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<mods:title id="9DBA3C80D127C62BCEC00F8661029B8D">Revision of New World Cosmorrhyncha Meyrick, 1913 (Lepidoptera: Tortricidae: Olethreutinae), with descriptions of five new species</mods:title>
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<mods:namePart id="38168057A1981A8F09360152EA8022FC">Brown, John W.</mods:namePart>
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<mods:namePart id="548BF191B4F72D3911CCF68B9FA67AA5">Razowski, Józef</mods:namePart>
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<mods:namePart id="D0785C26DA69E36EAF0D8F107BD8B6DF">Timm, Alicia E.</mods:namePart>
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<subSubSection id="ECDC65548E06AF544582FF23FD46FF45" box="[192,707,217,241]" pageId="11" pageNumber="10" type="nomenclature">
<paragraph id="A47936DF8E06AF544582FF23FD46FF45" blockId="11.[192,707,217,274]" box="[192,707,217,241]" pageId="11" pageNumber="10">
<heading id="FF3181B38E06AF544582FF23FD46FF45" bold="true" box="[192,707,217,241]" fontSize="10" level="2" pageId="11" pageNumber="10" reason="3">
<taxonomicName id="63C64D5C8E06AF544582FF23FD46FF45" ID-CoL="6B7N9" authority="(Walker)" baseAuthorityName="Walker" baseAuthorityYear="1863" box="[192,707,217,241]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E06AF544582FF23FD46FF45" bold="true" box="[192,707,217,241]" pageId="11" pageNumber="10">
<emphasis id="96B2EACD8E06AF544582FF23FDC3FF45" bold="true" box="[192,582,217,241]" italics="true" pageId="11" pageNumber="10">Cosmorrhyncha ocelliferana</emphasis>
(Walker)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECDC65548E06AF544582FF00FEDBFEA6" box="[192,350,250,274]" pageId="11" pageNumber="10" type="description">
<paragraph id="A47936DF8E06AF544582FF00FEDBFEA6" blockId="11.[192,707,217,274]" box="[192,350,250,274]" pageId="11" pageNumber="10">
<figureCitation id="3CFD2A5A8E06AF544582FF00FE8DFEA5" box="[192,264,250,274]" captionStart="Figures 613" captionStartId="19.[196,291,1694,1716]" captionTargetBox="[195,1444,291,1661]" captionTargetId="figure@19.[192,1444,291,1661]" captionTargetPageId="19" captionText="Figures 613. Adults of Cosmorrhyncha. 6) C. tonsana from Costa Rica. 7) C. ocelliferana from Brazil (image of lectotype from NHMUK). 8) C. landryi from French Guiana. 9) C. parintina from Brazil. 10) C. macrospina female from Brazil. 11) C. macrospina male from Brazil (apex of right forewing damaged). 12) C. albistrigulana from Brazil. 13) C. osana from Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3702991" httpUri="https://zenodo.org/record/3702991/files/figure.png" pageId="11" pageNumber="10">Fig. 7</figureCitation>
,
<figureCitation id="3CFD2A5A8E06AF544456FF00FEB1FEA6" box="[276,308,250,274]" captionStart="Figures 1421" captionStartId="20.[192,285,1792,1814]" captionTargetBox="[250,1378,214,1758]" captionTargetId="figure@20.[240,1391,210,1770]" captionTargetPageId="20" captionText="Figures 1421. Male genitalia of Cosmorrhyncha. 14) C. tonsana from Costa Rica, USNM slide 124,813. 15) C. ocelliferana from Brazil, USNM slide 124,821. 16) C. landryi from French Guiana, USNM slide 126,233. 17) C. parintina from Brazil, USNM slide 126,237. 18) C. macrospina from Brazil, USNM slide 124,792. 19) C. albistrigulana from Costa Rica, USNM slide 124,923. 20) C. albistrigulana from Brazil, USNM slide 124,818. 21) C. osana from Costa Rica, USNM slide 142,211." figureDoi="http://doi.org/10.5281/zenodo.3702993" httpUri="https://zenodo.org/record/3702993/files/figure.png" pageId="11" pageNumber="10">15</figureCitation>
,
<figureCitation id="3CFD2A5A8E06AF544403FF00FEDBFEA6" box="[321,350,250,274]" captionStart="Figures 2226" captionStartId="21.[192,285,1848,1870]" captionTargetBox="[216,1418,214,1818]" captionTargetId="figure@21.[215,1418,211,1821]" captionTargetPageId="21" captionText="Figures 2226. Female genitalia of Cosmorrhyncha. 22) C. tonsana from Brazil, USNM slide 145,538.23) C.ocelliferana from Brazil, USNM slide 124,775. 24) C. macrospina from Brazil, USNM slide 126,236. 25) C. albistrigulana from Guatemala, USNM slide 124,801. 26) C. osana from Costa Rica, USNM slide 142,202." figureDoi="http://doi.org/10.5281/zenodo.3702995" httpUri="https://zenodo.org/record/3702995/files/figure.png" pageId="11" pageNumber="10">23</figureCitation>
</paragraph>
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<treatmentCitation id="256710CE8E06AF544582FED2FD12FE8A" author="Walker, F." box="[192,663,296,318]" page="408" pageId="11" pageNumber="10" year="1863">
<taxonomicName id="63C64D5C8E06AF544582FED2FD12FE8A" ID-CoL="967C8" authority="Walker 1863: 408" authorityName="Walker" authorityPageNumber="408" authorityYear="1863" box="[192,663,296,318]" class="Insecta" family="Tortricidae" genus="Carpocapsa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E06AF544582FED2FE4FFE89" box="[192,458,296,317]" italics="true" pageId="11" pageNumber="10">Carpocapsa ocelliferana</emphasis>
<bibRefCitation id="C0574B2E8E06AF544492FED2FD12FE8A" author="Walker, F." box="[464,663,296,318]" journalOrPublisher="London" pageId="11" pageNumber="10" refId="ref15713" refString="Walker, F. 1863. List of specimens of lepidopterous insects in the collection of the British Museum, part. 28, Tortricites and Tineites: 287 - 561. London." title="List of specimens of lepidopterous insects in the collection of the British Museum, part. 28, Tortricites and Tineites: 287" type="book" year="1863">Walker 1863: 408</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<treatmentCitationGroup id="84D611F18E06AF544582FEBFFC4EFEEF" box="[192,971,325,347]" pageId="11" pageNumber="10">
<treatmentCitation id="256710CE8E06AF544582FEBFFD78FEEF" author="Powell, J. A. &amp; J. Razowski &amp; R. L. Brown" box="[192,765,325,347]" page="156" pageId="11" pageNumber="10" year="1995">
<taxonomicName id="63C64D5C8E06AF544582FEBFFD78FEEF" ID-CoL="6B7N9" authority=": Powell et al. 1995: 156" authorityName="Powell" authorityPageNumber="156" authorityYear="1995" box="[192,765,325,347]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E06AF544582FEBFFE71FEEE" box="[192,500,325,346]" italics="true" pageId="11" pageNumber="10">Cosmorrhyncha ocelliferana</emphasis>
:
<bibRefCitation id="C0574B2E8E06AF544740FEBFFD78FEEF" author="Powell, J. A. &amp; J. Razowski &amp; R. L. Brown" bookContentInfo="243 p." box="[514,765,325,347]" editor="Heppner, J. B." journalOrPublisher="Atlas of Neotropical Lepidoptera, checklist part II: Hyblaeoidea-Pyraloidea- Tortricoidea. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL" pageId="11" pageNumber="10" pagination="151 - 157" refId="ref15391" refString="Powell, J. A., J. Razowski, and R. L. Brown. 1995. Tortricidae: Olethreutinae. p. 151 - 157. In: J. B. Heppner, J. B. (ed.). Atlas of Neotropical Lepidoptera, checklist part II: Hyblaeoidea-Pyraloidea- Tortricoidea. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL. 243 p." title="Tortricidae: Olethreutinae" type="book chapter" volumeTitle="J. B" year="1995">Powell et al. 1995: 156</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="256710CE8E06AF54464AFEBCFC4DFEEF" author="Brown" box="[776,968,325,347]" page="218" pageId="11" pageNumber="10" year="2005">
<bibRefCitation id="C0574B2E8E06AF54464AFEBCFC4DFEEF" author="Brown" box="[776,968,325,347]" firstAuthor="Brown" pageId="11" pageNumber="10" refId="ref14316" refString="Brown, J. W. 2005. World catalogue of insects. Volume 5: Tortricidae (Lepidoptera). Apollo Books; Stenstrup. 741 p." type="book" year="2005">Brown 2005: 218</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="ECDC65548E06AF544582FE8EFDCAFC84" pageId="11" pageNumber="10" type="diagnosis">
<paragraph id="A47936DF8E06AF544582FE8EFB09FDF9" blockId="11.[192,1441,372,1042]" pageId="11" pageNumber="10">
<emphasis id="96B2EACD8E06AF544582FE8EFECAFE38" bold="true" box="[192,335,372,396]" pageId="11" pageNumber="10">Diagnosis.</emphasis>
Because the
<typeStatus id="7B7D887D8E06AF5444B1FE8EFDE4FE38" box="[499,609,372,396]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
lacks its abdomen, it is impossible to unequivocally associate other specimens with
<taxonomicName id="63C64D5C8E06AF5444C6FE6EFDB1FE18" box="[388,564,404,428]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E06AF5444C6FE6EFDB1FE18" box="[388,564,404,428]" italics="true" pageId="11" pageNumber="10">C. ocelliferana</emphasis>
</taxonomicName>
. Under this species name we include a large number of specimens from
<collectingCountry id="DCD1764F8E06AF544582FE4FFE8BFE79" box="[192,270,437,461]" name="Brazil" pageId="11" pageNumber="10">Brazil</collectingCountry>
,
<collectingCountry id="DCD1764F8E06AF54445FFE4FFE13FE79" box="[285,406,437,461]" name="Paraguay" pageId="11" pageNumber="10">Paraguay</collectingCountry>
, and
<collectingCountry id="DCD1764F8E06AF54449FFE4FFDD9FE79" box="[477,604,437,461]" name="Venezuela" pageId="11" pageNumber="10">Venezuela</collectingCountry>
that lack the distinctive white costal strigulae characteristic of
<taxonomicName id="63C64D5C8E06AF5440C7FE4FFEEAFE59" authorityName="Brown and Razowski" authorityYear="2020" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="albistrigulana">
<emphasis id="96B2EACD8E06AF5440C7FE4FFA1DFE78" box="[1413,1432,437,460]" italics="true" pageId="11" pageNumber="10">C</emphasis>
.
<emphasis id="96B2EACD8E06AF544582FE2FFEEAFE59" box="[192,367,469,493]" italics="true" pageId="11" pageNumber="10">albistrigulana</emphasis>
</taxonomicName>
and
<taxonomicName id="63C64D5C8E06AF5444E8FE2FFD96FE58" authorityName="Brown and Razowski" authorityYear="2020" box="[426,531,469,492]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="osana">
<emphasis id="96B2EACD8E06AF5444E8FE2FFD96FE58" box="[426,531,469,492]" italics="true" pageId="11" pageNumber="10">C. osana</emphasis>
</taxonomicName>
, and share similar male and female genitalia. The male genitalia are characterized by a more-or-less rounded outer margin of the valva and a flattened, finlike, hooked flange from the dorso-distal tip of the phallus. The female genitalia are characterized by a distinctly V-shaped posterior margin of sternum 7, similar to that of
<taxonomicName id="63C64D5C8E06AF544657FDCCFC40FDF9" authorityName="Brown and Razowski" authorityYear="2020" box="[789,965,566,589]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="macrospina">
<emphasis id="96B2EACD8E06AF544657FDCCFC40FDF9" box="[789,965,566,589]" italics="true" pageId="11" pageNumber="10">C. macrospina</emphasis>
</taxonomicName>
and
<taxonomicName id="63C64D5C8E06AF544140FDCCFB00FDF9" authorityName=": Brown" authorityYear="2005" box="[1026,1157,566,589]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="tonsana">
<emphasis id="96B2EACD8E06AF544140FDCCFB00FDF9" box="[1026,1157,566,589]" italics="true" pageId="11" pageNumber="10">C. tonsana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A47936DF8E06AF5445AAFDADFDCAFC84" blockId="11.[192,1441,372,1042]" pageId="11" pageNumber="10">
In the
<typeStatus id="7B7D887D8E06AF544479FDACFE2CFDDA" box="[315,425,598,622]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
, the bend in the distal 0.4 of the costa of the forewing is at a greater angle than in other specimens associated with the name; however, this feature appears to be somewhat variable among individuals throughout its geographic range.
<location id="A11960048E06AF5447B0FD6DFC99FD1A" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E06AF5447B0FD6DFC99FD1A" box="[754,796,663,686]" country="Brazil" municipality="Cacaulandia" name="For" pageId="11" pageNumber="10" stateProvince="Rondonia">For</location>
example, in a pair of specimens from
<collectingMunicipality id="441DACA58E06AF544043FD6DFA19FD1B" box="[1281,1436,663,687]" pageId="11" pageNumber="10">Cacaulandia</collectingMunicipality>
,
<collectingRegion id="6602F83D8E06AF544582FD4DFEB0FD7B" box="[192,309,695,719]" country="Brazil" name="Rondonia" pageId="11" pageNumber="10">Rondônia</collectingRegion>
,
<collectingCountry id="DCD1764F8E06AF544403FD4DFE08FD7B" box="[321,397,695,719]" name="Brazil" pageId="11" pageNumber="10">Brazil</collectingCountry>
, collected on the same date (Becker no. 80239), one has a distinctly angled costa of the forewing as in the
<typeStatus id="7B7D887D8E06AF5444D8FD2DFD80FD5B" box="[410,517,727,751]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="63C64D5C8E06AF544765FD2DFD57FD5B" box="[551,722,727,751]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E06AF544765FD2DFD57FD5B" box="[551,722,727,751]" italics="true" pageId="11" pageNumber="10">C. ocelliferana</emphasis>
</taxonomicName>
(
<figureCitation id="3CFD2A5A8E06AF5447A3FD2DFCA0FD5B" box="[737,805,727,751]" captionStart="Figures 613" captionStartId="19.[196,291,1694,1716]" captionTargetBox="[195,1444,291,1661]" captionTargetId="figure@19.[192,1444,291,1661]" captionTargetPageId="19" captionText="Figures 613. Adults of Cosmorrhyncha. 6) C. tonsana from Costa Rica. 7) C. ocelliferana from Brazil (image of lectotype from NHMUK). 8) C. landryi from French Guiana. 9) C. parintina from Brazil. 10) C. macrospina female from Brazil. 11) C. macrospina male from Brazil (apex of right forewing damaged). 12) C. albistrigulana from Brazil. 13) C. osana from Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3702991" httpUri="https://zenodo.org/record/3702991/files/figure.png" pageId="11" pageNumber="10">Fig. 7</figureCitation>
), whereas the other has a more evenly rounded costa.
<figureCitation id="3CFD2A5A8E06AF544582FD0DFEADFCBB" box="[192,296,759,783]" captionStart="Figures 613" captionStartId="19.[196,291,1694,1716]" captionTargetBox="[195,1444,291,1661]" captionTargetId="figure@19.[192,1444,291,1661]" captionTargetPageId="19" captionText="Figures 613. Adults of Cosmorrhyncha. 6) C. tonsana from Costa Rica. 7) C. ocelliferana from Brazil (image of lectotype from NHMUK). 8) C. landryi from French Guiana. 9) C. parintina from Brazil. 10) C. macrospina female from Brazil. 11) C. macrospina male from Brazil (apex of right forewing damaged). 12) C. albistrigulana from Brazil. 13) C. osana from Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3702991" httpUri="https://zenodo.org/record/3702991/files/figure.png" pageId="11" pageNumber="10">Figure 7</figureCitation>
, an image of the
<typeStatus id="7B7D887D8E06AF5444BBFD0DFDE2FCBB" box="[505,615,759,783]" pageId="11" pageNumber="10" type="lectotype">lectotype</typeStatus>
, NHMUK, is considerably darker brown than other images, but this is partially an artifact of lighting.
</paragraph>
</subSubSection>
<subSubSection id="ECDC65548E06AF5445AAFCC2FE78F8BC" pageId="11" pageNumber="10" type="description">
<paragraph id="A47936DF8E06AF5445AAFCC2FA25FBA6" blockId="11.[192,1441,372,1042]" pageId="11" pageNumber="10">
In the male genitalia of the specimens examined, there is subtle variation in the shape of the valva and in the number of spines along the sacculus, the latter often variable from the left to the right valva of the same specimen. In the female genitalia, the shape of the signum is somewhat variable, as well. Hence, it is likely that the specimens examined represent a species complex rather than a single species, and the trees (
<figureCitation id="3CFD2A5A8E06AF54458BFC43FEA2FC65" box="[201,295,953,977]" captionStart="Figures 13" captionStartId="7.[192,287,1743,1765]" captionTargetBox="[219,1418,229,1705]" captionTargetId="figure@7.[192,1440,213,1734]" captionTargetPageId="7" captionText="Figures 13. Neighbor-joining trees. 1) Tree based on all available sequences of Cosmorrhyncha (n = 28), regardless of sequence length. 2) Tree based on sequences excluding the the 200bp segment that could not be amplified for several specimens of Cosmorrhyncha (n = 24). 3) Tree based on all sequences longer than 500bp (n = 23). [Bootstrap values ≥70 shown at nodes; BOLD process IDs or GenBank accession numbers next to species names.]" figureDoi="http://doi.org/10.5281/zenodo.3702987" httpUri="https://zenodo.org/record/3702987/files/figure.png" pageId="11" pageNumber="10">Fig. 13</figureCitation>
) also suggests this, although based on short sequences. In specimens from
<collectingCountry id="DCD1764F8E06AF5441D1FC43FB59FC65" box="[1171,1244,953,977]" name="Brazil" pageId="11" pageNumber="10">Brazil</collectingCountry>
the signum may have a pair of long slender spines (
<figureCitation id="3CFD2A5A8E06AF544718FC23FD29FC45" box="[602,684,985,1009]" captionStart="Figures 2226" captionStartId="21.[192,285,1848,1870]" captionTargetBox="[216,1418,214,1818]" captionTargetId="figure@21.[215,1418,211,1821]" captionTargetPageId="21" captionText="Figures 2226. Female genitalia of Cosmorrhyncha. 22) C. tonsana from Brazil, USNM slide 145,538.23) C.ocelliferana from Brazil, USNM slide 124,775. 24) C. macrospina from Brazil, USNM slide 126,236. 25) C. albistrigulana from Guatemala, USNM slide 124,801. 26) C. osana from Costa Rica, USNM slide 142,202." figureDoi="http://doi.org/10.5281/zenodo.3702995" httpUri="https://zenodo.org/record/3702995/files/figure.png" pageId="11" pageNumber="10">Fig. 23</figureCitation>
) or a more typical spindle-shaped signum subtended by a small pair of short slender spines; rarely there is only a spindle-shaped signum (e.g., in specimens from
<collectingCountry id="DCD1764F8E06AF54405CFC00FA11FBA6" box="[1310,1428,1018,1042]" name="Paraguay" pageId="11" pageNumber="10">Paraguay</collectingCountry>
).
</paragraph>
<paragraph id="A47936DF8E06AF544582FBD3FE78F8BC" blockId="11.[192,1441,1065,1800]" pageId="11" pageNumber="10">
<emphasis id="96B2EACD8E06AF544582FBD3FE0DFBF5" bold="true" box="[192,392,1065,1089]" pageId="11" pageNumber="10">Redescription.</emphasis>
<emphasis id="96B2EACD8E06AF5444CCFBD3FE4AFBF5" box="[398,463,1065,1089]" italics="true" pageId="11" pageNumber="10">Head</emphasis>
. Reddish-brown to brown; labial palpus reddish brown, dorsum of second segment with bluish-black, narrow, longitudinal stripe; third segment mostly bluish black, slightly metallic.
<emphasis id="96B2EACD8E06AF544582FB90FE9DFB36" box="[192,280,1130,1154]" italics="true" pageId="11" pageNumber="10">Thorax</emphasis>
. Nota reddish-brown to brown. Forewing (
<figureCitation id="3CFD2A5A8E06AF544659FB90FCE5FB36" box="[795,864,1130,1154]" captionStart="Figures 613" captionStartId="19.[196,291,1694,1716]" captionTargetBox="[195,1444,291,1661]" captionTargetId="figure@19.[192,1444,291,1661]" captionTargetPageId="19" captionText="Figures 613. Adults of Cosmorrhyncha. 6) C. tonsana from Costa Rica. 7) C. ocelliferana from Brazil (image of lectotype from NHMUK). 8) C. landryi from French Guiana. 9) C. parintina from Brazil. 10) C. macrospina female from Brazil. 11) C. macrospina male from Brazil (apex of right forewing damaged). 12) C. albistrigulana from Brazil. 13) C. osana from Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3702991" httpUri="https://zenodo.org/record/3702991/files/figure.png" pageId="11" pageNumber="10">Fig. 7</figureCitation>
) length 6.07.0 mm (
<emphasis id="96B2EACD8E06AF544120FB91FB0DFB36" box="[1122,1160,1131,1154]" italics="true" pageId="11" pageNumber="10">n =</emphasis>
6), pale reddish-brown with faint refractive pale-blue striations and scale patches, a small rounded, upraised patch of silverwhite scales near distal end of discal cell, surrounded by small dark-brown circular patch; costa reflexed downward at approximately 0.6 length from base to apex; distal 0.2 of costa faint orange with few black strigulae; median part of the costa weakly tinged with orange; two pale-cream spots at mid-costa; costa without well-defined areas of dense silver-white striae. Fringe pale-orange tinged with brown, black at apex. Hindwing brown; male with long, well defined anal roll. Fringe brown, paler in the anal area.
<emphasis id="96B2EACD8E06AF544582FAB6FEB6FAD0" box="[192,307,1356,1380]" italics="true" pageId="11" pageNumber="10">Abdomen</emphasis>
. Male genitalia (
<figureCitation id="3CFD2A5A8E06AF544749FAB6FDE7FAD0" box="[523,610,1356,1380]" captionStart="Figures 1421" captionStartId="20.[192,285,1792,1814]" captionTargetBox="[250,1378,214,1758]" captionTargetId="figure@20.[240,1391,210,1770]" captionTargetPageId="20" captionText="Figures 1421. Male genitalia of Cosmorrhyncha. 14) C. tonsana from Costa Rica, USNM slide 124,813. 15) C. ocelliferana from Brazil, USNM slide 124,821. 16) C. landryi from French Guiana, USNM slide 126,233. 17) C. parintina from Brazil, USNM slide 126,237. 18) C. macrospina from Brazil, USNM slide 124,792. 19) C. albistrigulana from Costa Rica, USNM slide 124,923. 20) C. albistrigulana from Brazil, USNM slide 124,818. 21) C. osana from Costa Rica, USNM slide 142,211." figureDoi="http://doi.org/10.5281/zenodo.3702993" httpUri="https://zenodo.org/record/3702993/files/figure.png" pageId="11" pageNumber="10">Fig. 15</figureCitation>
) with uncus short, stout, apically bifid, with dense cluster of long spines on each bifurcation; tegumen with rounded dorsal half; socius digitate, slightly broader basally, pendant, finely hairy; valva mostly parallel-sided in basal 0.5, upturned, attenuate, densely spined in distal 0.5; cucullus with a few strong setae along ventral edge in basal half, one basal and one subbasal setae conspicuously larger; patch of long, slender, hairlike setae at venter immediately basad of cucullus; a distinct triangular subbasal process of valva bearing a single spine; juxta shield-shaped; phallus short stout, vesica lacking cornuti. Female genitalia (
<figureCitation id="3CFD2A5A8E06AF544603F9F4FC11F992" box="[833,916,1550,1574]" captionStart="Figures 2226" captionStartId="21.[192,285,1848,1870]" captionTargetBox="[216,1418,214,1818]" captionTargetId="figure@21.[215,1418,211,1821]" captionTargetPageId="21" captionText="Figures 2226. Female genitalia of Cosmorrhyncha. 22) C. tonsana from Brazil, USNM slide 145,538.23) C.ocelliferana from Brazil, USNM slide 124,775. 24) C. macrospina from Brazil, USNM slide 126,236. 25) C. albistrigulana from Guatemala, USNM slide 124,801. 26) C. osana from Costa Rica, USNM slide 142,202." figureDoi="http://doi.org/10.5281/zenodo.3702995" httpUri="https://zenodo.org/record/3702995/files/figure.png" pageId="11" pageNumber="10">Fig. 23</figureCitation>
) with papillae anales unmodified; sterigma mostly membranous with a strongly sclerotized, mesal, suborbicular plate and a pair of narrow lateral flanges postero-laterally; antrum long, spindle-shaped, occupying posterior 0.33 of ductus bursae; ductus bursae relatively narrow, membranous beyond antrum, gradually broadening to corpus bursae without distinct junction; corpus bursae irregularly pear-shaped, weakly and evenly punctuate throughout; signum variable with one or two spindle-shaped blades from an irregularly rounded, sclerotized patch; ventro-posterior margin of segment 7 with deep V-shaped excavation mesally, resulting in a pair of rounded, sublateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="ECDC65548E06AF534582F8DAFD3DFDA3" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="materials_examined">
<paragraph id="A47936DF8E06AF544582F8DAFBD2F88C" blockId="11.[192,1111,1824,1848]" box="[192,1111,1824,1848]" pageId="11" pageNumber="10">
<emphasis id="96B2EACD8E06AF544582F8DAFE9CF88C" bold="true" box="[192,281,1824,1848]" pageId="11" pageNumber="10">Types.</emphasis>
<materialsCitation id="14AE3C828E06AF544462F8DAFBD2F88C" ID-GBIF-Occurrence="2575116521" box="[288,1111,1824,1848]" collectionCode="NHMUK" country="Brazil" location="Amazonas" municipality="Ega" pageId="11" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="7B7D887D8E06AF544462F8DAFE12F88C" box="[288,407,1824,1848]" pageId="11" pageNumber="10" type="lectotype">Lectotype</typeStatus>
<specimenCount id="B2C0FD568E06AF5444DDF8DAFE36F883" box="[415,435,1824,1847]" pageId="11" pageNumber="10" type="male"></specimenCount>
(abdomen missing),
<collectingCountry id="DCD1764F8E06AF5447F2F8DAFD7BF88C" box="[688,766,1824,1848]" name="Brazil" pageId="11" pageNumber="10">Brazil</collectingCountry>
,
<location id="A11960048E06AF544648F8DAFC09F88C" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E06AF544648F8DAFC09F88C" box="[778,908,1824,1848]" country="Brazil" municipality="Ega" name="Amazonas" pageId="11" pageNumber="10">
<collectingRegion id="6602F83D8E06AF544648F8DAFC09F88C" box="[778,908,1824,1848]" country="Brazil" name="Amazonas" pageId="11" pageNumber="10">Amazonas</collectingRegion>
</location>
,
<collectingMunicipality id="441DACA58E06AF5446D5F8DAFC42F88C" box="[919,967,1824,1848]" pageId="11" pageNumber="10">Ega</collectingMunicipality>
(
<collectionCode id="C2D7AE1A8E06AF544695F8DAFBC9F88C" box="[983,1100,1824,1848]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="10" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="A47936DF8E06AF544582F8AAFA25F83C" blockId="11.[192,1440,1872,1929]" pageId="11" pageNumber="10">
<emphasis id="96B2EACD8E06AF544582F8AAFE70F8DC" bold="true" box="[192,501,1872,1896]" pageId="11" pageNumber="10">Additional specimens.</emphasis>
<materialsCitation id="14AE3C828E06AF544743F8AAFE02F83D" ID-GBIF-Occurrence="2575116546" collectingDate="1884-08" collectionCode="USNM" country="Brazil" location="Ceara" pageId="11" pageNumber="10" specimenCount="1" specimenCount-female="1" stateProvince="Ceara">
<collectingCountry id="DCD1764F8E06AF544743F8AAFDE9F8DC" box="[513,620,1872,1896]" name="Brazil" pageId="11" pageNumber="10">BRAZIL</collectingCountry>
:
<collectingRegion id="6602F83D8E06AF544738F8AAFD4DF8DC" box="[634,712,1872,1896]" country="Brazil" name="Ceara" pageId="11" pageNumber="10">Ceara</collectingRegion>
: [no further locality],
<date id="D078101F8E06AF5446A3F8AAFBDDF8DC" box="[993,1112,1872,1896]" pageId="11" pageNumber="10" value="1884-08">
<collectingDate id="C03CE9F78E06AF5446A3F8AAFBDDF8DC" box="[993,1112,1872,1896]" pageId="11" pageNumber="10" value="1884-08">Aug 1884</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E06AF54412DF8AAFB0BF8DC" box="[1135,1166,1872,1896]" pageId="11" pageNumber="10" type="female">1♀</specimenCount>
), Leech,
<collectionCode id="C2D7AE1A8E06AF544044F8AAFADEF8DC" box="[1286,1371,1872,1896]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="10" type="Museum">USNM</collectionCode>
slide 72,364 (
<collectionCode id="C2D7AE1A8E06AF544463F88BFEFEF83D" box="[289,379,1905,1929]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="10" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E06AF5444CCF88AFA25F83C" ID-GBIF-Occurrence="2575116559" box="[398,1440,1904,1929]" collectingDate="1985-03-15" collectionCode="USNM" collectorName="V. Becker" elevation="1000" location="Planaltina" municipality="Planaltina" pageId="11" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Distrito Federal">
<collectingRegion id="6602F83D8E06AF5444CCF88AFDD2F83C" box="[398,599,1904,1928]" country="Brazil" name="Distrito Federal" pageId="11" pageNumber="10">Distrito Federal</collectingRegion>
:
<collectingMunicipality id="441DACA58E06AF544720F88AFD63F83C" box="[610,742,1904,1928]" pageId="11" pageNumber="10">Planaltina</collectingMunicipality>
,
<quantity id="633E9B3A8E06AF5447B1F88BFCCAF83C" box="[755,847,1905,1929]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="10" unit="m" value="1000.0">
<elevation id="2FEBD1EC8E06AF5447B1F88BFCCAF83C" box="[755,847,1905,1929]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="10" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="D078101F8E06AF54461EF88AFC72F83C" box="[860,1015,1904,1928]" pageId="11" pageNumber="10" value="1985-03-15">
<collectingDate id="C03CE9F78E06AF54461EF88AFC72F83C" box="[860,1015,1904,1928]" pageId="11" pageNumber="10" value="1985-03-15">15 Mar 1985</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E06AF54414AF88BFBAEF83D" box="[1032,1067,1905,1929]" pageId="11" pageNumber="10" type="male">1♂</specimenCount>
),
<collectorName id="093353098E06AF544100F88BFB39F83C" box="[1090,1212,1904,1928]" pageId="11" pageNumber="10">V. Becker</collectorName>
, Becker No. 57384
</materialsCitation>
</paragraph>
<paragraph id="A47936DF8E01AF534582FF20FD3DFDA3" blockId="12.[192,1441,217,666]" pageId="12" pageNumber="11">
(
<collectionCode id="C2D7AE1A8E01AF53458AFF20FEA7FF46" box="[200,290,218,242]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
<materialsCitation id="14AE3C828E01AF534475FF23FEFAFEA6" ID-GBIF-Occurrence="2575116550" collectingDate="1990-11-19" collectingDateMax="1990-11-27" collectingDateMin="1990-11-19" collectionCode="USNM" collectorName="V. Becker &amp; G. Dubois" elevation="150" location="Acailandia" municipality="Maranhao" pageId="12" pageNumber="11" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="441DACA58E01AF534475FF23FE3CFF45" box="[311,441,217,241]" pageId="12" pageNumber="11">Maranhão</collectingMunicipality>
:
<location id="A11960048E01AF534484FF23FDC8FF45" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF534484FF23FDC8FF45" box="[454,589,217,241]" municipality="Maranhao" name="Acailandia" pageId="12" pageNumber="11">Acailandia</location>
,
<quantity id="633E9B3A8E01AF534718FF23FD2CFF45" box="[602,681,217,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="150.0">
<elevation id="2FEBD1EC8E01AF534718FF23FD2CFF45" box="[602,681,217,241]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="D078101F8E01AF5347F5FF20FC07FF46" box="[695,898,218,242]" pageId="12" pageNumber="11" value="1990-11-19" valueMax="1990-11-27" valueMin="1990-11-19">
<collectingDate id="C03CE9F78E01AF5347F5FF20FC07FF46" box="[695,898,218,242]" pageId="12" pageNumber="11" value="1990-11-19" valueMax="1990-11-27" valueMin="1990-11-19">1927 Nov 1990</collectingDate>
</date>
, (
<specimenCount id="B2C0FD568E01AF5346D9FF20FC3FFF46" box="[923,954,218,242]" pageId="12" pageNumber="11" type="female">1♀</specimenCount>
),
<collectorName id="093353098E01AF534696FF20FBC8FF45" box="[980,1101,217,241]" pageId="12" pageNumber="11">V. Becker</collectorName>
and
<collectorName id="093353098E01AF5341CFFF20FA8FFF45" box="[1165,1290,217,242]" pageId="12" pageNumber="11">G. Dubois</collectorName>
, Becker No. 77692 (
<collectionCode id="C2D7AE1A8E01AF53445AFF00FEF7FEA6" box="[280,370,250,274]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF5344C7FF01FD2CFE87" ID-GBIF-Occurrence="2575116561" collectingDate="1993-10-26" collectionCode="USNM" collectorName="V. Becker" location="Chapada dos Guimaraes" municipality="Mato Grosso" pageId="12" pageNumber="11" specimenCount="2" specimenCount-female="2">
<collectingMunicipality id="441DACA58E01AF5344C7FF01FDA7FEA6" box="[389,546,250,274]" pageId="12" pageNumber="11">Mato Grosso</collectingMunicipality>
:
<location id="A11960048E01AF53476EFF00FCDFFEA6" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF53476EFF00FCDFFEA6" box="[556,858,250,274]" municipality="Mato Grosso" name="Chapada dos Guimaraes" pageId="12" pageNumber="11">Chapada dos Guimaraes</location>
,
<date id="D078101F8E01AF534624FF00FC73FEA6" box="[870,1014,250,274]" pageId="12" pageNumber="11" value="1993-10-26">
<collectingDate id="C03CE9F78E01AF534624FF00FC73FEA6" box="[870,1014,250,274]" pageId="12" pageNumber="11" value="1993-10-26">26 Oct 1993</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF534145FF00FBACFEA6" box="[1031,1065,250,274]" pageId="12" pageNumber="11" type="female">2♀</specimenCount>
),
<collectorName id="093353098E01AF53417DFF01FB3DFEA6" box="[1087,1208,250,274]" pageId="12" pageNumber="11">V. Becker</collectorName>
, Becker No. 89332,
<collectionCode id="C2D7AE1A8E01AF534582FEE1FE90FE87" box="[192,277,283,307]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
slides 126,290, 126,374 (
<collectionCode id="C2D7AE1A8E01AF534700FEE1FD19FE87" box="[578,668,283,307]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF5347ECFEE1FCFAFEE7" ID-GBIF-Occurrence="2575116511" collectingDate="1983-11-03" collectionCode="USNM" collectorName="V. O. Becker" location="Unai" municipality="Minas Gerais" pageId="12" pageNumber="11" specimenCount="704" specimenCount-male="704">
<collectingMunicipality id="441DACA58E01AF5347ECFEE1FCD3FE87" box="[686,854,283,307]" pageId="12" pageNumber="11">Minas Gerais</collectingMunicipality>
:
<location id="A11960048E01AF534622FEE1FC25FE87" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF534622FEE1FC25FE87" box="[864,928,283,307]" municipality="Minas Gerais" name="Unai" pageId="12" pageNumber="11">Unai</location>
,
<quantity id="633E9B3A8E01AF5346E8FEE1FC73FE86" box="[938,1014,283,307]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="11" unit="m" value="700.0">
<specimenCount id="B2C0FD568E01AF5346E8FEE1FC73FE86" box="[938,1014,283,307]" pageId="12" pageNumber="11" type="male">700 m</specimenCount>
</quantity>
,
<date id="D078101F8E01AF534140FEE1FB0CFE87" box="[1026,1161,283,307]" pageId="12" pageNumber="11" value="1983-11-03">
<collectingDate id="C03CE9F78E01AF534140FEE1FB0CFE87" box="[1026,1161,283,307]" pageId="12" pageNumber="11" value="1983-11-03">3 Nov 1983</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF5341DAFEE1FB39FE87" box="[1176,1212,283,307]" pageId="12" pageNumber="11" type="male">4♂</specimenCount>
),
<collectorName id="093353098E01AF534193FEE1FAEEFE87" box="[1233,1387,283,307]" pageId="12" pageNumber="11">V. O. Becker</collectorName>
, col- lection no. 49881 (
<collectionCode id="C2D7AE1A8E01AF5344DBFEC1FE77FEE7" box="[409,498,315,339]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
),
<collectionCode id="C2D7AE1A8E01AF534747FEC1FDDFFEE7" box="[517,602,315,339]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
slides 124,795, 126,235.
</materialsCitation>
<materialsCitation id="14AE3C828E01AF5346C6FEC1FD95FEC0" ID-GBIF-Occurrence="2575116529" collectingDate="1984-01" collectionCode="USNM" collectorName="V. Becker" location="Belem" municipality="Para" pageId="12" pageNumber="11" specimenCount="22" specimenCount-male="22">
<collectingMunicipality id="441DACA58E01AF5346C6FEC1FC47FEE7" box="[900,962,315,339]" pageId="12" pageNumber="11">Pará</collectingMunicipality>
:
<location id="A11960048E01AF53468EFEC1FB98FEE7" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF53468EFEC1FB98FEE7" box="[972,1053,315,339]" municipality="Para" name="Belem" pageId="12" pageNumber="11">Belém</location>
,
<quantity id="633E9B3A8E01AF534165FEC1FBE1FEE7" box="[1063,1124,315,339]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="11" unit="m" value="20.0">
<specimenCount id="B2C0FD568E01AF534165FEC1FBE1FEE7" box="[1063,1124,315,339]" pageId="12" pageNumber="11" type="male">20 m</specimenCount>
</quantity>
,
<date id="D078101F8E01AF53412CFEC6FB58FEE7" box="[1134,1245,315,339]" pageId="12" pageNumber="11" value="1984-01">
<collectingDate id="C03CE9F78E01AF53412CFEC6FB58FEE7" box="[1134,1245,315,339]" pageId="12" pageNumber="11" value="1984-01">Jan 1984</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF5341A9FEC1FA94FEE7" box="[1259,1297,315,339]" pageId="12" pageNumber="11" type="male">2♂</specimenCount>
),
<collectorName id="093353098E01AF534061FEC6FA19FEE7" box="[1315,1436,315,339]" pageId="12" pageNumber="11">V. Becker</collectorName>
, Becker No. 47165 (
<collectionCode id="C2D7AE1A8E01AF5344EBFEA6FD86FEC0" box="[425,515,348,372]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF534755FEA6FC8FFE21" ID-GBIF-Occurrence="2575116549" collectingDate="1991-11" collectionCode="USNM" collectorName="V. Becker" elevation="140" location="Cacaulandia" municipality="Rondonia" pageId="12" pageNumber="11" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingMunicipality id="441DACA58E01AF534755FEA6FD0AFEC0" box="[535,655,348,372]" pageId="12" pageNumber="11">Rondônia</collectingMunicipality>
:
<location id="A11960048E01AF5347D9FEA6FCBDFEC0" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF5347D9FEA6FCBDFEC0" box="[667,824,348,372]" municipality="Rondonia" name="Cacaulandia" pageId="12" pageNumber="11">Cacaulandia</location>
,
<quantity id="633E9B3A8E01AF534607FEA6FC16FEC7" box="[837,915,348,372]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="11" unit="m" value="140.0">
<elevation id="2FEBD1EC8E01AF534607FEA6FC16FEC7" box="[837,915,348,372]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="11" unit="m" value="140.0">140 m</elevation>
</quantity>
,
<date id="D078101F8E01AF5346E2FEA6FB91FEC0" box="[928,1044,348,372]" pageId="12" pageNumber="11" value="1991-11">
<collectingDate id="C03CE9F78E01AF5346E2FEA6FB91FEC0" box="[928,1044,348,372]" pageId="12" pageNumber="11" value="1991-11">Nov 1991</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF534164FEA6FBCCFEC0" box="[1062,1097,348,372]" pageId="12" pageNumber="11" type="male">1♂</specimenCount>
,
<specimenCount id="B2C0FD568E01AF534115FEA6FBFDFEC0" box="[1111,1144,348,372]" pageId="12" pageNumber="11" type="female">1♀</specimenCount>
),
<collectorName id="093353098E01AF5341D0FEA6FA89FEC0" box="[1170,1292,348,372]" pageId="12" pageNumber="11">V. Becker</collectorName>
, Becker No. 80239,
<collectionCode id="C2D7AE1A8E01AF53445BFE87FEEBFE21" box="[281,366,381,405]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
slides 124,821, 126,285 (
<collectionCode id="C2D7AE1A8E01AF5347E6FE87FD7BFE21" box="[676,766,381,405]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF534650FE87FB9BFE01" ID-GBIF-Occurrence="2575116556" collectingDate="1961-11" collectionCode="USNM" collectorName="F. Plaumann" elevation="400" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" municipality="Santa Catarina" pageId="12" pageNumber="11" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="441DACA58E01AF534650FE87FC50FE21" box="[786,981,381,405]" pageId="12" pageNumber="11">Santa Catarina</collectingMunicipality>
:
<location id="A11960048E01AF5346A3FE87FB1FFE21" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF5346A3FE87FB1FFE21" box="[993,1178,381,405]" latitude="-27.183332" longLatPrecision="1238" longitude="-52.383335" municipality="Santa Catarina" name="Nova Teutonia" pageId="12" pageNumber="11">Nova Teutonia</location>
,
<quantity id="633E9B3A8E01AF5341E4FE86FAB5FE20" box="[1190,1328,380,404]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="11" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">
<elevation id="2FEBD1EC8E01AF5341E4FE86FAB5FE20" box="[1190,1328,380,404]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="11" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</elevation>
</quantity>
,
<geoCoordinate id="C1F250188E01AF53407FFE81FA1EFE20" box="[1341,1435,379,404]" degrees="27" direction="south" minutes="11" orientation="latitude" pageId="12" pageNumber="11" precision="925" value="-27.183332">27°11S</geoCoordinate>
,
<geoCoordinate id="C1F250188E01AF534582FE66FEADFE01" box="[192,296,412,437]" degrees="52" direction="west" minutes="23" orientation="longitude" pageId="12" pageNumber="11" precision="925" value="-52.383335">52°23W</geoCoordinate>
,
<date id="D078101F8E01AF534477FE64FE2FFE01" box="[309,426,413,437]" pageId="12" pageNumber="11" value="1961-11">
<collectingDate id="C03CE9F78E01AF534477FE64FE2FFE01" box="[309,426,413,437]" pageId="12" pageNumber="11" value="1961-11">Nov 1961</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF5344FFFE67FE58FE01" box="[445,477,413,437]" pageId="12" pageNumber="11" type="female">1♀</specimenCount>
),
<collectorName id="093353098E01AF5344B4FE64FD1FFE01" box="[502,666,413,437]" pageId="12" pageNumber="11">F. Plaumann</collectorName>
,
<collectionCode id="C2D7AE1A8E01AF5347EAFE67FD78FE01" box="[680,765,413,437]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
slide 124,784 (
<collectionCode id="C2D7AE1A8E01AF5346FAFE67FB97FE01" box="[952,1042,413,437]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF534164FE67FB83FE62" ID-GBIF-Occurrence="2575116506" collectingDate="1986-03-15" collectingDateMax="1986-03-18" collectingDateMin="1986-03-15" collectionCode="USNM" collectorName="M. Pogue &amp; M. Solis" location="Parque Nacional Ybycui" pageId="12" pageNumber="11" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="PARAGUAY">
<collectingRegion id="6602F83D8E01AF534164FE67FB43FE01" box="[1062,1222,413,437]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="11">PARAGUAY</collectingRegion>
:
<location id="A11960048E01AF534190FE64FE92FE62" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF534190FE64FE92FE62" name="Parque Nacional Ybycui" pageId="12" pageNumber="11" stateProvince="PARAGUAY">Parque Nacional Ybycui</location>
,
<date id="D078101F8E01AF534467FE44FE74FE62" box="[293,497,446,470]" pageId="12" pageNumber="11" value="1986-03-15" valueMax="1986-03-18" valueMin="1986-03-15">
<collectingDate id="C03CE9F78E01AF534467FE44FE74FE62" box="[293,497,446,470]" pageId="12" pageNumber="11" value="1986-03-15" valueMax="1986-03-18" valueMin="1986-03-15">1518 Mar 1986</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF534741FE44FDADFE62" box="[515,552,446,470]" pageId="12" pageNumber="11" type="male">5♂</specimenCount>
,
<specimenCount id="B2C0FD568E01AF534775FE44FDDCFE62" box="[567,601,446,470]" pageId="12" pageNumber="11" type="female">9♀</specimenCount>
),
<collectorName id="093353098E01AF534731FE44FD6DFE62" box="[627,744,446,470]" pageId="12" pageNumber="11">M. Pogue</collectorName>
and
<collectorName id="093353098E01AF53466BFE44FC0BFE62" box="[809,910,446,470]" pageId="12" pageNumber="11">M. Solis</collectorName>
(
<collectionCode id="C2D7AE1A8E01AF5346E2FE44FC7FFE62" box="[928,1018,446,470]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF53414DFE44FBE0FE43" ID-GBIF-Occurrence="2575116532" collectingDate="1969-02-22" collectingDateMax="1969-02-26" collectingDateMin="1969-02-22" collectionCode="USNM" collectorName="Duckworth &amp; Dietz" country="Venezuela" location="Barinitas" municipality="Barin" pageId="12" pageNumber="11" specimenCount="2" specimenCount-female="2">
<collectingCountry id="DCD1764F8E01AF53414DFE44FB45FE62" box="[1039,1216,446,470]" name="Venezuela" pageId="12" pageNumber="11">VENEZUELA</collectingCountry>
:
<collectingMunicipality id="441DACA58E01AF53418FFE44FA92FE62" box="[1229,1303,446,470]" pageId="12" pageNumber="11">Barin</collectingMunicipality>
:
<location id="A11960048E01AF534067FE44FA19FE62" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF534067FE44FA19FE62" box="[1317,1436,446,470]" country="Venezuela" municipality="Barin" name="Barinitas" pageId="12" pageNumber="11">Barinitas</location>
,
<date id="D078101F8E01AF534582FE24FE00FE42" box="[192,389,478,502]" pageId="12" pageNumber="11" value="1969-02-22" valueMax="1969-02-26" valueMin="1969-02-22">
<collectingDate id="C03CE9F78E01AF534582FE24FE00FE42" box="[192,389,478,502]" pageId="12" pageNumber="11" value="1969-02-22" valueMax="1969-02-26" valueMin="1969-02-22">2226 Feb 1969</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF5344DAFE25FE3FFE43" box="[408,442,479,503]" pageId="12" pageNumber="11" type="female">2♀</specimenCount>
),
<collectorName id="093353098E01AF534490FE24FDDEFE42" box="[466,603,478,502]" pageId="12" pageNumber="11">Duckworth</collectorName>
and
<collectorName id="093353098E01AF5347D9FE24FD5AFE42" box="[667,735,478,502]" pageId="12" pageNumber="11">Dietz</collectorName>
,
<collectionCode id="C2D7AE1A8E01AF5347AFFE25FCC7FE43" box="[749,834,479,503]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
slide 126,234 (
<collectionCode id="C2D7AE1A8E01AF5346BCFE25FBDDFE43" box="[1022,1112,479,503]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation id="14AE3C828E01AF53412FFE25FD3DFDA3" ID-GBIF-Occurrence="2575116508" collectingDate="1967-07-08" collectingDateMax="1967-07-14" collectingDateMin="1967-07-08" collectionCode="USNM" collectorName="R. W. Poole" location="Rancho Grande" pageId="12" pageNumber="11" specimenCount="4" specimenCount-male="4" stateProvince="Aragua">
<collectingRegion id="6602F83D8E01AF53412FFE25FB49FE43" box="[1133,1228,479,503]" country="Venezuela" name="Aragua" pageId="12" pageNumber="11">Aragua</collectingRegion>
:
<location id="A11960048E01AF53419AFE24FA19FE42" LSID="urn:lsid:plazi:treatment:2C6F87C98E06AF534582FF23FBF6FD2D:A11960048E01AF53419AFE24FA19FE42" box="[1240,1436,478,502]" name="Rancho Grande" pageId="12" pageNumber="11" stateProvince="Aragua">Rancho Grande</location>
,
<date id="D078101F8E01AF534582FE05FEE9FDA3" box="[192,364,511,535]" pageId="12" pageNumber="11" value="1967-07-08" valueMax="1967-07-14" valueMin="1967-07-08">
<collectingDate id="C03CE9F78E01AF534582FE05FEE9FDA3" box="[192,364,511,535]" pageId="12" pageNumber="11" value="1967-07-08" valueMax="1967-07-14" valueMin="1967-07-08">814 Jul 1967</collectingDate>
</date>
(
<specimenCount id="B2C0FD568E01AF53443EFE05FE24FDA3" box="[380,417,511,535]" pageId="12" pageNumber="11" type="male">4♂</specimenCount>
),
<collectorName id="093353098E01AF5344F4FDFAFDC6FDA3" box="[438,579,511,535]" pageId="12" pageNumber="11">R. W. Poole</collectorName>
(
<collectionCode id="C2D7AE1A8E01AF534710FE05FD29FDA3" box="[594,684,511,535]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="11" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECDC65548E01AF5345AAFDE5FBF6FD2D" pageId="12" pageNumber="11" type="distribution">
<paragraph id="A47936DF8E01AF5345AAFDE5FBF6FD2D" blockId="12.[192,1441,217,666]" pageId="12" pageNumber="11">
<emphasis id="96B2EACD8E01AF5345AAFDE5FDBDFD83" bold="true" box="[232,568,543,568]" pageId="12" pageNumber="11">Distribution and biology.</emphasis>
We record this species throughout much of
<collectingCountry id="DCD1764F8E01AF534175FDDAFB06FD8C" box="[1079,1155,544,568]" name="Brazil" pageId="12" pageNumber="11">Brazil</collectingCountry>
, south to
<collectingCountry id="DCD1764F8E01AF5341B0FDDAFAEDFD8C" box="[1266,1384,544,568]" name="Paraguay" pageId="12" pageNumber="11">Paraguay</collectingCountry>
, and northwest to
<collectingCountry id="DCD1764F8E01AF534422FDBAFE5AFDEC" box="[352,479,576,600]" name="Venezuela" pageId="12" pageNumber="11">Venezuela</collectingCountry>
, from approximately
<quantity id="633E9B3A8E01AF5347A3FDBAFC09FDEC" box="[737,908,576,600]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="12" pageNumber="11" unit="m" value="550.0" valueMax="1000.0" valueMin="100.0">100 to 1000 m</quantity>
elevation. A specimen in the NHMUK from
<collectingCountry id="DCD1764F8E01AF534582FD9BFF7FFDCC" box="[192,250,609,632]" name="Peru" pageId="12" pageNumber="11">Peru</collectingCountry>
may or may not be conspecific with
<taxonomicName id="63C64D5C8E01AF5347EDFD9BFCDFFDCC" box="[687,858,608,632]" class="Insecta" family="Tortricidae" genus="Cosmorrhyncha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="ocelliferana">
<emphasis id="96B2EACD8E01AF5347EDFD9BFCDFFDCC" box="[687,858,608,632]" italics="true" pageId="12" pageNumber="11">C. ocelliferana</emphasis>
</taxonomicName>
. Nothing is known of the early stages. Captures throughout the year suggest that the species may have continuous generations.
</paragraph>
</subSubSection>
</treatment>
</document>