber="11">COLOMBIA</collectingCountry>
–
<emphasis id="8BFFEAF4B70B0E168A80E11EFB614F59" bold="true" box="[392,1219,801,828]" pageId="10" pageNumber="11">
<collectingRegion id="7B4FF804B70B0E168A80E11EFE474F5E" box="[392,485,801,828]" pageId="10" pageNumber="11">Bolívar</collectingRegion>
,
<location id="BC54603DB70B0E168AF3E11EFCB24F59" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168AF3E11EFCB24F59" box="[507,784,801,828]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Cartagena de Indias" pageId="10" pageNumber="11" stateProvince="Bolivar">Cartagena de Indias</location>
,
<location id="BC54603DB70B0E16882FE11DFBAF4F59" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E16882FE11DFBAF4F59" box="[807,1037,801,828]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Islas del Rosario" pageId="10" pageNumber="11" stateProvince="Bolivar">Islas del Rosario</location>
,
<location id="BC54603DB70B0E168F2BE11DFB614F59" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168F2BE11DFB614F59" box="[1059,1219,801,828]" country="Colombia" county="Playa Libre" latitude="10.181194" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" name="Isla Grande" pageId="10" pageNumber="11" stateProvince="Bolivar">Isla Grande</location>
</emphasis>
•
<specimenCount id="AF8DFD6FB70B0E168FE6E11DFA814F5E" box="[1262,1315,802,828]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
(parts in micropreparations); same collection data as for holotype;
<specimenCode id="E92D9E9DB70B0E1688B4E17AFB2C4F3D" box="[956,1166,837,863]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 128</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168FBEE17AFB914FE0" ID-GBIF-Occurrence="3464759350" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; Borja-Arrieta &amp; Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Bolivar" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 129" specimenCount="2" specimenCount-female="2" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
<specimenCount id="AF8DFD6FB70B0E168FBEE17AFB5D4F3D" box="[1206,1279,837,863]" pageId="10" pageNumber="11" type="female">2 ♀♀</specimenCount>
(parts in micropreparations); same collection data as for holotype;
<specimenCode id="E92D9E9DB70B0E168860E157FB914FE0" box="[872,1075,872,898]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 129</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168F46E156FCB14FC7" ID-GBIF-Occurrence="3464759395" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; Borja-Arrieta &amp; Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Bolivar" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 130" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
many
<specimenCount id="AF8DFD6FB70B0E168F9FE156FB614FE1" box="[1175,1219,873,899]" pageId="10" pageNumber="11" type="male">♂♂</specimenCount>
and
<specimenCount id="AF8DFD6FB70B0E168FF6E156FA894FE1" box="[1278,1323,873,899]" pageId="10" pageNumber="11" type="female">♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="E92D9E9DB70B0E16894CE1B4FCB14FC7" box="[580,787,907,933]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 130</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E16883CE1B3FAB64FAB" ID-GBIF-Occurrence="3464759323" collectingDate="2017-10-06" collectionCode="R" collectorName="El Silencio" country="Colombia" county="Playa Libre" latitude="10.180362" location="El Silencio" longLatPrecision="1" longitude="-75.73797" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 131" specimenCount="20" specimenCount-female="13" specimenCount-male="7" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
<specimenCount id="AF8DFD6FB70B0E16883CE1B3FCDE4FC4" box="[820,892,908,934]" pageId="10" pageNumber="11" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="AF8DFD6FB70B0E168885E1B4FC414FC4" box="[909,995,907,934]" pageId="10" pageNumber="11" type="female">13 ♀♀</specimenCount>
;
<location id="BC54603DB70B0E1688FDE1B4FBDC4FC7" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E1688FDE1B4FBDC4FC7" box="[1013,1150,907,933]" country="Colombia" latitude="10.180362" longLatPrecision="1" longitude="-75.73797" name="El Silencio" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">El Silencio</location>
,
<location id="BC54603DB70B0E168F85E1B4FAD04FC7" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168F85E1B4FAD04FC7" box="[1165,1394,907,934]" country="Colombia" latitude="10.180362" longLatPrecision="1" longitude="-75.73797" name="Playa La Cocotera" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Playa La Cocotera</location>
;
<geoCoordinate id="DCBF5021B70B0E168BB5E192FECB4FAB" box="[189,361,941,969]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="49.3" value="10.180362">10°10′49.3″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B70B0E168A7DE192FD844FAB" box="[373,550,941,969]" degrees="75" direction="west" minutes="44" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="16.7" value="-75.73797">75°44′16.7″ W</geoCoordinate>
;
<date id="CD351026B70B0E16893AE191FD1D4FAB" box="[562,703,942,969]" pageId="10" pageNumber="11" value="2017-10-06">
<collectingDate id="DD71E9CEB70B0E16893AE191FD1D4FAB" box="[562,703,942,969]" pageId="10" pageNumber="11" value="2017-10-06">6 Oct. 2017</collectingDate>
</date>
; same collectors as for holotype;
<specimenCode id="E92D9E9DB70B0E168F41E190FAB64FAB" box="[1097,1300,943,969]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 131</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168E25E190FE2A486D" ID-GBIF-Occurrence="3464759368" collectingDate="2017-10-05" collectionCode="R" collectorName="La Punta &amp; El Terminal" country="Colombia" county="Playa Libre" latitude="10.182153" location="La Punta" longLatPrecision="1" longitude="-75.72682" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 132" specimenCount="18" specimenCount-female="10" specimenCount-male="8" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
<specimenCount id="AF8DFD6FB70B0E168E25E190FAD34FAB" box="[1325,1393,943,969]" pageId="10" pageNumber="11" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="AF8DFD6FB70B0E168BB5E1EDFEB14F8E" box="[189,275,978,1004]" pageId="10" pageNumber="11" type="female">10 ♀♀</specimenCount>
;
<location id="BC54603DB70B0E168A2EE1EDFE354F8E" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168A2EE1EDFE354F8E" box="[294,407,978,1004]" country="Colombia" latitude="10.182153" longLatPrecision="1" longitude="-75.72682" name="La Punta" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">La Punta</location>
,
<location id="BC54603DB70B0E168AAFE1EDFD984F8E" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168AAFE1EDFD984F8E" box="[423,570,978,1004]" country="Colombia" latitude="10.182153" longLatPrecision="1" longitude="-75.72682" name="El Terminal" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">El Terminal</location>
;
<geoCoordinate id="DCBF5021B70B0E168942E1EFFCAB4F8E" box="[586,777,976,1004]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="55.75" value="10.182153">10°10′55.75″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B70B0E168812E1EFFC7C4F8E" box="[794,990,976,1004]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="36.57" value="-75.72682">75°43′36.57″ W</geoCoordinate>
; same collectors as for holotype;
<specimenCode id="E92D9E9DB70B0E168BB5E1CAFE2A486D" box="[189,392,1013,1039]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 132</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168AAAE1CAFD8E4850" ID-GBIF-Occurrence="3464759426" collectingDate="2017-09-07" collectionCode="R" collectorName="Playa Libre" country="Colombia" county="Playa Libre" latitude="10.181194" location="Playa Libre" longLatPrecision="1" longitude="-75.73214" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 133" specimenCount="5" specimenCount-female="5" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
<specimenCount id="AF8DFD6FB70B0E168AAAE1CAFE444872" box="[418,486,1013,1040]" pageId="10" pageNumber="11" type="female">5 ♀♀</specimenCount>
;
<location id="BC54603DB70B0E168AFDE1CAFD23486D" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168AFDE1CAFD23486D" box="[501,641,1013,1039]" country="Colombia" latitude="10.181194" longLatPrecision="1" longitude="-75.73214" name="Playa Libre" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Playa Libre</location>
;
<geoCoordinate id="DCBF5021B70B0E168985E1CCFC9B4872" box="[653,825,1011,1040]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="52.3" value="10.181194">10°10′52.3″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B70B0E16884EE1CCFC554872" box="[838,1015,1011,1040]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="55.7" value="-75.73214">75°43′55.7″ W</geoCoordinate>
;
<date id="CD351026B70B0E168F0CE1C9FB31486D" box="[1028,1171,1013,1040]" pageId="10" pageNumber="11" value="2017-09-07">
<collectingDate id="DD71E9CEB70B0E168F0CE1C9FB31486D" box="[1028,1171,1013,1040]" pageId="10" pageNumber="11" value="2017-09-07">7 Sep. 2017</collectingDate>
</date>
; same collectors as for holotype;
<specimenCode id="E92D9E9DB70B0E168A57E627FD8E4850" box="[351,556,1048,1074]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 133</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E16894FE627FAB24850" ID-GBIF-Occurrence="3464759333" box="[583,1296,1048,1075]" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; Borja-Arrieta &amp; Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Bolivar" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 134" specimenCount="1" specimenCount-female="1" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="holotype">
<specimenCount id="AF8DFD6FB70B0E16894FE627FDD44851" box="[583,630,1048,1075]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
; same collection data as for
<typeStatus id="66308844B70B0E1688C4E627FB944850" box="[972,1078,1048,1074]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
;
<specimenCode id="E92D9E9DB70B0E168F4CE627FAB24850" box="[1092,1296,1048,1074]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 134</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168E24E627FC5F4837" ID-GBIF-Occurrence="3464759324" collectingDate="2017-10-05" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; Borja-Arrieta &amp; Diaz" country="Colombia" county="Playa Libre" latitude="10.181194" location="Bolivar" longLatPrecision="1" longitude="-75.73189" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="holotype">
<specimenCount id="AF8DFD6FB70B0E168E24E627FAD34851" box="[1324,1393,1048,1075]" pageId="10" pageNumber="11" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="AF8DFD6FB70B0E168BB5E603FF5C4834" box="[189,254,1084,1110]" pageId="10" pageNumber="11" type="female">3 ♀♀</specimenCount>
; same collection data as for
<typeStatus id="66308844B70B0E16894BE603FD0F4834" box="[579,685,1084,1110]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
;
<date id="CD351026B70B0E1689BFE604FCE34834" box="[695,833,1083,1110]" pageId="10" pageNumber="11" value="2017-10-05">
<collectingDate id="DD71E9CEB70B0E1689BFE604FCE34834" box="[695,833,1083,1110]" pageId="10" pageNumber="11" value="2017-10-05">5 Oct. 2017</collectingDate>
</date>
; ICN-CR-is 262
</materialsCitation>
•
<materialsCitation id="09E33CBBB70B0E168F1BE603FA914818" ID-GBIF-Occurrence="3464759434" collectingDate="2017-10-06" collectionCode="R" collectorName="La Punta &amp; Hotel Cocoliso" country="Colombia" county="Playa Libre" latitude="10.178387" location="La Punta" longLatPrecision="1" longitude="-75.7258" municipality="Isla Grande" pageId="10" pageNumber="11" specimenCode="CUDC-CRU 135" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande" typeStatus="paratype">
<specimenCount id="AF8DFD6FB70B0E168F1BE603FBE24834" box="[1043,1088,1084,1110]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
;
<location id="BC54603DB70B0E168F43E603FB174834" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168F43E603FB174834" box="[1099,1205,1084,1110]" country="Colombia" latitude="10.178387" longLatPrecision="1" longitude="-75.7258" name="La Punta" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">La Punta</location>
,
<location id="BC54603DB70B0E168FB7E603FAD04834" LSID="urn:lsid:plazi:treatment:312287F0B70B0E108A86E337FBAC4F58:BC54603DB70B0E168FB7E603FAD04834" box="[1215,1394,1084,1110]" country="Colombia" latitude="10.178387" longLatPrecision="1" longitude="-75.7258" name="Hotel Cocoliso" pageId="10" pageNumber="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Hotel Cocoliso</location>
;
<geoCoordinate id="DCBF5021B70B0E168BB5E662FED5481B" box="[189,375,1117,1145]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="42.19" value="10.178387">10°10′42.19″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B70B0E168A8BE662FDE1481B" box="[387,579,1117,1145]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="32.88" value="-75.7258">75°43′32.88″ W</geoCoordinate>
;
<date id="CD351026B70B0E168947E660FD7E481B" box="[591,732,1119,1145]" pageId="10" pageNumber="11" value="2017-10-06">
<collectingDate id="DD71E9CEB70B0E168947E660FD7E481B" box="[591,732,1119,1145]" pageId="10" pageNumber="11" value="2017-10-06">6 Oct. 2017</collectingDate>
</date>
; same collectors as for holotype;
<specimenCode id="E92D9E9DB70B0E168F6DE660FA914818" box="[1125,1331,1119,1146]" collectionCode="CUDC-CRU" pageId="10" pageNumber="11">CUDC-CRU 135</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F191656DB70B0E108BB5E69FFB2E4DC9" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="B93436E6B70B0E168BB5E69FFEED48D8" blockId="10.[189,335,1184,1210]" box="[189,335,1184,1210]" pageId="10" pageNumber="11">
<heading id="E27C818AB70B0E168BB5E69FFEED48D8" bold="true" box="[189,335,1184,1210]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="8BFFEAF4B70B0E168BB5E69FFEED48D8" bold="true" box="[189,335,1184,1210]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E6EFFC564888" blockId="10.[189,1012,1231,1258]" box="[189,1012,1231,1258]" pageId="10" pageNumber="11">
MEASUREMENTS. Maximum body length: male
<quantity id="7E739B03B70B0E1689D2E6F0FC8D4888" box="[730,815,1231,1258]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="10.0">10 mm</quantity>
; female
<quantity id="7E739B03B70B0E16889BE6F0FC524888" box="[915,1008,1231,1258]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="11" unit="mm" value="8.1">8.1 mm</quantity>
.
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E72FFB28492F" blockId="10.[189,1398,1296,1357]" pageId="10" pageNumber="11">
BODY. Colour yellowish with many irregular dark spots (
<figureCitation id="21B02A63B70B0E16889AE72FFC544948" box="[914,1014,1296,1322]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[296,1291,264,1755]" captionTargetId="figure-113@9.[296,1291,264,1755]" captionTargetPageId="9" captionText="Fig. 2. A. Tylos marcuzzii Giordani Soika, 1954.B. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos- Filho sp. nov. C. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. D. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989.E. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. F. Armadilloniscus ninae Schultz, 1984. G. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. H. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. I. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). J. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986. K. Agnara madagascariensis (Budde-Lund, 1885). L. Ctenorillo tuberosus (Budde-Lund, 1904). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6037898" httpUri="https://zenodo.org/record/6037898/files/figure.png" pageId="10" pageNumber="11">Fig. 2B</figureCitation>
). Body outline as in
<figureCitation id="21B02A63B70B0E168E06E72FFAD04948" box="[1294,1394,1296,1322]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
. Endoantennal conglobation. Dorsum covered with lanceolate scale-setae (
<figureCitation id="21B02A63B70B0E168F28E70CFBDE492F" box="[1056,1148,1331,1357]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E74BFB1E49EC" blockId="10.[189,1212,1396,1422]" box="[189,1212,1396,1422]" pageId="10" pageNumber="11">
CEPHALON. Frontal lamina triangular (
<figureCitation id="21B02A63B70B0E16897EE74BFD5549EC" box="[630,759,1396,1422]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3C–D</figureCitation>
); eyes consisting of 26–27 ommatidia.
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E78AFBAD4A76" blockId="10.[187,1398,1461,1557]" pageId="10" pageNumber="11">
PEREON. Pereonite 1 epimera grooved on lateral margins, schisma with inner lobe triangular, slightly surpassing outer lobe (
<figureCitation id="21B02A63B70B0E168AC9E7E8FDB94993" box="[449,539,1495,1521]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3E</figureCitation>
); pereonite 2 with ventral lobe triangular, shorter than outer lobe (
<figureCitation id="21B02A63B70B0E168E07E7E8FACA4993" box="[1295,1384,1495,1521]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3E</figureCitation>
); epimera of pereonites 2–4 subtriangular, 5–7 subquadrangular (
<figureCitation id="21B02A63B70B0E1688ABE7C5FBA34A76" box="[931,1025,1530,1556]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E403FB744AF9" blockId="10.[189,1399,1595,1691]" pageId="10" pageNumber="11">
PLEON. Epimera of pleonites 3–5 with rounded distal margin (
<figureCitation id="21B02A63B70B0E1688A4E404FBAB4A37" box="[940,1033,1595,1621]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3F</figureCitation>
). Ventral phylacomera present on pleonites 3–5; phylacomera 5 oar-shaped, with wide distal part, medial margin oblique and slightly sinuous (
<figureCitation id="21B02A63B70B0E168A2FE4BFFE244AF9" box="[295,390,1664,1691]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3G</figureCitation>
). Telson rectangular, posterior margin slightly folded upwards (
<figureCitation id="21B02A63B70B0E168F67E4BFFB6A4AF9" box="[1135,1224,1664,1691]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E4FDFE714A9C" blockId="10.[189,1398,1729,1790]" pageId="10" pageNumber="11">
ANTENNULA. Composed of one article, flattened and triangular, several aesthetascs inserted on slightly rounded lobe (
<figureCitation id="21B02A63B70B0E168A6FE4DBFE674A9C" box="[359,453,1764,1790]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="10" pageNumber="11">Fig. 3H</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70B0E168BB5E51AFE904B00" blockId="10.[189,1399,1829,1890]" pageId="10" pageNumber="11">
ANTENNA. Short and thickset, flagellum of three articles, apical article as long as first and second articles (
<figureCitation id="21B02A63B70B0E168BCEE577FE874B00" box="[198,293,1864,1890]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="10" pageNumber="11">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70B0E178BB5E5B6FC1C4DD2" blockId="10.[187,1399,1929,2025]" lastBlockId="11.[187,1399,265,432]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
MOUTH. Mandibles with strong molar process, left mandible (
<figureCitation id="21B02A63B70B0E168876E5B6FC724BC1" box="[894,976,1929,1955]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="10" pageNumber="11">Fig 4B</figureCitation>
) with two penicils on molar process, two on hairy lobe and five free penicils; right mandible (
<figureCitation id="21B02A63B70B0E168857E594FC1F4BA4" box="[863,957,1963,1990]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="10" pageNumber="11">Fig. 4C</figureCitation>
) with two penicils on molar process, one on hairy lobe and five free penicils. Maxillula (
<figureCitation id="21B02A63B70B0E168804E5F1FCC54B8A" box="[780,871,1998,2024]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="10" pageNumber="11">Fig. 4D</figureCitation>
) inner endite with three long penicils densely covered with thin setae, apical penicil longest; outer endite composed of 5+5 simple teeth plus one sensory seta, inner set bearing two teeth slightly serrate, median portion bearing three short setae. Maxilla (
<figureCitation id="21B02A63B70A0E178BCCE370FEBE4D0B" box="[196,284,335,361]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="11" pageNumber="12">Fig. 4E</figureCitation>
) of two lobes, inner lobe rounded, covered with thick setae, outer lobe covered with thin setae. Maxilliped (
<figureCitation id="21B02A63B70A0E178A58E34DFE0B4DEF" box="[336,425,370,397]" captionStart="Fig" captionStartId="12.[189,232,1961,1987]" captionTargetBox="[202,1389,904,1903]" captionTargetId="figure-203@12.[189,1399,888,1914]" captionTargetPageId="12" captionText="Fig. 4. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♀ (CDUC- CRU 129). A. Antenna. B. Left mandible. C. Right mandible. D. Maxillula. E. Maxilla. F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.6037904" httpUri="https://zenodo.org/record/6037904/files/figure.png" pageId="11" pageNumber="12">Fig. 4F</figureCitation>
) basis with rounded outer margin, palp proximal article triangular without setae; endite rectangular, bearing five apical and two subapical penicils.
</paragraph>
<paragraph id="B93436E6B70A0E178BB5E3E7FE674D93" blockId="11.[189,453,471,498]" box="[189,453,471,498]" pageId="11" pageNumber="12">
UROPOD. As in
<figureCitation id="21B02A63B70A0E178A64E3E8FE674D93" box="[364,453,471,497]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[194,1373,756,1837]" captionTargetId="figure-192@11.[189,1399,742,1850]" captionTargetPageId="11" captionText="Fig. 3. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Habitus, lateral view. B. Dorsal scale-seta. C. Cephalon, frontal view. D. Cephalon, dorsal view. E. Epimera of pereonites 1–3, ventral view. F. Pleonites 4, 5 and telson, dorsal view. G. Pleon, ventral view. H. Antennula." figureDoi="http://doi.org/10.5281/zenodo.6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="11" pageNumber="12">Fig 3G.</figureCitation>
</paragraph>
<paragraph id="B93436E6B70A0E178BB5E026FCEE4E1B" blockId="11.[189,1398,536,634]" pageId="11" pageNumber="12">
PEREOPODS. Pereopod 1 (
<figureCitation id="21B02A63B70A0E178AE3E026FDE94E51" box="[491,587,536,563]" captionStart="Fig" captionStartId="13.[189,232,1227,1253]" captionTargetBox="[201,1386,280,1164]" captionTargetId="figure-122@13.[189,1399,265,1186]" captionTargetPageId="13" captionText="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.6037906" httpUri="https://zenodo.org/record/6037906/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
) basis with triangular latero-apical process. Pereopod 7 (
<figureCitation id="21B02A63B70A0E178E07E026FACF4E51" box="[1295,1389,536,563]" captionStart="Fig" captionStartId="13.[189,232,1227,1253]" captionTargetBox="[201,1386,280,1164]" captionTargetId="figure-122@13.[189,1399,265,1186]" captionTargetPageId="13" captionText="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.6037906" httpUri="https://zenodo.org/record/6037906/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
) propodus slightly inflated; dactylus of one claw, dactylar seta stout, not surpassing claw and bearing several small sensory setae on median to apical portion.
</paragraph>
<caption id="EDF4666EB70A0E178BB5E55CFDA04B84" ID-DOI="http://doi.org/10.5281/zenodo.6037900" ID-Zenodo-Dep="6037900" httpUri="https://zenodo.org/record/6037900/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1891,1917]" targetBox="[194,1373,756,1837]" targetPageId="11">
<paragraph id="B93436E6B70A0E178BB5E55CFDA04B84" blockId="11.[188,1399,1891,2023]" pageId="11" pageNumber="12">
<emphasis id="8BFFEAF4B70A0E178BB5E55CFEB04B1F" bold="true" box="[189,274,1891,1917]" pageId="11" pageNumber="12">Fig. 3.</emphasis>
<taxonomicName id="7E8B4D65B70A0E178A28E55CFC524B1F" authority="Lopez-Orozco, Carpio-Diaz &amp; Campos-Filho" authorityName="Lopez-Orozco, Carpio-Diaz &amp; Campos-Filho" authorityYear="2022" box="[288,1008,1891,1917]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negroi" status="sp. nov.">
<emphasis id="8BFFEAF4B70A0E178A28E55CFE1A4B1F" box="[288,440,1891,1917]" italics="true" pageId="11" pageNumber="12">Tylos negroi</emphasis>
López-Orozco, Carpio-Díaz &amp; Campos-Filho
</taxonomicName>
<taxonomicNameLabel id="90CC578FB70A0E1788F6E55CFBC34B1F" box="[1022,1121,1891,1917]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CDUC- CRU 128).
<emphasis id="8BFFEAF4B70A0E178A49E5B9FEF44BC2" bold="true" box="[321,342,1926,1952]" pageId="11" pageNumber="12">A</emphasis>
. Habitus, lateral view.
<emphasis id="8BFFEAF4B70A0E178968E5B9FDD64BC2" bold="true" box="[608,628,1926,1952]" pageId="11" pageNumber="12">B</emphasis>
. Dorsal scale-seta.
<emphasis id="8BFFEAF4B70A0E17885CE5B9FCCB4BC2" bold="true" box="[852,873,1926,1952]" pageId="11" pageNumber="12">C</emphasis>
. Cephalon, frontal view.
<emphasis id="8BFFEAF4B70A0E178F85E5B9FB004BC2" bold="true" box="[1165,1186,1926,1952]" pageId="11" pageNumber="12">D</emphasis>
. Cephalon, dorsal view.
<emphasis id="8BFFEAF4B70A0E178A0CE596FEBA4BA1" bold="true" box="[260,280,1961,1987]" pageId="11" pageNumber="12">E</emphasis>
. Epimera of pereonites 1–3, ventral view.
<emphasis id="8BFFEAF4B70A0E17881FE596FC854BA1" bold="true" box="[791,807,1961,1987]" pageId="11" pageNumber="12">F</emphasis>
. Pleonites 4, 5 and telson, dorsal view.
<emphasis id="8BFFEAF4B70A0E178E0FE596FABE4BA1" bold="true" box="[1287,1308,1961,1987]" pageId="11" pageNumber="12">G</emphasis>
. Pleon, ventral view.
<emphasis id="8BFFEAF4B70A0E178A54E5F2FED14B85" bold="true" box="[348,371,1997,2023]" pageId="11" pageNumber="12">H</emphasis>
. Antennula.
</paragraph>
</caption>
<paragraph id="B93436E6B70D0E108BE2E336FE884D41" blockId="12.[189,958,265,326]" box="[234,298,265,291]" pageId="12" pageNumber="13">
<emphasis id="8BFFEAF4B70D0E108BE2E336FE884D41" bold="true" box="[234,298,265,291]" pageId="12" pageNumber="13">
<collectingRegion id="7B4FF804B70D0E108BE2E336FE884D41" box="[234,298,265,291]" country="Maldives" name="Maale" pageId="12" pageNumber="13">Male</collectingRegion>
</emphasis>
</paragraph>
<paragraph id="B93436E6B70D0E108BB5E313FC1C4D24" blockId="12.[189,958,265,326]" box="[189,958,299,326]" pageId="12" pageNumber="13">
PEREOPODS 1 AND 7. Without any sexual dimorphism (
<figureCitation id="21B02A63B70D0E108838E313FC104D24" box="[816,946,299,326]" captionStart="Fig" captionStartId="13.[189,232,1227,1253]" captionTargetBox="[201,1386,280,1164]" captionTargetId="figure-122@13.[189,1399,265,1186]" captionTargetPageId="13" captionText="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.6037906" httpUri="https://zenodo.org/record/6037906/files/figure.png" pageId="12" pageNumber="13">Fig. 5A–B</figureCitation>
).
</paragraph>
<paragraph id="B93436E6B70D0E108BB5E351FB2E4DC9" blockId="12.[189,1399,365,427]" pageId="12" pageNumber="13">
PLEOPODS. Pleopod 2 exopod rectangular (
<figureCitation id="21B02A63B70D0E1089B9E352FCAE4DEA" box="[689,780,365,392]" captionStart="Fig" captionStartId="13.[189,232,1227,1253]" captionTargetBox="[201,1386,280,1164]" captionTargetId="figure-122@13.[189,1399,265,1186]" captionTargetPageId="13" captionText="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.6037906" httpUri="https://zenodo.org/record/6037906/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
) with respiratory folds; endopod with distal portion slightly directed inwards, apex covered with cleft scales of 2–4 branches (
<figureCitation id="21B02A63B70D0E108F2AE3AFFBDD4DC9" box="[1058,1151,400,427]" captionStart="Fig" captionStartId="13.[189,232,1227,1253]" captionTargetBox="[201,1386,280,1164]" captionTargetId="figure-122@13.[189,1399,265,1186]" captionTargetPageId="13" captionText="Fig. 5. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov., paratype, ♂ (CDUC- CRU 128). A. Pereopod 1. B. Pereopod 7. C. Pleopod 2." figureDoi="http://doi.org/10.5281/zenodo.6037906" httpUri="https://zenodo.org/record/6037906/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F191656DB70D0E108BB5E3EDFB2C4EA9" pageId="12" pageNumber="13" type="discussion">
<paragraph id="B93436E6B70D0E108BB5E3EDFE924D8E" blockId="12.[189,304,466,492]" box="[189,304,466,492]" pageId="12" pageNumber="13">
<heading id="E27C818AB70D0E108BB5E3EDFE924D8E" bold="true" box="[189,304,466,492]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="8BFFEAF4B70D0E108BB5E3EDFE924D8E" bold="true" box="[189,304,466,492]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B70D0E108BB5E03FFB2C4EA9" blockId="12.[188,1399,512,715]" pageId="12" pageNumber="13">
Presently, the genus
<taxonomicName id="7E8B4D65B70D0E108AA3E03FFE4B4E78" authorityName="Audouin" authorityYear="1826" box="[427,489,512,538]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="8BFFEAF4B70D0E108AA3E03FFE4B4E78" box="[427,489,512,538]" italics="true" pageId="12" pageNumber="13">Tylos</emphasis>
</taxonomicName>
has a worldwide distribution and comprises 21 coastal species (
<bibRefCitation id="DD1A4B17B70D0E108FD8E03FFEFC4E5C" author="Schmalfuss H. &amp; Vergara K." pageId="12" pageNumber="13" pagination="1 - 42" refId="ref25254" refString="Schmalfuss H. &amp; Vergara K. 2000. The isopod genus Tylos (Oniscidea: Tylidae) in Chile, with bibliographies of all described species of the genus. Stuttgarter Beitrage zur Naturkunde A 612: 1 - 42." type="journal article" year="2000">Schmalfuss &amp; Vergara 2000</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B70D0E108A64E01CFD9F4E5C" author="Schmalfuss H." box="[364,573,547,574]" pageId="12" pageNumber="13" pagination="1 - 341" refId="ref25160" refString="Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341." type="journal article" year="2003">Schmalfuss 2003</bibRefCitation>
). In
<collectingCountry id="C19C7676B70D0E10897CE01CFD494E5F" box="[628,747,547,573]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
, only
<taxonomicName id="7E8B4D65B70D0E108831E01BFB2F4E5C" authority="Budde-Lund, 1885" authorityName="Budde-Lund" authorityYear="1885" box="[825,1165,547,574]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="niveus">
<emphasis id="8BFFEAF4B70D0E108831E01BFCE54E5F" box="[825,839,548,573]" italics="true" pageId="12" pageNumber="13">T</emphasis>
.
<emphasis id="8BFFEAF4B70D0E108850E01BFC064E5F" box="[856,932,548,573]" italics="true" pageId="12" pageNumber="13">niveus</emphasis>
<bibRefCitation id="DD1A4B17B70D0E1088A5E01CFB2F4E5C" author="Budde-Lund G." box="[941,1165,547,574]" pageId="12" pageNumber="13" refId="ref21297" refString="Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen &amp; Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885</bibRefCitation>
</taxonomicName>
was recorded from the Caribbean region (
<bibRefCitation id="DD1A4B17B70D0E108AC7E078FD564E00" author="Carpio-Diaz Y. M. &amp; Lopez-Orozco C. M. &amp; Herrera-Medina Y. &amp; Navas G. R. &amp; Bermudez A." box="[463,756,583,610]" pageId="12" pageNumber="13" pagination="433 - 437" refId="ref21642" refString="Carpio-Diaz Y. M., Lopez-Orozco C. M., Herrera-Medina Y., Navas G. R. &amp; Bermudez A. 2016. Primer registro de Tylos niveus y nuevo reporte de Porcellionides pruinosus (Oniscidea: Tylidae y Porcellionidae) para Colombia. Revista de la Academia colombiana de Ciencias exactas fisicas y naturales 40 (156): 433 - 437. https: // doi. org / 10.18257 / raccefyn. 343" type="journal article" year="2016">
Carpio-Díaz
<emphasis id="8BFFEAF4B70D0E108962E078FD0B4E03" box="[618,681,583,609]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2016
</bibRefCitation>
).
<taxonomicName id="7E8B4D65B70D0E108803E078FC024E02" authority="López-Orozco &amp; Carpio-Díaz &amp; Borja-Arrieta &amp; Navas-S &amp; Campos-Filho &amp; Taiti &amp; Mateos &amp; Olazaran &amp; Caballero &amp; Jotty &amp; Gómez-Estrada &amp; Hurtado, 2022" authorityName="López-Orozco &amp; Carpio-Díaz &amp; Borja-Arrieta &amp; Navas-S &amp; Campos-Filho &amp; Taiti &amp; Mateos &amp; Olazaran &amp; Caballero &amp; Jotty &amp; Gómez-Estrada &amp; Hurtado" authorityYear="2022" box="[779,928,583,609]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="negroi" status="sp. nov.">
<emphasis id="8BFFEAF4B70D0E108803E078FC024E02" box="[779,928,583,609]" italics="true" pageId="12" pageNumber="13">Tylos negroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90CC578FB70D0E1088A3E078FBAB4E03" box="[939,1033,583,609]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all species of the genus by the shape of the ventral phylacomera 5. Moreover, it differs from
<taxonomicName id="7E8B4D65B70D0E108FCEE054FA8C4EE6" authorityName="Budde-Lund" authorityYear="1885" box="[1222,1326,619,644]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="niveus">
<emphasis id="8BFFEAF4B70D0E108FCEE054FA8C4EE6" box="[1222,1326,619,644]" italics="true" pageId="12" pageNumber="13">T. niveus</emphasis>
</taxonomicName>
in the pereonite 1 epimeron having the inner lobe of the schisma surpassing the outer lobe (vs inner lobe not surpassing outer lobe) and the pereopod 7 propodus being inflated (vs not inflated).
</paragraph>
</subSubSection>
<subSubSection id="F191656DB70D0E108BB5E0CEFBAC4F58" box="[189,1038,753,826]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="B93436E6B70D0E108BB5E0CEFEFA4F69" blockId="12.[189,344,753,779]" box="[189,344,753,779]" pageId="12" pageNumber="13">
<heading id="E27C818AB70D0E108BB5E0CEFEFA4F69" bold="true" box="[189,344,753,779]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="8BFFEAF4B70D0E108BB5E0CEFEFA4F69" bold="true" box="[189,344,753,779]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B70D0E108BB5E11FFBAC4F58" blockId="12.[189,1038,800,826]" box="[189,1038,800,826]" pageId="12" pageNumber="13">
Presently known only from
<collectingCountry id="C19C7676B70D0E10890DE11FFD8D4F58" box="[517,559,800,826]" name="Iceland" pageId="12" pageNumber="13">Isla</collectingCountry>
Grande, Cartagena de Indias,
<collectingCountry id="C19C7676B70D0E10889BE11FFBA94F58" box="[915,1035,800,826]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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