146 lines
12 KiB
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146 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.797.29510" ID-GBIF-Dataset="17a988bc-106e-4cd5-af88-8a606353b34f" ID-PMC="PMC6255853" ID-Pensoft-Pub="1313-2970-797-19" ID-PubMed="30505161" ID-ZBK="A5403588F43D40FF81EE2009E49823B4" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-797-19" ModsDocOrigin="ZooKeys 797" ModsDocTitle="A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1542677994046" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai & Panha, Somsak" docDate="2018" docId="095D147E436CDE04BA4757B82F9B36C8" docLanguage="en" docName="ZooKeys 797: 19-69" docOrigin="ZooKeys 797" docSource="http://dx.doi.org/10.3897/zookeys.797.29510" docTitle="Spinaxytes palmata Srisonchai, Enghoff & Panha, sp. n." docType="treatment" docUuid="2A4B8447-3443-44D7-B463-0A6A2AB840C9" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="7C43FFACB225AA2193482143FFF2FF98" masterDocTitle="A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="69" masterPageNumber="19" originalUpdateDomain="Boston" originalUpdateTime="1542677994046" originalUpdateUser="pensoft" pageNumber="19" updateTime="1668166441925" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>797</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.797.29510</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.797.29510</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-797-19</mods:identifier>
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<mods:identifier type="ZBK">A5403588F43D40FF81EE2009E49823B4</mods:identifier>
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<mods:identifier type="ZooBank">A5403588F43D40FF81EE2009E49823B4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="149736186" LSID="urn:lsid:zoobank.org:act:2A4B8447-3443-44D7-B463-0A6A2AB840C9" httpUri="http://treatment.plazi.org/id/095D147E436CDE04BA4757B82F9B36C8" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="http://zoobank.org/2A4B8447-3443-44D7-B463-0A6A2AB840C9" authority="Srisonchai, Enghoff & Panha" class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes palmata" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="palmata">Spinaxytes palmata Srisonchai, Enghoff & Panha</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="19">sp. n.</taxonomicNameLabel>
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Figs 2; 4N, O; 5F; 21, 22, 23
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Holotype. ♂, THAILAND, Surat Thani Province, Phanom District, Wat Tham Wararam,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.885278">8°53'07"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="98.66694">98°40'01"E</geoCoordinate>
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, ca. 51 m a.s.l., 6 Aug. 2016, ASRU members leg. (CUMZ-pxDGT00215). Paratypes. 23 ♂♂, 22 ♀♀, same data as for holotype (CUMZ-pxDGT00216); 1 ♂, 1 ♀, same data as for holotype (ZMUC00040254); 1 ♂, 1 ♀, same data as for holotype (ZMUM); 1 ♂, 1 ♀, same data as for holotype (NHMW9425); 1 ♂, 1 ♀, same data as for holotype (NHMUK). Further specimens, not paratypes. THAILAND: 1 ♂, 2 ♀♀, Surat Thani Province, Phanom District, Wat Tham Wararam,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.885278">8°53'07"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="98.66694">98°40'01"E</geoCoordinate>
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, ca. 51 m a.s.l., 5 Aug. 2014, ASRU members leg. (CUMZ).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="etymology">
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<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="19">The species name is a Latin adjective, referring to the tip of lamina medialis which is somewhat hand-shaped.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Male femora without modification. Similar in this respect to
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<taxonomicName lsidName="S. efefi" pageId="0" pageNumber="19" rank="species" species="efefi">S. efefi</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="S. hasta" pageId="0" pageNumber="19" rank="species" species="hasta">S. hasta</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="S. sutchariti" pageId="0" pageNumber="19" rank="species" species="sutchariti">S. sutchariti</taxonomicName>
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sp. n. and
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<taxonomicName lsidName="S. tortioverpa" pageId="0" pageNumber="19" rank="species" species="tortioverpa">S. tortioverpa</taxonomicName>
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sp. n., but differs from them by having: anterior part of sternal lobe between male coxae 4 bifurcate, fork-like; tip of lamina medialis expanded, hand-shaped.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">SIZE. Length 26-30 mm (male), 27-32 mm (female); width of midbody metazona 1.9-2.2 mm (male), 2.0-2.4 mm (female). Width of collum = 2 = 3 = 4 <head <5-16, thereafter body gradually tapering towards telson.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Colour (Figure 21
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Specimens in life with body black; paraterga orange; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona, metaterga (except white spines in posterior row) and surface below paraterga black; sterna and legs brown; epiproct pale brown; a few basal podomeres whitish brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">Antennae (Figure 22M). Reaching to body ring 8 (male) and 6 or 7 (female) when stretched dorsally.</paragraph>
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<paragraph pageId="0" pageNumber="19">Collum (Figure 22A). With three transverse rows of setiferous tubercles/cones, 4+4 in anterior row, 1(0)+1(0) in intermediate row and 2(1)+2(1) in posterior row; with one conspicuous setiferous notch at lateral margin; paraterga spiniform, long, tip sharp, elevated at ca. 15°-20° (male) 10°-15° (female), directed almost laterad.</paragraph>
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<paragraph pageId="0" pageNumber="19">Tegument. Very shining; collum coarsely microgranulate; metaterga and surface below paraterga smooth.</paragraph>
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<paragraph pageId="0" pageNumber="19">Metaterga (Figure 22A, C, E). With two transverse rows of setiferous cones and setiferous spines; metaterga 2-19 with 2+2 cones in anterior row and 2+2 spines in posterior row; lateral cones/spines of posterior row bigger and longer than mesal ones, gradually reduced in size and length on the following rings.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Paraterga (Figure 22
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, H). Very long; directed dorsolaterad on body rings 2-17, elevated at ca. 50°-60° (male) 45°-60° (female), directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">Telson (Figure 22E, H, I). Epiproct quite short; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subtriangular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles.</paragraph>
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<paragraph pageId="0" pageNumber="19">Sterna (Figs 4N; 22F, G). Sternal lobe between male coxae 4 with two parts; anterior part bifurcate, tuning-fork-like, long, tips sharp, in situ directed ventroanteriad; posterior margin of anterior part bearing 1 pore; posterior part swollen, short.</paragraph>
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<paragraph pageId="0" pageNumber="19">Legs (Figs 4O, 22J). Male femora without modification.</paragraph>
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<paragraph pageId="0" pageNumber="19">Gonopods (Figs 5F, 23). Coxa shorter than femur. Prefemoral part ca. half as long as femur. Femur not enlarged distally. Postfemoral part broad. Mesal sulcus and lateral sulcus wide. Solenophore a bit bigger than postfemoral part: lamina lateralis small, oval, tip round: lamina medialis long; basally enlarged and slightly attenuated near the tip; tip fringed, hand-shaped; tip curving down, in situ resting close to solenomere. Solenomere curved and twisted, compressed in transverse section, tip directed posteriad.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="distribution">
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<paragraph pageId="0" pageNumber="19">Distribution and habitat</paragraph>
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<paragraph pageId="0" pageNumber="19">
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(Figure 21D).
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<taxonomicName lsidName="S. palmata" pageId="0" pageNumber="19" rank="species" species="palmata">S. palmata</taxonomicName>
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sp. n. is known only from the type locality. We regard this species as endemic for the Thai fauna. The new species can be found in syntopy with
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes corythosaurus" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="corythosaurus">Desmoxytes corythosaurus</taxonomicName>
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Srisonchai, Enghoff & Panha, 2018, crawling on humid rock walls.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="remarks">
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<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="19">There are variations in the hypoproct: subtrapeziform in some specimens, subtriangular in the others; caudal margin in some individuals round, angular in the others.</paragraph>
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<caption pageId="0" pageNumber="19">
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<paragraph pageId="0" pageNumber="19">
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Figure 13.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes hasta" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hasta">Spinaxytes hasta</taxonomicName>
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sp. n., ♂ paratype, CUMZ-pxDGT00210 A, B anterior body part C, D body rings 8-10 E, H, I posteriormost body rings and telson F, G sternal lobe between coxae 4 J legs 3-7 K legs 1-3 L gonopods M left antenna N sculpture of body ring 10.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="19">
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<paragraph pageId="0" pageNumber="19">
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Figure 14.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes hasta" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="hasta">Spinaxytes hasta</taxonomicName>
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sp. n., paratype, CUMZ-pxDGT00210 - right gonopod A lateral view B mesal view C ventral view D mesodorsal view E dorsal view (arrowhead points to lobe on lamina medialis) F laterodorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |