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<taxonomicName id="7E8B4D65B7090E148931E76DFC59490F" ID-CoL="7CZQY" authority="Giordani Soika, 1954" authorityName="Giordani Soika" authorityYear="1954" box="[569,1019,1362,1389]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marcuzzii">
<emphasis id="8BFFEAF4B7090E148931E76DFD5A490E" bold="true" box="[569,760,1362,1389]" italics="true" pageId="8" pageNumber="9">Tylos marcuzzii</emphasis>
<bibRefCitation id="DD1A4B17B7090E1489F7E76DFC59490F" author="Giordani Soika A." box="[767,1019,1362,1389]" pageId="8" pageNumber="9" pagination="63 - 83" refId="ref22264" refString="Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83." type="journal article" year="1954">Giordani Soika, 1954</bibRefCitation>
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<figureCitation id="21B02A63B7090E1489D5E748FC8449F0" box="[733,806,1399,1426]" captionStart="Fig" captionStartId="3.[189,232,1142,1168]" captionTargetBox="[198,1394,269,1093]" captionTargetId="figure-182@3.[189,1399,265,1102]" captionTargetPageId="3" captionText="Fig. 1. Map of the study areas and distribution of species of terrestrial isopods from Isla Grande. 1. Ligia baudiniana Milne Edwards, 1840. 2. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos-Filho sp. nov. 3. Tylos marcuzzii Giordani Soika, 1954. 4. Tylos niveus Budde-Lund, 1885. 5. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. 6. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989. 7. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. 8. Armadilloniscus ninae Schultz, 1984. 9. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. 10. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. 11. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). 12. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986.13. Trichorhina heterophthalma Lemos de Castro, 1964.14. Trichorhina bermudezae Carpio-Díaz, López-Orozco &amp; Campos-Filho, 2018. 15. Porcellionides pruinosus (Brandt, 1833). 16. Agnara madagascariensis (Budde-Lund, 1885). 17. Ctenorillo tuberosus (Budde-Lund, 1904)." figureDoi="http://doi.org/10.5281/zenodo.6037894" httpUri="https://zenodo.org/record/6037894/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
,
<figureCitation id="21B02A63B7090E14883CE747FCFA49F0" box="[820,856,1400,1426]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[296,1291,264,1755]" captionTargetId="figure-113@9.[296,1291,264,1755]" captionTargetPageId="9" captionText="Fig. 2. A. Tylos marcuzzii Giordani Soika, 1954.B. Tylos negroi López-Orozco, Carpio-Díaz &amp; Campos- Filho sp. nov. C. Stenoniscus nestori López-Orozco, Taiti &amp; Campos-Filho sp. nov. D. Armadilloniscus caraibicus Paoletti &amp; Stinner, 1989.E. Armadilloniscus luisi Carpio-Díaz, Taiti &amp; Campos-Filho sp. nov. F. Armadilloniscus ninae Schultz, 1984. G. Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. H. Littorophiloscia amphindica Taiti &amp; Ferrara, 1986. I. Littorophiloscia denticulata (Ferrara &amp; Taiti, 1982). J. Littorophiloscia tropicalis Taiti &amp; Ferrara, 1986. K. Agnara madagascariensis (Budde-Lund, 1885). L. Ctenorillo tuberosus (Budde-Lund, 1904). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6037898" httpUri="https://zenodo.org/record/6037898/files/figure.png" pageId="8" pageNumber="9">2A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F191656DB7090E148BB5E784FD174A2A" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="B93436E6B7090E148BB5E784FCBE49B7" blockId="8.[189,796,1467,1494]" box="[189,796,1467,1494]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="999B11C8B7090E148BB5E784FCBE49B7" box="[189,796,1467,1494]" pageId="8" pageNumber="9">
<treatmentCitation id="382A10F7B7090E148BB5E784FD0549B7" author="Giordani Soika A." box="[189,679,1467,1494]" page="79" pageId="8" pageNumber="9" year="1954">
<taxonomicName id="7E8B4D65B7090E148BB5E784FD0549B7" ID-CoL="7CZQY" authority="Giordani Soika, 1954: 79" authorityName="Giordani Soika" authorityPageNumber="79" authorityYear="1954" box="[189,679,1467,1494]" class="Malacostraca" family="Tylidae" genus="Tylos" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marcuzzii">
<emphasis id="8BFFEAF4B7090E148BB5E784FED749B7" box="[189,373,1467,1493]" italics="true" pageId="8" pageNumber="9">Tylos marcuzzii</emphasis>
<bibRefCitation id="DD1A4B17B7090E148A74E784FD0549B7" author="Giordani Soika A." box="[380,679,1467,1494]" pageId="8" pageNumber="9" pagination="63 - 83" refId="ref22264" refString="Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83." type="journal article" year="1954">Giordani Soika, 1954: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 10/1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B93436E6B7090E148BB5E7C1FE6F4A7A" blockId="8.[189,461,1534,1560]" box="[189,461,1534,1560]" pageId="8" pageNumber="9">
<heading id="E27C818AB7090E148BB5E7C1FE6F4A7A" bold="true" box="[189,461,1534,1560]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="8BFFEAF4B7090E148BB5E7C1FE6F4A7A" bold="true" box="[189,461,1534,1560]" pageId="8" pageNumber="9">Additional references</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7090E148BB5E411FD174A2A" blockId="8.[189,693,1582,1608]" box="[189,693,1582,1608]" pageId="8" pageNumber="9">
<bibRefCitation id="DD1A4B17B7090E148BB5E411FECC4A2A" author="Schultz G. A." box="[189,366,1582,1608]" pageId="8" pageNumber="9" pagination="162 - 173" refId="ref25643" refString="Schultz G. A. 1974. Terrestrial isopod crustaceans (Oniscoidea) mainly from the West Indies and adjacent regions: 1. Tylos and Ligia. Studies on the fauna of Cura ҫao and other Caribbean islands 149: 162 - 173. Available from http: // www. repository. naturalis. nl / document / 549820 [accessed 7 Jan. 2022]." type="journal article" year="1974">Schultz (1974)</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E148A72E411FD134A2A" author="Kensley B. &amp; Schotte M." box="[378,689,1582,1608]" pageId="8" pageNumber="9" refId="ref23014" refString="Kensley B. &amp; Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022]." type="url" year="1989">Kensley &amp; Schotte (1989)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F191656DB7090E148BB5E44EFBDC4B49" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="B93436E6B7090E148BB5E44EFE0F4AE9" blockId="8.[189,429,1649,1675]" box="[189,429,1649,1675]" pageId="8" pageNumber="9">
<heading id="E27C818AB7090E148BB5E44EFE0F4AE9" bold="true" box="[189,429,1649,1675]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="8BFFEAF4B7090E148BB5E44EFE0F4AE9" bold="true" box="[189,429,1649,1675]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7090E148BB5E49DFBDC4B49" blockId="8.[189,1399,1697,1835]" pageId="8" pageNumber="9">
<materialsCitation id="09E33CBBB7090E148BB5E49DFDB04B64" ID-GBIF-Occurrence="3464759399" collectingDate="2017-09-07" collectionCode="R" collectorName="C. M. Lopez-Orozco &amp; R. Borja-Arrieta &amp; Y. M. Carpio- Diaz" country="Colombia" latitude="10.181223" location="Cartagena de Indias" longLatPrecision="1" longitude="-75.732254" pageId="8" pageNumber="9" specimenCode="CUDC-CRU 136" specimenCount="1" specimenCount-female="1" stateProvince="Bolivar">
<collectingCountry id="C19C7676B7090E148BB5E49DFEFA4ADE" box="[189,344,1698,1724]" name="Colombia" pageId="8" pageNumber="9">COLOMBIA</collectingCountry>
–
<emphasis id="8BFFEAF4B7090E148A73E49EFB214ADE" bold="true" box="[379,1155,1697,1724]" pageId="8" pageNumber="9">
<collectingRegion id="7B4FF804B7090E148A73E49EFE7A4ADE" box="[379,472,1697,1724]" pageId="8" pageNumber="9">Bolívar</collectingRegion>
,
<location id="BC54603DB7090E148AE0E49DFD534ADE" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E148AE0E49DFD534ADE" box="[488,753,1698,1724]" country="Colombia" latitude="10.181223" longLatPrecision="1" longitude="-75.732254" name="Cartagena de Indias" pageId="8" pageNumber="9" stateProvince="Bolivar">Cartagena de Indias</location>
,
<location id="BC54603DB7090E148809E49DFC784ADE" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E148809E49DFC784ADE" box="[769,986,1698,1724]" country="Colombia" latitude="10.181223" longLatPrecision="1" longitude="-75.732254" name="Islas del Rosario" pageId="8" pageNumber="9" stateProvince="Bolivar">Islas del Rosario</location>
,
<location id="BC54603DB7090E1488E2E49DFB214ADE" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E1488E2E49DFB214ADE" box="[1002,1155,1698,1724]" country="Colombia" latitude="10.181223" longLatPrecision="1" longitude="-75.732254" name="Isla Grande" pageId="8" pageNumber="9" stateProvince="Bolivar">Isla Grande</location>
</emphasis>
•
<specimenCount id="AF8DFD6FB7090E148FAAE49DFB704ADE" box="[1186,1234,1698,1724]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
;
<location id="BC54603DB7090E148FEBE49DFAD04ADE" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E148FEBE49DFAD04ADE" box="[1251,1394,1698,1724]" country="Colombia" latitude="10.181223" longLatPrecision="1" longitude="-75.732254" name="Playa Libre" pageId="8" pageNumber="9" stateProvince="Bolivar">Playa Libre</location>
;
<geoCoordinate id="DCBF5021B7090E148BB5E4FAFEC94A83" box="[189,363,1733,1761]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="52.4" value="10.181223">10°10′52.4″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B7090E148A71E4FAFD8E4A83" box="[377,556,1733,1761]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="56.1" value="-75.732254">75°43′56.1″ W</geoCoordinate>
;
<date id="CD351026B7090E148932E4F8FD6F4A83" box="[570,717,1735,1761]" pageId="8" pageNumber="9" value="2017-09-07">
<collectingDate id="DD71E9CEB7090E148932E4F8FD6F4A83" box="[570,717,1735,1761]" pageId="8" pageNumber="9" value="2017-09-07">7 Sep. 2017</collectingDate>
</date>
;
<collectorName id="147E5330B7090E1489D3E4F8FC6F4A83" box="[731,973,1735,1761]" pageId="8" pageNumber="9">C.M. López-Orozco</collectorName>
,
<collectorName id="147E5330B7090E1488D3E4F8FB3F4A83" box="[987,1181,1735,1761]" pageId="8" pageNumber="9">R. Borja-Arrieta</collectorName>
and
<collectorName id="147E5330B7090E148FD1E4F8FF564B64" pageId="8" pageNumber="9">Y.M. Carpio- Díaz</collectorName>
leg.;
<specimenCode id="E92D9E9DB7090E148A49E4D3FDB04B64" box="[321,530,1772,1798]" collectionCode="CUDC-CRU" pageId="8" pageNumber="9">CUDC-CRU 136</specimenCode>
</materialsCitation>
•
<materialsCitation id="09E33CBBB7090E14893EE4D3FBDB4B49" ID-GBIF-Occurrence="3464759418" collectingDate="2017-10-06" collectionCode="R" collectorName="La Punta" country="Colombia" latitude="10.182337" location="La Punta" longLatPrecision="1" longitude="-75.725975" municipality="La Punta" pageId="8" pageNumber="9" specimenCode="CUDC-CRU 137" specimenCount="51" specimenCount-female="39" specimenCount-male="11" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">
<specimenCount id="AF8DFD6FB7090E14893EE4D3FD2F4B64" box="[566,653,1772,1798]" pageId="8" pageNumber="9" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="AF8DFD6FB7090E148997E4D3FD554B64" box="[671,759,1772,1798]" pageId="8" pageNumber="9" type="female">39 ♀♀</specimenCount>
,
<specimenCount id="AF8DFD6FB7090E148802E4D3FCF64B64" box="[778,852,1772,1798]" pageId="8" pageNumber="9" type="generic">1 juv.</specimenCount>
;
<location id="BC54603DB7090E14886DE4D3FC754B64" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E14886DE4D3FC754B64" box="[869,983,1772,1798]" country="Colombia" latitude="10.182337" longLatPrecision="1" longitude="-75.725975" municipality="La Punta" name="La Punta" pageId="8" pageNumber="9" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">La Punta</location>
,
<location id="BC54603DB7090E1488E0E4D3FB034B64" LSID="urn:lsid:plazi:treatment:312287F0B7090E148931E76DFEC14B8A:BC54603DB7090E1488E0E4D3FB034B64" box="[1000,1185,1772,1798]" country="Colombia" latitude="10.182337" longLatPrecision="1" longitude="-75.725975" municipality="La Punta" name="Hotel Majagua" pageId="8" pageNumber="9" stateProvince="Bolivar, Cartagena de Indias, Islas del Rosario, Isla Grande">Hotel Majagua</location>
;
<geoCoordinate id="DCBF5021B7090E148FBAE4D5FAD04B64" box="[1202,1394,1770,1798]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="56.41" value="10.182337">10°10′56.41″ N</geoCoordinate>
,
<geoCoordinate id="DCBF5021B7090E148BB5E530FEDF4B49" box="[189,381,1807,1835]" degrees="75" direction="west" minutes="43" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="33.51" value="-75.725975">75°43′33.51″ W</geoCoordinate>
;
<date id="CD351026B7090E148A81E52EFDB44B49" box="[393,534,1809,1835]" pageId="8" pageNumber="9" value="2017-10-06">
<collectingDate id="DD71E9CEB7090E148A81E52EFDB44B49" box="[393,534,1809,1835]" pageId="8" pageNumber="9" value="2017-10-06">6 Oct. 2017</collectingDate>
</date>
; same collectors as for preceding;
<specimenCode id="E92D9E9DB7090E1488A3E52EFBDB4B49" box="[939,1145,1809,1835]" collectionCode="CUDC-CRU" pageId="8" pageNumber="9">CUDC-CRU 137</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F191656DB7090E148BB5E56BFEC14B8A" pageId="8" pageNumber="9" type="distribution">
<paragraph id="B93436E6B7090E148BB5E56BFEFA4B0C" blockId="8.[189,344,1876,1902]" box="[189,344,1876,1902]" pageId="8" pageNumber="9">
<heading id="E27C818AB7090E148BB5E56BFEFA4B0C" bold="true" box="[189,344,1876,1902]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="8BFFEAF4B7090E148BB5E56BFEFA4B0C" bold="true" box="[189,344,1876,1902]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B93436E6B7090E148BB5E5BBFEC14B8A" blockId="8.[189,1398,1924,2024]" pageId="8" pageNumber="9">
Florida,
<collectingCountry id="C19C7676B7090E148A2CE5BBFE314BFC" box="[292,403,1924,1950]" name="Bahamas" pageId="8" pageNumber="9">Bahamas</collectingCountry>
, Leeward Islands,
<collectingCountry id="C19C7676B7090E148971E5BBFD654BFC" box="[633,711,1924,1950]" name="Belize" pageId="8" pageNumber="9">Belize</collectingCountry>
,
<collectingCountry id="C19C7676B7090E1489DDE5BBFCEC4BFC" box="[725,846,1924,1950]" name="Venezuela" pageId="8" pageNumber="9">Venezuela</collectingCountry>
and
<collectingCountry id="C19C7676B7090E148886E5BBFC6E4BFC" box="[910,972,1924,1950]" name="Cuba" pageId="8" pageNumber="9">Cuba</collectingCountry>
(
<bibRefCitation id="DD1A4B17B7090E1488D7E5BBFB204BFC" author="Schultz G. A." box="[991,1154,1924,1950]" pageId="8" pageNumber="9" pagination="162 - 173" refId="ref25643" refString="Schultz G. A. 1974. Terrestrial isopod crustaceans (Oniscoidea) mainly from the West Indies and adjacent regions: 1. Tylos and Ligia. Studies on the fauna of Cura ҫao and other Caribbean islands 149: 162 - 173. Available from http: // www. repository. naturalis. nl / document / 549820 [accessed 7 Jan. 2022]." type="journal article" year="1974">Schultz 1974</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E148F9AE5BBFF594BA1" author="Kensley B. &amp; Schotte M." pageId="8" pageNumber="9" refId="ref23014" refString="Kensley B. &amp; Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022]." type="url" year="1989">Kensley &amp; Schotte 1989</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E148A00E596FDF74BA1" author="Schmalfuss H. &amp; Vergara K." box="[264,597,1961,1988]" pageId="8" pageNumber="9" pagination="1 - 42" refId="ref25254" refString="Schmalfuss H. &amp; Vergara K. 2000. The isopod genus Tylos (Oniscidea: Tylidae) in Chile, with bibliographies of all described species of the genus. Stuttgarter Beitrage zur Naturkunde A 612: 1 - 42." type="journal article" year="2000">Schmalfuss &amp; Vergara 2000</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E148969E596FCAB4BA1" author="Schmidt C." box="[609,777,1961,1987]" pageId="8" pageNumber="9" pagination="1 - 12" refId="ref25296" refString="Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12." type="journal article" year="2001">Schmidt 2001</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E14881DE596FC404BA1" author="Schmalfuss H." box="[789,994,1961,1987]" pageId="8" pageNumber="9" pagination="1 - 341" refId="ref25160" refString="Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341." type="journal article" year="2003">Schmalfuss 2003</bibRefCitation>
;
<bibRefCitation id="DD1A4B17B7090E1488E7E596FB784BA1" author="Hurtado L. A. &amp; Lee E. J. &amp; Mateos M. &amp; Taiti S." box="[1007,1242,1961,1987]" pageId="8" pageNumber="9" pagination="94081" refId="ref22475" refString="Hurtado L. A., Lee E. J., Mateos M. &amp; Taiti S. 2014. Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PloS ONE 9 (4): e 94081. https: // doi. org / 10.1371 / journal. pone. 0094081" type="journal article" year="2014">
Hurtado
<emphasis id="8BFFEAF4B7090E148F5FE595FB314BA1" box="[1111,1171,1961,1987]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2014
</bibRefCitation>
). First record for
<collectingCountry id="C19C7676B7090E148BEFE5F1FEFD4B8A" box="[231,351,1998,2024]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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