treatments-xml/data/08/69/49/0869494AFFDFFFFCFF3EFB75FAF3FD41.xml

<document id="442AC22A030778CF2A6FB4AE5C368F00" ID-CLB-Dataset="20622" ID-DOI="10.11646/zootaxa.5099.5.4" ID-GBIF-Dataset="528fb8bb-8347-4f38-8278-80f0f40a4e01" ID-ISSN="1175-5326" ID-Zenodo-Dep="6110242" ID-ZooBank="9F61D437-C30A-4C26-9453-E8EB76DAC651" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644998194949" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Miller, Allison K. &amp; Malay, Maria Celia D." docDate="2022" docId="0869494AFFDFFFFCFF3EFB75FAF3FD41" docLanguage="en" docName="zootaxa.5099.5.4.pdf" docOrigin="Zootaxa 5099 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nematopagurus marianicus Komai &amp; Miller &amp; Malay 2022, n. sp." docType="treatment" docUuid="04CC8B85-19FD-43F4-8BD8-E4FB6E58FC6E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="576" masterDocId="F4503132FFD8FFF1FFA9FFBAFF9CFFC4" masterDocTitle="Three new species of pagurid hermit crabs (Decapoda: Anomura: Paguroidea) from the Northern Mariana Islands, Micronesia" masterLastPageNumber="585" masterPageNumber="563" pageNumber="570" updateTime="1699312104795" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4A4DE15E67EF20C5653FC8D181BE9BAF">Three new species of pagurid hermit crabs (Decapoda: Anomura: Paguroidea) from the Northern Mariana Islands, Micronesia</mods:title>
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<mods:namePart id="03831BBC7A1DADC8E63573614483291F">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="2B09489780DEC93A501582CD5E52671E">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba 260 - 8682, Japan.</mods:affiliation>
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<mods:namePart id="EA86D0C8AC50CD28A295C1916258D96F">Miller, Allison K.</mods:namePart>
<mods:affiliation id="B3A1A39CB28F2285C24165351B204C2E">Anatomy Department, University of Otago, Dunedin, Otago, New Zealand.</mods:affiliation>
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<mods:namePart id="A5ADD530B7E01215A1238A7EDF690B73">Malay, Maria Celia D.</mods:namePart>
<mods:nameIdentifier id="61DA67539CB9CA45878CBA93BE2D360C" type="ORCID">0000-0003-0219-4788</mods:nameIdentifier>
<mods:affiliation id="A33C59FDA1706DA6F1B87F61973B5205">Marine Biodiversity and Evolution Laboratory, Division of Biological Sciences, University of the Philippines Visayas, Miag-ao, Iloilo, Philippines. mdmalay @ up. edu. ph; https: // orcid. org / 0000 - 0003 - 0219 - 4788</mods:affiliation>
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<emphasis id="B2B4244EFFDFFFF6FF3EFB75FE75FB2C" bold="true" box="[151,489,1230,1257]" italics="true" pageId="7" pageNumber="570">Nematopagurus marianicus</emphasis>
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(
<figureCitation id="18FBE4D9FFDFFFF6FF36FB48FE97FAC9" box="[159,267,1266,1293]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="9.[151,250,1876,1900]" captionStartId-1="10.[151,250,1957,1981]" captionStartId-2="11.[151,250,1454,1478]" captionStartId-3="12.[151,250,1567,1591]" captionTargetBox-0="[164,1421,388,1836]" captionTargetBox-1="[165,1421,496,1921]" captionTargetBox-2="[164,1416,193,1427]" captionTargetBox-3="[161,1423,195,1529]" captionTargetId-0="figure-128@9.[151,1436,381,1853]" captionTargetId-1="figure-139@10.[151,1436,489,1934]" captionTargetId-2="figure-19@11.[151,1436,181,1430]" captionTargetId-3="figure-19@12.[151,1436,181,1544]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." captionText-1="FIGURE 6. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm). A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted)." captionText-2="FIGURE 7. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view." captionText-3="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi-0="http://doi.org/10.5281/zenodo.6110254" figureDoi-1="http://doi.org/10.5281/zenodo.6110256" figureDoi-2="http://doi.org/10.5281/zenodo.6110258" figureDoi-3="http://doi.org/10.5281/zenodo.6110260" httpUri-0="https://zenodo.org/record/6110254/files/figure.png" httpUri-1="https://zenodo.org/record/6110256/files/figure.png" httpUri-2="https://zenodo.org/record/6110258/files/figure.png" httpUri-3="https://zenodo.org/record/6110260/files/figure.png" pageId="7" pageNumber="570">Figs. 5–8</figureCitation>
, 
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)
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<emphasis id="B2B4244EFFDFFFF6FF3EFA80FE1DFA90" bold="true" box="[151,385,1338,1364]" pageId="7" pageNumber="570">Material examined.</emphasis>
<materialsCitation id="30A8F201FFDFFFF6FE21FA80FE24FABD" ID-GBIF-Occurrence="3467114304" accessionNumber="C14523" collectingDate="2015-06-17" collectorName="Agrihan &amp; No. S" pageId="7" pageNumber="570" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F7B46FEFFDFFFF6FE21FA80FE68FA90" box="[392,500,1338,1364]" pageId="7" pageNumber="570" type="holotype">Holotype</typeStatus>
. SIO-BIC C14523, male (sl 
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, no depth record, 
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, 
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, field 
<collectorName id="2D359D8AFFDFFFF6FEBFFAE5FEC8FABD" box="[278,340,1374,1401]" pageId="7" pageNumber="570">No. S</collectorName>
25722.4.
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<typeStatus id="5F7B46FEFFDFFFF6FF6EFA39FEA4FA59" box="[199,312,1411,1437]" pageId="7" pageNumber="570" type="paratype">Paratypes</typeStatus>
. SIO-BIC C14238, 
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(sl 
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), same data as holotype, field 
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SIO-BIC C14512, 
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(sl 
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), station AKM-PAG-04, 
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, 
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, 
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, field 
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25713.1;
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SIO-BIC C14513, 
<specimenCount id="96C633D5FFDFFFF6FDB9FA70FDF9FA20" box="[528,613,1482,1508]" pageId="7" pageNumber="570" type="male">2 males</specimenCount>
(sl 2.3, 
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; including DNA voucher), 2 ovigerous females (sl 2.1, 
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SIO-BIC C14517, 1 ovigerous female (sl 
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, 
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, 
<quantity id="473855B9FFDFFFF6FE1FF9A8FD81F9E9" box="[438,541,1554,1581]" metricMagnitude="1" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="7" pageNumber="570" unit="m" value="42.0" valueMax="44.0" valueMin="40.0">
<elevation id="0BED1F6FFFDFFFF6FE1FF9A8FD81F9E9" box="[438,541,1554,1581]" metricMagnitude="1" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="7" pageNumber="570" unit="m" value="42.0" valueMax="44.0" valueMin="40.0">40–44 m</elevation>
</quantity>
, 
<date id="F47EDE9CFFDFFFF6FD83F9A8FD5AF9E9" box="[554,710,1554,1581]" pageId="7" pageNumber="570" value="2015-06-23">
<collectingDate id="E43A2774FFDFFFF6FD83F9A8FD5AF9E9" box="[554,710,1554,1581]" pageId="7" pageNumber="570" value="2015-06-23">23 June 2015</collectingDate>
</date>
, Buoy 
<collectingCountry id="F8D7B8CCFFDFFFF6FCB1F9A8FCC0F9E8" box="[792,860,1554,1580]" name="Guam" pageId="7" pageNumber="570">Guam</collectingCountry>
(a 
<collectingMethod id="5981804BFFDFFFF6FC2AF9A8FBA5F9E9" box="[899,1081,1554,1581]" pageId="7" pageNumber="570">kind of ring net</collectingMethod>
), field 
<collectorName id="2D359D8AFFDFFFF6FB23F9A9FB56F9E9" box="[1162,1226,1554,1581]" pageId="7" pageNumber="570">No. S</collectorName>
25750.4;
</materialsCitation>
<materialsCitation id="30A8F201FFDFFFF6FA9EF9A8FE6DF9B1" ID-GBIF-Occurrence="3467114308" accessionNumber="C14238, C14512, C14513, C14517, C14518, C14519, C14520, C14522, S25717, C14524" collectingDate="2015-06-17" collectingDateMax="2015-06-23" collectingDateMin="2015-06-17" country="Guam" elevation="74" latitude="18.01" location="Pagan" longLatPrecision="766" longitude="145.04" pageId="7" pageNumber="570" specimenCount="11" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="9" typeStatus="paratype">
SIO-BIC C14518, 
<specimenCount id="96C633D5FFDFFFF6FEA9F98CFED1F994" box="[256,333,1590,1616]" pageId="7" pageNumber="570" type="male">1 male</specimenCount>
(sl 
<quantity id="473855B9FFDFFFF6FED5F98CFE45F995" box="[380,473,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="570" unit="mm" value="2.8">2.8 mm</quantity>
), Guguan, station AKM-GUG-03, 
<geoCoordinate id="E5F49E9BFFDFFFF6FCCFF98CFC7FF994" box="[870,995,1590,1616]" degrees="17.6093" direction="north" orientation="latitude" pageId="7" pageNumber="570" precision="5" value="17.6093">17.6093°N</geoCoordinate>
, 
<geoCoordinate id="E5F49E9BFFDFFFF6FC58F98CFBE4F995" box="[1009,1144,1590,1617]" degrees="145.8101" direction="east" orientation="longitude" pageId="7" pageNumber="570" precision="5" value="145.8101">145.8101°E</geoCoordinate>
, 
<quantity id="473855B9FFDFFFF6FB2FF98CFB71F995" box="[1158,1261,1590,1617]" metricMagnitude="1" metricUnit="m" metricValue="6.6" metricValueMax="8.4" metricValueMin="4.8" pageId="7" pageNumber="570" unit="m" value="66.0" valueMax="84.0" valueMin="48.0">
<elevation id="0BED1F6FFFDFFFF6FB2FF98CFB71F995" box="[1158,1261,1590,1617]" metricMagnitude="1" metricUnit="m" metricValue="6.6" metricValueMax="8.4" metricValueMin="4.8" pageId="7" pageNumber="570" unit="m" value="66.0" valueMax="84.0" valueMin="48.0">48–84 m</elevation>
</quantity>
, 
<date id="F47EDE9CFFDFFFF6FB52F98CFA04F995" box="[1275,1432,1590,1617]" pageId="7" pageNumber="570" value="2015-06-22">
<collectingDate id="E43A2774FFDFFFF6FB52F98CFA04F995" box="[1275,1432,1590,1617]" pageId="7" pageNumber="570" value="2015-06-22">22 June 2015</collectingDate>
</date>
, 
<collectingMethod id="5981804BFFDFFFF6FF3EF9E0FE94F9B1" box="[151,264,1626,1653]" pageId="7" pageNumber="570">crab traps</collectingMethod>
, field 
<collectorName id="2D359D8AFFDFFFF6FEE7F9E1FE10F9B1" box="[334,396,1626,1653]" pageId="7" pageNumber="570">No. S</collectorName>
25745.2;
</materialsCitation>
<materialsCitation id="30A8F201FFDFFFF6FE51F9E0FEE9F95D" ID-GBIF-Occurrence="3467114305" accessionNumber="C14238, C14512, C14513, C14517, C14518, C14519, C14520, C14522, S25717, C14524" collectingDate="2015-06-17" collectingDateMax="2015-06-23" collectingDateMin="2015-06-17" country="Guam" elevation="74" latitude="18.01" location="Pagan" longLatPrecision="766" longitude="145.04" pageId="7" pageNumber="570" specimenCount="11" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="9" typeStatus="paratype">
SIO-BIC C14519, 1 ovigerous female (sl 
<quantity id="473855B9FFDFFFF6FC64F9E0FBB4F9B1" box="[973,1064,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="570" unit="mm" value="2.1">2.1 mm</quantity>
), same data as SIO-BIC C14518, field 
<collectorName id="2D359D8AFFDFFFF6FF7BF9C5FE8DF95D" box="[210,273,1662,1689]" pageId="7" pageNumber="570">No. S</collectorName>
25745.3;
</materialsCitation>
<materialsCitation id="30A8F201FFDFFFF6FED4F9C4FE40F979" ID-GBIF-Occurrence="3467114310" accessionNumber="C14238, C14512, C14513, C14517, C14518, C14519, C14520, C14522, S25717, C14524" collectingDate="2015-06-17" collectingDateMax="2015-06-23" collectingDateMin="2015-06-17" country="Guam" elevation="74" latitude="18.01" location="Pagan" longLatPrecision="766" longitude="145.04" pageId="7" pageNumber="570" specimenCount="11" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="9" typeStatus="paratype">
SIO-BIC C14520, 
<specimenCount id="96C633D5FFDFFFF6FDF8F9C4FD34F95C" box="[593,680,1662,1688]" pageId="7" pageNumber="570" type="male">3 males</specimenCount>
(sl 2.2, 2.5, 
<quantity id="473855B9FFDFFFF6FC9EF9C4FC0EF95D" box="[823,914,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="570" unit="mm" value="2.8">2.8 mm</quantity>
), 
<specimenCount id="96C633D5FFDFFFF6FC0CF9C4FB9BF95C" box="[933,1031,1662,1688]" pageId="7" pageNumber="570" type="female">1 female</specimenCount>
(sl 
<quantity id="473855B9FFDFFFF6FB9BF9C4FB11F95D" box="[1074,1165,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="570" unit="mm" value="1.9">1.9 mm</quantity>
), same data as SIO-BIC C14518, field 
<collectorName id="2D359D8AFFDFFFF6FE90F919FEE4F979" box="[313,376,1698,1725]" pageId="7" pageNumber="570">No. S</collectorName>
25745.4;
</materialsCitation>
<materialsCitation id="30A8F201FFDFFFF6FE4AF918FD72F925" ID-GBIF-Occurrence="3467114302" accessionNumber="C14238, C14512, C14513, C14517, C14518, C14519, C14520, C14522, S25717, C14524" collectingDate="2015-06-17" collectingDateMax="2015-06-23" collectingDateMin="2015-06-17" country="Guam" elevation="74" latitude="18.01" location="Pagan" longLatPrecision="766" longitude="145.04" pageId="7" pageNumber="570" specimenCount="11" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="9" typeStatus="paratype">
SIO-BIC C14522, 
<specimenCount id="96C633D5FFDFFFF6FD1EF918FCABF978" box="[695,823,1698,1724]" pageId="7" pageNumber="570" type="immature">1 immature</specimenCount>
male (sl 
<quantity id="473855B9FFDFFFF6FC09F918FC67F979" box="[928,1019,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="570" unit="mm" value="1.7">1.7 mm</quantity>
), 
<collectorName id="2D359D8AFFDFFFF6FBA7F919FBC9F979" box="[1038,1109,1699,1725]" pageId="7" pageNumber="570">Pagan</collectorName>
, 
<geoCoordinate id="E5F49E9BFFDFFFF6FBC8F918FB5EF978" box="[1121,1218,1698,1724]" degrees="18.01" direction="north" orientation="latitude" pageId="7" pageNumber="570" precision="555" value="18.01">18.01°N</geoCoordinate>
, 
<geoCoordinate id="E5F49E9BFFDFFFF6FB67F918FAA6F979" box="[1230,1338,1698,1725]" degrees="145.04" direction="east" orientation="longitude" pageId="7" pageNumber="570" precision="555" value="145.04">145.04°E</geoCoordinate>
, 
<date id="F47EDE9CFFDFFFF6FAEFF918FF4EF925" pageId="7" pageNumber="570" value="2015-06-20">
<collectingDate id="E43A2774FFDFFFF6FAEFF918FF4EF925" pageId="7" pageNumber="570" value="2015-06-20">20 June 2015</collectingDate>
</date>
, ring nets for trapping crabs, field 
<collectorName id="2D359D8AFFDFFFF6FDF6F97DFD15F925" box="[607,649,1735,1761]" pageId="7" pageNumber="570">No.</collectorName>
<accessionNumber id="9F9365BFFFDFFFF6FD38F97CFD76F925" box="[657,746,1734,1761]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S25717" pageId="7" pageNumber="570" type="EnaNcbi">S25717</accessionNumber>
;
</materialsCitation>
<materialsCitation id="30A8F201FFDFFFF6FD5EF97CFD75F8C1" ID-GBIF-Occurrence="3467114307" accessionNumber="C14238, C14512, C14513, C14517, C14518, C14519, C14520, C14522, S25717, C14524" collectingDate="2015-06-17" collectingDateMax="2015-06-23" collectingDateMin="2015-06-17" country="Guam" elevation="74" latitude="18.01" location="Pagan" longLatPrecision="766" longitude="145.04" pageId="7" pageNumber="570" specimenCount="11" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="9" typeStatus="paratype">
SIO-BIC C14524, 
<specimenCount id="96C633D5FFDFFFF6FC65F97CFB85F924" box="[972,1049,1734,1760]" pageId="7" pageNumber="570" type="male">1 male</specimenCount>
(sl 
<quantity id="473855B9FFDFFFF6FBEFF97CFB3EF925" box="[1094,1186,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="570" unit="mm" value="2.3">2.3 mm</quantity>
), 
<collectorName id="2D359D8AFFDFFFF6FB1FF97DFB61F925" box="[1206,1277,1735,1761]" pageId="7" pageNumber="570">Pagan</collectorName>
, depth not recorded, 
<date id="F47EDE9CFFDFFFF6FF58F950FE16F8C1" box="[241,394,1770,1797]" pageId="7" pageNumber="570" value="2015-06-17">
<collectingDate id="E43A2774FFDFFFF6FF58F950FE16F8C1" box="[241,394,1770,1797]" pageId="7" pageNumber="570" value="2015-06-17">17 June 2015</collectingDate>
</date>
, 
<collectingMethod id="5981804BFFDFFFF6FE3CF950FD99F8C1" box="[405,517,1770,1797]" pageId="7" pageNumber="570">crab traps</collectingMethod>
, field 
<collectorName id="2D359D8AFFDFFFF6FDE2F951FD15F8C1" box="[587,649,1770,1797]" pageId="7" pageNumber="570">No. S</collectorName>
25722.3
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8DAABD7FFDFFFFDFF6EF8B4FE60F90C" lastPageId="12" lastPageNumber="575" pageId="7" pageNumber="570" type="description">
<paragraph id="807FF85CFFDFFFF6FF6EF8B4FDC1F8EC" blockId="7.[151,1437,1338,2013]" box="[199,605,1806,1832]" pageId="7" pageNumber="570">
<emphasis id="B2B4244EFFDFFFF6FF6EF8B4FEC5F8EC" bold="true" box="[199,345,1806,1832]" pageId="7" pageNumber="570">Description.</emphasis>
Gills biserial, 11 pairs.
</paragraph>
<paragraph id="807FF85CFFDFFFF6FF6EF888FD3FF818" blockId="7.[151,1437,1338,2013]" pageId="7" pageNumber="570">
Shield (
<figureCitation id="18FBE4D9FFDFFFF6FEB6F888FEEBF889" box="[287,375,1842,1869]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="7" pageNumber="570">Fig. 5A</figureCitation>
) approximately as long as wide, dorsal surface with longitudinal row of setae posterior to anterolateral projection on either side; anterior margin between rostral lobe and lateral projections weakly interrupted. Rostrum as broadly triangular or rounded rostral lobe, not quite reaching to level of lateral projections, unarmed, but with tuft of short setae on either side of midline. Lateral projections obtusely triangular, with or without submarginal spine. Lateral carapace lobe moderately broad.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFF2DFBA8FED9" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Ocular peduncles (
<figureCitation id="18FBE4D9FFD0FFF9FE37FF2DFE64FF75" box="[414,504,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="8" pageNumber="571">Fig. 5A</figureCitation>
) relatively stout, 0.8 length of shield, subcylindrical; dorsal surface tufts of short setae at corneal indentation; mesial surface with small, low setose protuberance proximal to corneal base; cornea substantially dilated, corneal width about half of peduncular length. Ocular acicles narrowly triangular, terminally bunt and with submarginal spine, separated widely. Interocular lobe fully exposed.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFE9DFB7BFE4D" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Antennular peduncles (
<figureCitation id="18FBE4D9FFD0FFF9FE65FE9DFDB8FE85" box="[460,548,294,321]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="8" pageNumber="571">Fig. 5A</figureCitation>
) slightly overreaching distal corneal margin by tip of ultimate article. Ultimate article without tuft of setae at distolateral margin, dorsal surface almost glabrous. Penultimate article unarmed. Basal article without spine laterally on statocyst lobe, ventrodistal angle not particularly produced.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFE29FEB7FD6D" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Antennal peduncles (
<figureCitation id="18FBE4D9FFD0FFF9FE13FE29FD88FE69" box="[442,532,402,429]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="8" pageNumber="571">Fig. 5A</figureCitation>
) falling slightly short of distal corneal margin. Articles 3–5 unarmed but each with few tufts or individual stiff setae. Article 2 with dorsolateral distal angle produced beyond proximal margin of article 4, terminating in small spine and sometimes with additional subterminal spine, mesial and/or lateral margins with few tufts of setae; dorsomesial distal angle with small spine. Article 1 with ventrolateral margin prominently produced, unarmed. Antennal acicles reaching to or beyond midlength of peduncular article 5, terminating in small spine; dorsomesial margin with several tufts of stiff setae proximally and row of stiff setae in distal half. Antennal flagella long, far overreaching outstretched chelipeds; each article with one or two minute bristle-like setae, at least in distal half.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFD09FCBEFCD1" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Maxilliped 3 (
<figureCitation id="18FBE4D9FFD0FFF9FEC1FD09FE7DFD09" box="[360,481,690,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="8" pageNumber="571">Fig. 5B, C</figureCitation>
) endopod moderately slender. Carpus and merus unarmed. Ischium with well-developed crista dentata consisting of row of closely set, corneous-tipped teeth and with 1 well-developed accessory tooth located subdistally. Exopod reaching midlength of carpus.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFCA5FAE8FAB3" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Right cheliped (
<figureCitation id="18FBE4D9FFD0FFF9FED2FCA5FE69FCFE" box="[379,501,799,826]" captionStart="FIGURE 6" captionStartId="10.[151,250,1957,1981]" captionTargetBox="[165,1421,496,1921]" captionTargetId="figure-139@10.[151,1436,489,1934]" captionTargetPageId="10" captionText="FIGURE 6. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm). A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.6110256" httpUri="https://zenodo.org/record/6110256/files/figure.png" pageId="8" pageNumber="571">Fig. 6A–D</figureCitation>
) larger and slightly longer than left. Chela subovate in general outline, 2.2 times as long as wide. Dactylus 0.8 length of palm, bearing row of small spines on dorsomesial margin, not extending to tip; mesial face with several tufts of short to long simple setae and few spinules proximally. Palm subequal in length to carpus, with dorsomesial and dorsolateral marginal row of small spines, latter extending onto proximal half of fixed finger; dorsal surface with very weak median row of small spines, distinct as row only proximally, and with additional numerous small spines or spinules and short, distally curved setae, spines extending onto proximal half of fixed finger; mesial and lateral faces each with sparse tufts of simple setae, longer and more numerous on mesial and ventral faces; occlusal margin of fixed finger with row of 5 obtuse calcareous teeth and minute, fused corneous teeth in distal 0.2, terminating in small corneous claw. Carpus slightly widened distally, 1.8 times as long as distal width, with row of moderately large spines on dorsomesial margin and row of smaller spines on dorsal surface laterally, dorsodistal margin with 1 small spine; mesial and lateral faces each with numerous tufts of long simple setae, partially obscuring faces; ventrolateral distal margin armed with 1 small spine, ventromesial distal margin unarmed; gently convex ventral surface with scattered long setae. Merus dorsal surface with subdistal transverse ridge but otherwise smooth, almost glabrous; dorsodistal margin with 1 small spine mesially, obscured by row of moderately long setae; lateral surface with few, short transverse rows of minute setae, ventrolateral margin with 2 widely spaced spines in distal 0.3; mesial surface almost glabrous except for few tufts of short setae, ventromesial margin with 3 or 4 spines accompanied by few long setae; ventral surface with short, setose transverse ridges. Ischium unarmed.
</paragraph>
<paragraph id="807FF85CFFD0FFF9FF6EFA3AFD22F883" blockId="8.[151,1437,150,2043]" pageId="8" pageNumber="571">
Left cheliped (
<figureCitation id="18FBE4D9FFD0FFF9FEC2FA3AFE78FA5F" box="[363,484,1408,1435]" captionStart="FIGURE 7" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[164,1416,193,1427]" captionTargetId="figure-19@11.[151,1436,181,1430]" captionTargetPageId="11" captionText="FIGURE 7. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6110258" httpUri="https://zenodo.org/record/6110258/files/figure.png" pageId="8" pageNumber="571">Fig. 7A–D</figureCitation>
) moderately slender, reaching level of base of dactylus of right cheliped. Chela elongate subovate in general outline, 2.6 times as long as wide. Dactylus with row of small spines on dorsomesial margin in proximal half. Dorsomesial and dorsolateral margins of palm each with row of spines, dorsal surface slightly elevated in midline and with row of moderately small spines, additional small spines and short setae scattered on dorsal surface, and median longitudinal row of spinules extending onto fixed finger; mesial and lateral faces each with few sparse tufts of short to long simple setae. Carpus with row of small spines on dorsomesial and dorsolateral margins, dorsodistal margin with 1 small median spine; lateral and mesial faces with sparse tufts or short vertical rows of long simple setae, partially obscuring faces (particularly on dorsal half of lateral face); ventral surface with few low protuberances and sparse tufts of long setae. Merus with transverse row of numerous setae on dorsodistal margin; dorsal surface glabrous; lateral face almost glabrous except for few tufts of setae adjacent to ventral margin, ventrolateral margin with 4 or 5 spines (distalmost one strongest); mesial face also almost glabrous with few tufts of short setae, ventromesial margin with 3 or 4 small spines accompanied with tufts of setae; ventral surface with short, setose transverse ridges. Ischium unarmed.
</paragraph>
<paragraph id="807FF85CFFD0FFF8FF6EF8EBFB49FEA1" blockId="8.[151,1437,150,2043]" lastBlockId="9.[151,1437,151,358]" lastPageId="9" lastPageNumber="572" pageId="8" pageNumber="571">
Pereopods 2 and 3 (
<figureCitation id="18FBE4D9FFD0FFF9FE0FF8EBFDBDF8AF" box="[422,545,1873,1899]" captionStart="FIGURE 8" captionStartId="12.[151,250,1567,1591]" captionTargetBox="[161,1423,195,1529]" captionTargetId="figure-19@12.[151,1436,181,1544]" captionTargetPageId="12" captionText="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="8" pageNumber="571">Fig. 8A–D</figureCitation>
) generally similar in armature and ornamentation, third slightly longer. Dactyli slender (length 13 times of width in second, 14 times of width in third), 1.3 times as long as propodus in second, 
<quantity id="473855B9FFD0FFF9FF3EF823FF4BF876" box="[151,215,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="8" pageNumber="571" unit="in" value="1.8">1.8 in</quantity>
third; dorsal and ventral margins each with row of sparse setae, which are easily broken off; lateral faces with few short setae; mesial faces each with dorsal (arranged in single or double row, 
<quantity id="473855B9FFD0FFF9FBBCF807FBE4F812" box="[1045,1144,1980,2007]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="5.08" metricValueMin="3.556" pageId="8" pageNumber="571" unit="in" value="17.0" valueMax="20.0" valueMin="14.0">14–20 in</quantity>
number) and ventral rows (arranged in single row, 
<quantity id="473855B9FFD0FFF9FE0DF85AFE64F83E" box="[420,504,2016,2042]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.54" metricValueMin="1.524" pageId="8" pageNumber="571" unit="in" value="8.0" valueMax="10.0" valueMin="6.0">6–10 in</quantity>
number) of moderately spaced small, slender corneous spines (
<figureCitation id="18FBE4D9FFD0FFF9FB11F85AFAADF83F" box="[1208,1329,2016,2043]" captionStart="FIGURE 8" captionStartId="12.[151,250,1567,1591]" captionTargetBox="[161,1423,195,1529]" captionTargetId="figure-19@12.[151,1436,181,1544]" captionTargetPageId="12" captionText="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="8" pageNumber="571">Fig. 8B, D</figureCitation>
). Propodi 1.4–1.8 lengths of carpi; rounded dorsal surfaces each with row of sparse short, stiff setae; lateral and mesial faces smooth, glabrous; ventral surfaces glabrous, with 1 or 2 small corneous spines at ventrodistal margin (
<figureCitation id="18FBE4D9FFD1FFF8FABDFF01FA0CFF11" box="[1300,1424,187,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1567,1591]" captionTargetBox="[161,1423,195,1529]" captionTargetId="figure-19@12.[151,1436,181,1544]" captionTargetPageId="12" captionText="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="9" pageNumber="572">Fig. 8B, D</figureCitation>
). Carpi slightly more than 0.5 length of meri; dorsal surfaces each with dorsodistal and dorsoproximal spines and row of stiff setae; lateral, mesial and ventral faces almost glabrous. Meri each with dorsal row of tufts of setae (fewer in third), ventrolateral distal angle with small spine (second) or unarmed; ventral margins each with few small spines (second) or unarmed (third). Ischia each with few tufts of setae on dorsal and/or ventral surfaces.
</paragraph>
<caption id="D4BFA8D4FFD1FFF8FF3EF8EEFBD4F81D" ID-DOI="http://doi.org/10.5281/zenodo.6110254" ID-Zenodo-Dep="6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="9" pageNumber="572" startId="9.[151,250,1876,1900]" targetBox="[164,1421,388,1836]" targetPageId="9">
<paragraph id="807FF85CFFD1FFF8FF3EF8EEFBD4F81D" blockId="9.[151,1437,1876,2009]" pageId="9" pageNumber="572">
<emphasis id="B2B4244EFFD1FFF8FF3EF8EEFE88F8A9" bold="true" box="[151,276,1876,1901]" pageId="9" pageNumber="572">FIGURE 5.</emphasis>
<taxonomicName id="47C083DFFFD1FFF8FEB2F8EFFDAAF8A8" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[283,566,1877,1900]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="572" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD1FFF8FEB2F8EFFDAAF8A8" box="[283,566,1877,1900]" italics="true" pageId="9" pageNumber="572">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD1FFF8FD94F8EFFDEBF8A9" bold="true" box="[573,631,1877,1901]" pageId="9" pageNumber="572">
<taxonomicNameLabel id="A9879935FFD1FFF8FD94F8EFFDEBF8A9" box="[573,631,1877,1901]" pageId="9" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view.
</paragraph>
</caption>
<paragraph id="807FF85CFFD2FFFBFF6EFF2DFA86FF11" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="573">
Pereopods 4 semichelate (
<figureCitation id="18FBE4D9FFD2FFFBFE44FF2DFDDBFF75" box="[493,583,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="10" pageNumber="573">Fig. 5D</figureCitation>
); dactylus gently curved, terminating in long corneous claw, ventral margin with row of minute corneous spinules; propodal rasps each consisting of single row of corneous scales.
</paragraph>
<paragraph id="807FF85CFFD2FFFBFF6EFF65FC82FED9" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="573">
Pereopods 5 weakly semichelate. 
<collectingRegion id="420436BEFFD2FFFBFDE1FF65FD1EFF3D" box="[584,642,223,249]" country="Maldives" name="Maale" pageId="10" pageNumber="573">Male</collectingRegion>
with elongate, soft sexual tube extruded from gonopore of right coxa (
<figureCitation id="18FBE4D9FFD2FFFBFF09FEB9FF6AFED9" box="[160,246,258,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="10" pageNumber="573">Fig. 5E</figureCitation>
); left with gonopore, but not protruded (
<figureCitation id="18FBE4D9FFD2FFFBFD13FEB9FC8CFED9" box="[698,784,258,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="10" pageNumber="573">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="807FF85CFFD2FFFBFF6EFE9DFD48FEA1" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="573">
Sternite 6 anterior lobe (
<figureCitation id="18FBE4D9FFD2FFFBFE70FE9DFDB0FE85" box="[473,556,294,321]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="10" pageNumber="573">Fig. 5F</figureCitation>
) subcircular, anterior margin with row of short to moderately long setae. Sternite 8 (
<figureCitation id="18FBE4D9FFD2FFFBFF4FFEF1FEA5FEA1" box="[230,313,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1760,1784]" captionTargetBox="[164,1424,192,1722]" captionTargetId="figure-19@3.[151,1436,181,1737]" captionTargetPageId="3" captionText="FIGURE 1. Catapagurus tenuilamina n. sp., holotype, SIO-BIC C14512, male (sl 2.7 mm).A, shield and cephalic appendages, dorsal view; B, distal two peduncular articles and flagella of left antennule, lateral view; C, left maxilliped 3, lateral view; D, same, basis and ischium, ventral view; E, coxae of chelipeds, ventral view; F, dactylus, propodus and carpus of left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, crossing dorsal surface of anterior part of pleon; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6110244" httpUri="https://zenodo.org/record/6110244/files/figure.png" pageId="10" pageNumber="573">Fig. 1E</figureCitation>
) with pair of circular protuberances.
</paragraph>
<paragraph id="807FF85CFFD2FFFBFF6EFED5FEE1FE15" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="573">
Pleon dextrally twisted. 
<collectingRegion id="420436BEFFD2FFFBFE72FED5FD89FE4D" box="[475,533,367,393]" country="Maldives" name="Maale" pageId="10" pageNumber="573">Male</collectingRegion>
with unpaired left pleopods 3–5, pleopods 3 and 4 greatly unequally biramous, pleopod 5 uniramous. Female with paired and modified first pleopods typical of genus; unpaired left pleopods 2–5 unequally biramous.
</paragraph>
<caption id="D4BFA8D4FFD2FFFBFF3EF81FFA9FF825" ID-DOI="http://doi.org/10.5281/zenodo.6110256" ID-Zenodo-Dep="6110256" httpUri="https://zenodo.org/record/6110256/files/figure.png" pageId="10" pageNumber="573" startId="10.[151,250,1957,1981]" targetBox="[165,1421,496,1921]" targetPageId="10">
<paragraph id="807FF85CFFD2FFFBFF3EF81FFA9FF825" blockId="10.[151,1436,1957,2018]" pageId="10" pageNumber="573">
<emphasis id="B2B4244EFFD2FFFBFF3EF81FFE8FF879" bold="true" box="[151,275,1957,1981]" pageId="10" pageNumber="573">FIGURE 6.</emphasis>
<taxonomicName id="47C083DFFFD2FFFBFEB3F81CFDA9F879" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[282,565,1958,1981]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="573" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD2FFFBFEB3F81CFDA9F879" box="[282,565,1958,1981]" italics="true" pageId="10" pageNumber="573">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD2FFFBFD95F81CFDE9F87A" bold="true" box="[572,629,1958,1982]" pageId="10" pageNumber="573">
<taxonomicNameLabel id="A9879935FFD2FFFBFD95F81CFDE9F87A" box="[572,629,1958,1982]" pageId="10" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, SIO-BIC C14523, male (sl 2.8 mm). A, right cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view (setae omitted).
</paragraph>
</caption>
<caption id="D4BFA8D4FFD3FFFAFF3EFA14FBDBFA2F" ID-DOI="http://doi.org/10.5281/zenodo.6110258" ID-Zenodo-Dep="6110258" httpUri="https://zenodo.org/record/6110258/files/figure.png" pageId="11" pageNumber="574" startId="11.[151,250,1454,1478]" targetBox="[164,1416,193,1427]" targetPageId="11">
<paragraph id="807FF85CFFD3FFFAFF3EFA14FBDBFA2F" blockId="11.[151,1437,1454,1515]" pageId="11" pageNumber="574">
<emphasis id="B2B4244EFFD3FFFAFF3EFA14FE8FFA03" bold="true" box="[151,275,1454,1479]" pageId="11" pageNumber="574">FIGURE 7.</emphasis>
<taxonomicName id="47C083DFFFD3FFFAFEB0FA15FDAEFA02" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[281,562,1455,1478]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="574" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD3FFFAFEB0FA15FDAEFA02" box="[281,562,1455,1478]" italics="true" pageId="11" pageNumber="574">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD3FFFAFD91FA15FDEDFA03" bold="true" box="[568,625,1455,1479]" pageId="11" pageNumber="574">
<taxonomicNameLabel id="A9879935FFD3FFFAFD91FA15FDEDFA03" box="[568,625,1455,1479]" pageId="11" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, SIO-BIC C14523, male (sl 2.8 mm). A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view.
</paragraph>
</caption>
<paragraph id="807FF85CFFD3FFFAFF6EF9A3FD26F9BF" blockId="11.[151,1437,1561,2020]" pageId="11" pageNumber="574">
Telson (
<figureCitation id="18FBE4D9FFD3FFFAFE8BF9A3FE0EF9F7" box="[290,402,1561,1588]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="11" pageNumber="574">Fig. 5G, I</figureCitation>
) divided into anterior and posterior portions by distinct lateral indentations; posterior lobes slightly asymmetrical, separated by moderately deep median cleft; terminal margins each with 2–6 principal spines and sometimes with additional spinules laterally.
</paragraph>
<paragraph id="807FF85CFFD3FFFAFF6EF93FFDFCF8B3" blockId="11.[151,1437,1561,2020]" pageId="11" pageNumber="574">
<emphasis id="B2B4244EFFD3FFFAFF6EF93FFE37F95B" bold="true" box="[199,427,1669,1695]" pageId="11" pageNumber="574">Colouration in life.</emphasis>
<figureCitation id="18FBE4D9FFD3FFFAFE1BF93FFD84F95B" box="[434,536,1669,1695]" captionStart="FIGURE 12" captionStartId="18.[151,250,1933,1957]" captionTargetBox="[176,1412,188,1910]" captionTargetId="figure-19@18.[176,1412,181,1910]" captionTargetPageId="18" captionText="FIGURE 12. A, Catapagurus tenuilamina n. sp., holotype, SIO-BIC C14512, male (sl 2.7 mm)., habitus in lateral view, showing living colouration; B, Nematopagurus marianicus n. sp., paratypes, 3 males (sl 2.2, 2.5, 2.8 mm), 1 female (sl 1.9 mm), SIO-BIC C14520, showing living colouration." figureDoi="http://doi.org/10.5281/zenodo.6110270" httpUri="https://zenodo.org/record/6110270/files/figure.png" pageId="11" pageNumber="574">Fig. 12B</figureCitation>
. Shield generally pink; posterior carapace darker reddish. Ocular peduncles red, with dark ring basally; cornea brown-gray. Antennular and antennal peduncles red, antennal flagellum uniformly reddish translucent. Right chela mottled with red and pink, distal parts of fingers whitish; carpus with red and pink longitudinal stripes; merus also with alternate red and pink longitudinal stripes. Left cheliped similarly coloured to right cheliped. Ambulatory dactyli pink to violet, subdistally with red ring, distally white; propodi, carpi and meri with alternate red and pink longitudinal stripes (4 red stripes visible on respective article). Pereopods 4 and 5 also reddish. Pleon reddish, semi-translucent.
</paragraph>
<paragraph id="807FF85CFFD3FFFAFF6EF83BFC3CF827" blockId="11.[151,1437,1561,2020]" pageId="11" pageNumber="574">
<emphasis id="B2B4244EFFD3FFFAFF6EF83BFEA3F85F" bold="true" box="[199,319,1921,1947]" pageId="11" pageNumber="574">Variation.</emphasis>
The setation on chelipeds in the small, immature male specimen (sl 
<quantity id="473855B9FFD3FFFAFBE5F83BFB3AF85F" box="[1100,1190,1921,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="574" unit="mm" value="1.7">1.7 mm</quantity>
; SIO-BIC C14522) is sparse compared with adult specimens, suggesting that the setation becomes denser with growth. Spination on the terminal margins of the telson is also variable, as figured (
<figureCitation id="18FBE4D9FFD3FFFAFC8BF873FC0EF827" box="[802,914,1993,2020]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="11" pageNumber="574">Fig. 5G, I</figureCitation>
).
</paragraph>
<caption id="D4BFA8D4FFD4FFFDFF3EF9A5FD52F944" ID-DOI="http://doi.org/10.5281/zenodo.6110260" ID-Zenodo-Dep="6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="12" pageNumber="575" startId="12.[151,250,1567,1591]" targetBox="[161,1423,195,1529]" targetPageId="12">
<paragraph id="807FF85CFFD4FFFDFF3EF9A5FD52F944" blockId="12.[151,1436,1567,1664]" pageId="12" pageNumber="575">
<emphasis id="B2B4244EFFD4FFFDFF3EF9A5FE8EF9FC" bold="true" box="[151,274,1567,1592]" pageId="12" pageNumber="575">FIGURE 8.</emphasis>
<taxonomicName id="47C083DFFFD4FFFDFEB1F99AFDADF9F3" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[280,561,1568,1591]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD4FFFDFEB1F99AFDADF9F3" box="[280,561,1568,1591]" italics="true" pageId="12" pageNumber="575">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD4FFFDFD9FF99AFDF2F9FC" bold="true" box="[566,622,1568,1592]" pageId="12" pageNumber="575">
<taxonomicNameLabel id="A9879935FFD4FFFDFD9FF99AFDF2F9FC" box="[566,622,1568,1592]" pageId="12" pageNumber="575" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, SIO-BIC C14523, male (sl 2.8 mm). A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view.
</paragraph>
</caption>
<paragraph id="807FF85CFFD4FFFDFF6EF914FE60F90C" blockId="12.[151,1437,1710,2025]" box="[199,508,1710,1736]" pageId="12" pageNumber="575">
<emphasis id="B2B4244EFFD4FFFDFF6EF914FEAEF90C" bold="true" box="[199,306,1710,1736]" pageId="12" pageNumber="575">Housing.</emphasis>
Gastropod shells.
</paragraph>
</subSubSection>
<subSubSection id="C8DAABD7FFD4FFFDFF6EF968FA82F928" box="[199,1310,1746,1772]" pageId="12" pageNumber="575" type="etymology">
<paragraph id="807FF85CFFD4FFFDFF6EF968FA82F928" blockId="12.[151,1437,1710,2025]" box="[199,1310,1746,1772]" pageId="12" pageNumber="575">
<emphasis id="B2B4244EFFD4FFFDFF6EF968FED2F928" bold="true" box="[199,334,1746,1772]" pageId="12" pageNumber="575">Etymology.</emphasis>
Derived from the locality where the 
<typeStatus id="5F7B46FEFFD4FFFDFD45F968FC80F928" box="[748,796,1746,1772]" pageId="12" pageNumber="575">type</typeStatus>
specimens of the new species were collected.
</paragraph>
</subSubSection>
<subSubSection id="C8DAABD7FFD4FFFDFF6EF94CFB77F8D4" box="[199,1259,1782,1808]" pageId="12" pageNumber="575" type="distribution">
<paragraph id="807FF85CFFD4FFFDFF6EF94CFB77F8D4" blockId="12.[151,1437,1710,2025]" box="[199,1259,1782,1808]" pageId="12" pageNumber="575">
<emphasis id="B2B4244EFFD4FFFDFF6EF94CFEFDF8D4" bold="true" box="[199,353,1782,1808]" pageId="12" pageNumber="575">Distribution.</emphasis>
Presently known only from the 
<collectingCountry id="F8D7B8CCFFD4FFFDFD60F94CFC70F8D4" box="[713,1004,1782,1808]" name="Northern Mariana Islands" pageId="12" pageNumber="575">Northern Mariana Islands</collectingCountry>
, at depths of 
<quantity id="473855B9FFD4FFFDFB28F94CFB7BF8D4" box="[1153,1255,1782,1808]" metricMagnitude="1" metricUnit="m" metricValue="6.45" metricValueMax="8.9" metricValueMin="4.0" pageId="12" pageNumber="575" unit="m" value="64.5" valueMax="89.0" valueMin="40.0">
40– 
<specimenCount id="96C633D5FFD4FFFDFB05F94CFB7BF8D4" box="[1196,1255,1782,1808]" pageId="12" pageNumber="575" type="generic">89 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C8DAABD7FFD4FFFCFF6EF8A0FAF3FD41" lastPageId="13" lastPageNumber="576" pageId="12" pageNumber="575" type="discussion">
<paragraph id="807FF85CFFD4FFFDFF6EF8A0FBC5F800" blockId="12.[151,1437,1710,2025]" pageId="12" pageNumber="575">
<emphasis id="B2B4244EFFD4FFFDFF6EF8A0FEA7F8F0" bold="true" box="[199,315,1818,1844]" pageId="12" pageNumber="575">Remarks.</emphasis>
<bibRefCitation id="E45185ADFFD4FFFDFEECF8A0FDADF8F1" author="McLaughlin, P. A." box="[325,561,1818,1845]" pageId="12" pageNumber="575" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin (2004b)</bibRefCitation>
reviewed species of 
<taxonomicName id="47C083DFFFD4FFFDFC8FF8A1FC45F8F0" authorityName="A. Milne-Edwards &amp; Bouvier" authorityYear="1892" box="[806,985,1819,1844]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B2B4244EFFD4FFFDFC8FF8A1FC45F8F0" box="[806,985,1819,1844]" italics="true" pageId="12" pageNumber="575">Nematopagurus</emphasis>
</taxonomicName>
, recognizing 26 species, including five new species. Since then two new species have been described in the genus (
<bibRefCitation id="E45185ADFFD4FFFDFC5FF884FAA5F89C" author="McLaughlin, P. A. &amp; Okuno, J." box="[1014,1337,1854,1880]" pageId="12" pageNumber="575" pagination="1997" refId="ref12360" refString="McLaughlin, P. A. &amp; Okuno, J. (2010) A new and distinctive Japanese species of Nematopagurus (Decapoda, Paguridae), and a new record for Nematopagurus alcocki McLaughlin, 1997. In: Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (Eds.), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs. Vol. 14. Brill, Leiden, pp. 473 - 482." type="book chapter" year="2010">McLaughlin &amp; Okuno 2010</bibRefCitation>
; 
<bibRefCitation id="E45185ADFFD4FFFDFAECF884FF65F8B8" author="Lima, D. &amp; Tavares, M. &amp; Mendonca, J. B. de Jr." pageId="12" pageNumber="575" pagination="1 - 63" refId="ref11833" refString="Lima, D., Tavares, M. &amp; Mendonca, J. B. de Jr. (2019) Paguroids (Decapoda: Anomura: Diogenidae and Paguridae) of the remote oceanic Archipelago Trindade and Martin Vaz, off southeast Brazil, with new records, description of three new species and zoogeographical notes. Zootaxa, 4694 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4694.1.1" type="journal article" year="2019">
Lima 
<emphasis id="B2B4244EFFD4FFFDFA21F885FF31F8B8" italics="true" pageId="12" pageNumber="575">et al</emphasis>
. 2019
</bibRefCitation>
). Of the 28 presently recognized species, 26 occur in the Indo-West Pacific: 
<taxonomicName id="47C083DFFFD4FFFDFBF4F8D9FE01F864" authority="A. Milne-Edwards &amp; Bouvier, 1892" authorityName="A. Milne-Edwards &amp; Bouvier" authorityYear="1892" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B2B4244EFFD4FFFDFBF4F8D9FB62F8B8" box="[1117,1278,1890,1916]" italics="true" pageId="12" pageNumber="575">N. longicornis</emphasis>
A. 
<bibRefCitation id="E45185ADFFD4FFFDFA81F8D8FE01F864" author="Milne-Edwards, A. &amp; Bouvier, E. - L." pageId="12" pageNumber="575" pagination="185 - 226" refId="ref12658" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1892) Observations preliminaires sur les paguriens recueillis par les expeditions du Travailleur et du Talisman. Annales des Sciences Naturelles, Zoologie et Paleontologie, Series 7, 13, 185 - 226." type="journal article" year="1892">Milne-Edwards &amp; Bouvier, 1892</bibRefCitation>
</taxonomicName>
is restricted to the eastern Atlantic, and 
<taxonomicName id="47C083DFFFD4FFFDFCC8F83DFA00F864" authority="Lima, Tavares &amp; de Mendonca, 2019" authorityName="Lima, Tavares &amp; de Mendonca" authorityYear="2019" box="[865,1436,1926,1952]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="micheleae">
<emphasis id="B2B4244EFFD4FFFDFCC8F83DFC68F864" box="[865,1012,1926,1952]" italics="true" pageId="12" pageNumber="575">N. micheleae</emphasis>
<bibRefCitation id="E45185ADFFD4FFFDFC52F83CFA00F864" author="Lima, D. &amp; Tavares, M. &amp; Mendonca, J. B. de Jr." box="[1019,1436,1926,1952]" pageId="12" pageNumber="575" pagination="1 - 63" refId="ref11833" refString="Lima, D., Tavares, M. &amp; Mendonca, J. B. de Jr. (2019) Paguroids (Decapoda: Anomura: Diogenidae and Paguridae) of the remote oceanic Archipelago Trindade and Martin Vaz, off southeast Brazil, with new records, description of three new species and zoogeographical notes. Zootaxa, 4694 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4694.1.1" type="journal article" year="2019">Lima, Tavares &amp; de Mendonça, 2019</bibRefCitation>
</taxonomicName>
is known only from Trindade Island, off 
<collectingRegion id="420436BEFFD4FFFDFDF7F810FC9DF800" box="[606,769,1962,1988]" country="Brazil" name="Espirito Santo" pageId="12" pageNumber="575">Espirito Santo</collectingRegion>
, 
<collectingCountry id="F8D7B8CCFFD4FFFDFCA5F810FCCFF800" box="[780,851,1962,1988]" name="Brazil" pageId="12" pageNumber="575">Brazil</collectingCountry>
, southwestern Atlantic.
</paragraph>
<paragraph id="807FF85CFFD4FFFCFF6EF875FAF6FE31" blockId="12.[151,1437,1710,2025]" lastBlockId="13.[151,1437,151,646]" lastPageId="13" lastPageNumber="576" pageId="12" pageNumber="575">
<taxonomicName id="47C083DFFFD4FFFDFF6EF875FE60F82C" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[199,508,1999,2024]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD4FFFDFF6EF875FE60F82C" box="[199,508,1999,2024]" italics="true" pageId="12" pageNumber="575">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD4FFFDFDA9F874FDA7F82C" bold="true" box="[512,571,1998,2024]" pageId="12" pageNumber="575">
<taxonomicNameLabel id="A9879935FFD4FFFDFDA9F874FDA7F82C" box="[512,571,1998,2024]" pageId="12" pageNumber="575" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="47C083DFFFD4FFFDFD1AF875FC4DF82D" authority="Alcock, 1905" authorityName="Alcock" authorityYear="1905" box="[691,977,1998,2025]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B2B4244EFFD4FFFDFD1AF875FCA6F82C" box="[691,826,1998,2024]" italics="true" pageId="12" pageNumber="575">N.gardineri</emphasis>
<bibRefCitation id="E45185ADFFD4FFFDFC97F874FC4DF82D" author="Alcock, A." box="[830,977,1998,2025]" pageId="12" pageNumber="575" pagination="1900" refId="ref10825" refString="Alcock, A. (1905) Marine Crustaceans. 14. Paguridae. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes, being an Account of the Work Carried on and the Collections made by an Expedition during the Years 1899 and 1900. Vol. 2. Cambridge University Press, Cambridge, pp. 827 - 835." type="book chapter" year="1905">Alcock, 1905</bibRefCitation>
</taxonomicName>
(junior synonyms: 
<taxonomicName id="47C083DFFFD4FFFCFB0AF875FE1CFF75" authority="McLaughlin &amp; Hogarth, 1998" authorityName="McLaughlin &amp; Hogarth" authorityYear="1998" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="13" lastPageNumber="576" order="Decapoda" pageId="12" pageNumber="575" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B2B4244EFFD4FFFDFB0AF875FABEF82C" box="[1187,1314,1998,2024]" italics="true" pageId="12" pageNumber="575">N.holthuisi</emphasis>
<bibRefCitation id="E45185ADFFD4FFFCFA8FF874FE1CFF75" author="McLaughlin, P. A. &amp; Hogarth, P. J." lastPageId="13" lastPageNumber="576" pageId="12" pageNumber="575" pagination="1 - 48" refId="ref12252" refString="McLaughlin, P. A. &amp; Hogarth, P. J. (1998) Hermit crabs (Decapoda: Anomura: Paguridea) from the Seychelles. Zoologische Verhandelingen, 318, 1 - 48." type="journal article" year="1998">McLaughlin &amp; Hogarth, 1998</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="47C083DFFFD5FFFCFE25FF2DFD05FF75" authority="Komai, 1999" authorityName="Komai" authorityYear="1999" box="[396,665,151,177]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B2B4244EFFD5FFFCFE25FF2DFE61FF75" box="[396,509,151,177]" italics="true" pageId="13" pageNumber="576">N. pilosus</emphasis>
<bibRefCitation id="E45185ADFFD5FFFCFDACFF2DFD05FF75" author="Komai, T." box="[517,665,151,177]" pageId="13" pageNumber="576" pagination="1 - 66" refId="ref11197" refString="Komai, T. (1999) Hermit crabs of the families Diogenidae and Paguridae (Crustacea: Decapoda: Anomura) collected during the Shin'yo-maru cruise to the Ogasawara Islands and Torishima Island, oceanic islands in Japan. Natural History Research, Special Issue 6, 1 - 66." type="journal article" year="1999">Komai, 1999</bibRefCitation>
</taxonomicName>
) and 
<taxonomicName id="47C083DFFFD5FFFCFD70FF2DFB88FF75" authority="Turkay, 1986" authorityName="Turkay" authorityYear="1986" box="[729,1044,151,177]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="lewinsohni">
<emphasis id="B2B4244EFFD5FFFCFD70FF2DFCEFFF75" box="[729,883,151,177]" italics="true" pageId="13" pageNumber="576">N. lewinsohni</emphasis>
<bibRefCitation id="E45185ADFFD5FFFCFCD2FF2DFB88FF75" author="Turkay, M." box="[891,1044,151,177]" pageId="13" pageNumber="576" pagination="123 - 185" refId="ref13079" refString="Turkay, M. (1986) Crustacea Decapoda Reptantia der Tiefsee des Roten Meeres. Senckenbergiana Maritima, 18 (3 / 6), 123 - 185." type="journal article" year="1986">Türkay, 1986</bibRefCitation>
</taxonomicName>
. Shared characters are: dactylus of right cheliped armed with several spines on dorsal surface; dorsal surfaces of palms of chelae armed with generally even covering of pointed spines over entire surface, but devoid of drop-like sensory structures. 
<taxonomicName id="47C083DFFFD5FFFCFB40FF65FE89FED8" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD5FFFCFB40FF65FE89FED8" italics="true" pageId="13" pageNumber="576">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD5FFFCFEB7FEB9FEC2FED9" bold="true" box="[286,350,259,285]" pageId="13" pageNumber="576">
<taxonomicNameLabel id="A9879935FFD5FFFCFEB7FEB9FEC2FED9" box="[286,350,259,285]" pageId="13" pageNumber="576" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinguished from 
<taxonomicName id="47C083DFFFD5FFFCFDF6FEB9FD77FED9" authorityName="Alcock" authorityYear="1905" box="[607,747,259,285]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B2B4244EFFD5FFFCFDF6FEB9FD77FED9" box="[607,747,259,285]" italics="true" pageId="13" pageNumber="576">N. gardineri</emphasis>
</taxonomicName>
by the following particulars: the cornea is distinctly dilated, of which the width is about half of the ocular peduncle length in 
<taxonomicName id="47C083DFFFD5FFFCFCD3FE9DFB86FE84" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[890,1050,295,320]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD5FFFCFCD3FE9DFB86FE84" box="[890,1050,295,320]" italics="true" pageId="13" pageNumber="576">N. marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD5FFFCFB8BFE9DFBFEFE85" bold="true" box="[1058,1122,295,321]" pageId="13" pageNumber="576">
<taxonomicNameLabel id="A9879935FFD5FFFCFB8BFE9DFBFEFE85" box="[1058,1122,295,321]" pageId="13" pageNumber="576" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="18FBE4D9FFD5FFFCFBDBFE9DFB50FE85" box="[1138,1228,294,321]" captionStart="FIGURE 5" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[164,1421,388,1836]" captionTargetId="figure-128@9.[151,1436,381,1853]" captionTargetPageId="9" captionText="FIGURE 5. Nematopagurus marianicus n. sp.: A–G, holotype, SIO-BIC C14523, male (sl 2.8 mm); H, I, paratype, SIO-BIC C14513, female (sl 2.1 mm). A, shield and cephalic appendages, dorsal view; B, left maxilliped 3, lateral view; C, same, basis and ischium, ventral view; D, left pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, thoracic sternite 6, ventral view; G, I, telson, dorsal view; H, paired pleopods 1, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6110254" httpUri="https://zenodo.org/record/6110254/files/figure.png" pageId="13" pageNumber="576">Fig. 5A</figureCitation>
), whereas it is not particularly dilated, of which the width is 
<quantity id="473855B9FFD5FFFCFDCFFEF1FD4AFEA1" box="[614,726,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" metricValueMax="10.16" metricValueMin="7.62" pageId="13" pageNumber="576" unit="in" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 in</quantity>
<taxonomicName id="47C083DFFFD5FFFCFD72FEF1FCF8FEA1" authorityName="Alcock" authorityYear="1905" box="[731,868,331,357]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B2B4244EFFD5FFFCFD72FEF1FCF8FEA1" box="[731,868,331,357]" italics="true" pageId="13" pageNumber="576">N. gardineri</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="E45185ADFFD5FFFCFC3FFEF1FBECFEA1" author="McLaughlin, P. A." box="[918,1136,331,357]" pageId="13" pageNumber="576" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin 2004b</bibRefCitation>
: fig. 23a); the carpus of the left cheliped bears a dense covering of long iridescent setae on the upper part of the lateral face in 
<taxonomicName id="47C083DFFFD5FFFCFB49FED5FAE3FE4C" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[1248,1407,367,392]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD5FFFCFB49FED5FAE3FE4C" box="[1248,1407,367,392]" italics="true" pageId="13" pageNumber="576">N. marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD5FFFCFA2FFED5FF24FE69" bold="true" pageId="13" pageNumber="576">
<taxonomicNameLabel id="A9879935FFD5FFFCFA2FFED5FF24FE69" pageId="13" pageNumber="576" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="18FBE4D9FFD5FFFCFF6EFE29FEA3FE69" box="[199,319,403,429]" captionStart="FIGURE 7" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[164,1416,193,1427]" captionTargetId="figure-19@11.[151,1436,181,1430]" captionTargetPageId="11" captionText="FIGURE 7. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, left cheliped, mesial view; B, same, lateral view; C, same, chela, dorsal view (setae omitted); D, same, carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6110258" httpUri="https://zenodo.org/record/6110258/files/figure.png" pageId="13" pageNumber="576">Fig. 7B, D</figureCitation>
), while there is no such setal field on the left cheliped carpus in 
<taxonomicName id="47C083DFFFD5FFFCFBAAFE29FB10FE69" authorityName="Alcock" authorityYear="1905" box="[1027,1164,403,429]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B2B4244EFFD5FFFCFBAAFE29FB10FE69" box="[1027,1164,403,429]" italics="true" pageId="13" pageNumber="576">N. gardineri</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="E45185ADFFD5FFFCFB17FE29FA04FE69" author="McLaughlin, P. A." box="[1214,1432,403,429]" pageId="13" pageNumber="576" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin 2004b</bibRefCitation>
: fig. 23b, c; see also 
<bibRefCitation id="E45185ADFFD5FFFCFED0FE0DFD95FE15" author="Komai, T." box="[377,521,439,465]" pageId="13" pageNumber="576" pagination="1 - 66" refId="ref11197" refString="Komai, T. (1999) Hermit crabs of the families Diogenidae and Paguridae (Crustacea: Decapoda: Anomura) collected during the Shin'yo-maru cruise to the Ogasawara Islands and Torishima Island, oceanic islands in Japan. Natural History Research, Special Issue 6, 1 - 66." type="journal article" year="1999">Komai 1999</bibRefCitation>
: figs. 16E, 18E, as 
<taxonomicName id="47C083DFFFD5FFFCFD4AFE0DFCC8FE15" authorityName="Komai" authorityYear="1999" box="[739,852,439,465]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B2B4244EFFD5FFFCFD4AFE0DFCC8FE15" box="[739,852,439,465]" italics="true" pageId="13" pageNumber="576">N. pilosus</emphasis>
</taxonomicName>
); and the dactyli of the pereopods 2 and 3 are more elongate and slender in 
<taxonomicName id="47C083DFFFD5FFFCFE08FE61FDDCFE30" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[417,576,475,500]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus">
<emphasis id="B2B4244EFFD5FFFCFE08FE61FDDCFE30" box="[417,576,475,500]" italics="true" pageId="13" pageNumber="576">N. marianicus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="47C083DFFFD5FFFCFD32FE61FCB8FE31" authorityName="Alcock" authorityYear="1905" box="[667,804,475,501]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="gardineri">
<emphasis id="B2B4244EFFD5FFFCFD32FE61FCB8FE31" box="[667,804,475,501]" italics="true" pageId="13" pageNumber="576">N. gardineri</emphasis>
</taxonomicName>
(
<figureCitation id="18FBE4D9FFD5FFFCFC9AFE61FC30FE31" box="[819,940,475,501]" captionStart="FIGURE 8" captionStartId="12.[151,250,1567,1591]" captionTargetBox="[161,1423,195,1529]" captionTargetId="figure-19@12.[151,1436,181,1544]" captionTargetPageId="12" captionText="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="13" pageNumber="576">Fig. 8A–D</figureCitation>
versus 
<bibRefCitation id="E45185ADFFD5FFFCFBABFE61FB41FE31" author="McLaughlin, P. A." box="[1026,1245,475,501]" pageId="13" pageNumber="576" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin 2004b</bibRefCitation>
: fig. 23d, e).
</paragraph>
<paragraph id="807FF85CFFD5FFFCFF6EFE45FAF3FD41" blockId="13.[151,1437,151,646]" pageId="13" pageNumber="576">
<taxonomicName id="47C083DFFFD5FFFCFF6EFE45FD9DFDDC" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[199,513,511,536]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD5FFFCFF6EFE45FD9DFDDC" box="[199,513,511,536]" italics="true" pageId="13" pageNumber="576">Nematopagurus marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD5FFFCFDA3FE45FDD6FDDD" bold="true" box="[522,586,511,537]" pageId="13" pageNumber="576">
<taxonomicNameLabel id="A9879935FFD5FFFCFDA3FE45FDD6FDDD" box="[522,586,511,537]" pageId="13" pageNumber="576" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is easily differentiated from 
<taxonomicName id="47C083DFFFD5FFFCFC3EFE45FBAFFDDD" authorityName="Turkay" authorityYear="1986" box="[919,1075,511,537]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="lewinsohni">
<emphasis id="B2B4244EFFD5FFFCFC3EFE45FBAFFDDD" box="[919,1075,511,537]" italics="true" pageId="13" pageNumber="576">N. lewinsohni</emphasis>
</taxonomicName>
by the lack of a row of bristlelike setae on the dorsal surface of each ambulatory dactylus (
<figureCitation id="18FBE4D9FFD5FFFCFCC9FD99FC43FDF9" box="[864,991,547,573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1567,1591]" captionTargetBox="[161,1423,195,1529]" captionTargetId="figure-19@12.[151,1436,181,1544]" captionTargetPageId="12" captionText="FIGURE 8. Nematopagurus marianicus n. sp., holotype, SIO-BIC C14523, male (sl 2.8 mm).A, right pereopod 2, lateral view; B, same, carpus, mesial view; C, same, dactylus and distal part of propodus, mesial view; D, left pereopod 3, lateral view; E, same, dactylus and distal part of propodus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.6110260" httpUri="https://zenodo.org/record/6110260/files/figure.png" pageId="13" pageNumber="576">Fig. 8A–D</figureCitation>
), which is present in the latter species (
<bibRefCitation id="E45185ADFFD5FFFCFF36FDFDFEE1FDA5" author="McLaughlin, P. A." box="[159,381,583,609]" pageId="13" pageNumber="576" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin 2004b</bibRefCitation>
: fig. 22c). The carpus of the left cheliped lacks a dense setal field on the lateral face in 
<taxonomicName id="47C083DFFFD5FFFCFAF4FDFDFF60FD41" authorityName="Turkay" authorityYear="1986" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="lewinsohni">
<emphasis id="B2B4244EFFD5FFFCFAF4FDFDFF60FD41" italics="true" pageId="13" pageNumber="576">N. lewinsohni</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="E45185ADFFD5FFFCFE99FDD1FDD8FD41" author="McLaughlin, P. A." box="[304,580,619,645]" pageId="13" pageNumber="576" pagination="151 - 229" refId="ref12127" refString="McLaughlin, P. A. (2004 b) A review of the hermit crab genus Nematopagurus A. Milne-Edwards &amp; Bouvier, 1892 and the descriptions of five new species. In: Marshall, B. &amp; Richer de Forges, B. (Eds.), Tropical Deep-Sea Benthos. Vol. 23. Memoirs du Museum National d'Histoire Naturelle, 191, pp. 151 - 229." type="journal article" year="2004">McLaughlin 2004b: 210</bibRefCitation>
, fig. 22b), which is present in 
<taxonomicName id="47C083DFFFD5FFFCFC3EFDD1FBA9FD40" authority="Komai &amp; Miller &amp; Malay, 2022" authorityName="Komai &amp; Miller &amp; Malay" authorityYear="2022" box="[919,1077,619,644]" class="Malacostraca" family="Paguridae" genus="Nematopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="576" phylum="Arthropoda" rank="species" species="marianicus" status="sp. nov.">
<emphasis id="B2B4244EFFD5FFFCFC3EFDD1FBA9FD40" box="[919,1077,619,644]" italics="true" pageId="13" pageNumber="576">N. marianicus</emphasis>
</taxonomicName>
<emphasis id="B2B4244EFFD5FFFCFB95FDD1FBE7FD41" bold="true" box="[1084,1147,619,645]" pageId="13" pageNumber="576">
<taxonomicNameLabel id="A9879935FFD5FFFCFB95FDD1FBE7FD41" box="[1084,1147,619,645]" pageId="13" pageNumber="576" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, as mentioned above.
</paragraph>
</subSubSection>
</treatment>
</document>