256 lines
27 KiB
XML
256 lines
27 KiB
XML
<document id="3EE610218249E79B8A6467714BA490D0" ID-DOI="10.26107/RBZ-2019-0044" ID-ISSN="2345-7600" ID-Zenodo-Dep="5343685" ID-ZooBank="102C469A-1F3A-4660-92DB-DE59CCCCAC6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630197404672" checkinUser="diego" docAuthor="Páll-Gergely, Barna & Grego, Jozef" docDate="2019" docId="0843F96EB46B3350FF7807E10AB8AA44" docLanguage="en" docName="RBZ-2019-0044.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Vargapupa biheli subsp. viridis Páll-Gergely & Grego 2019, new subspecies" docType="treatment" docVersion="3" lastPageNumber="591" masterDocId="F47A8116B46E3355FF9A00740955AD52" masterDocTitle="Notes on Pseudopomatias Möllendorff, 1885 and Vargapupa Páll- Gergely, 2015 (Gastropoda: Caenogastropoda: Pupinidae)" masterLastPageNumber="594" masterPageNumber="586" pageNumber="591" updateTime="1699183092722" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="B2719FE234AD38892C27649F1134FA56">
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<mods:title id="3928CE959EEA02EC2CF18C054AD55D03">Notes on Pseudopomatias Möllendorff, 1885 and Vargapupa Páll- Gergely, 2015 (Gastropoda: Caenogastropoda: Pupinidae)</mods:title>
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<mods:namePart id="6A57CCAC1E5458EC8914311F3771D08C">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="ECC767669721B5AE2D67A79095804924">Grego, Jozef</mods:namePart>
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<mods:typeOfResource id="1D78F94500C0F366D780C6E9A69AC385">text</mods:typeOfResource>
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<mods:title id="10FE4BE3DC52C2E6114AF97276F10FFE">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="E66240A063E7EBE21BFCB8269B42A80B">
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<mods:date id="762840DEA6898E7B0E6311FDDDDFAC3B">2019</mods:date>
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<mods:number id="EA9B9DA62EC33E1438D2E06CDB2A861F">2019-11-21</mods:number>
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<mods:number id="A2030074D5299589FDA9958C334D88A3">67</mods:number>
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<mods:classification id="92FA7B04B84033BED1FA279D7B825092">journal article</mods:classification>
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<mods:identifier id="4004E2E8CC8FA058DCA3CE0CF4D95983" type="DOI">10.26107/RBZ-2019-0044</mods:identifier>
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<mods:identifier id="4E4FC042A3C7226F5EDE02020C89EB2A" type="ISSN">2345-7600</mods:identifier>
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<treatment id="0843F96EB46B3350FF7807E10AB8AA44" LSID="urn:lsid:plazi:treatment:0843F96EB46B3350FF7807E10AB8AA44" httpUri="http://treatment.plazi.org/id/0843F96EB46B3350FF7807E10AB8AA44" lastPageNumber="591" pageId="5" pageNumber="591">
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<subSubSection id="C8F01BF3B46B3350FF7807E10BFBAAFE" box="[226,686,1940,1965]" pageId="5" pageNumber="591" type="nomenclature">
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<paragraph id="80554878B46B3350FF7807E10BFBAAFE" blockId="5.[226,686,1940,1997]" box="[226,686,1940,1965]" pageId="5" pageNumber="591">
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<heading id="DB1DFF14B46B3350FF7807E10BFBAAFE" bold="true" box="[226,686,1940,1965]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="591" reason="2">
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<emphasis id="B29E946AB46B3350FF7807E10BFBAAFE" bold="true" box="[226,686,1940,1965]" pageId="5" pageNumber="591">
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<taxonomicName id="47EA33FBB46B3350FF7807E108A4AAFE" ID-CoL="7V42B" authority="Páll-Gergely & Grego, 2019" authorityName="Páll-Gergely & Grego" authorityYear="2019" box="[226,497,1940,1964]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="subSpecies" species="biheli" status="subsp. nov." subSpecies="viridis">
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||
<emphasis id="B29E946AB46B3350FF7807E108A4AAFE" bold="true" box="[226,497,1940,1964]" italics="true" pageId="5" pageNumber="591">Vargapupa biheli viridis</emphasis>
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A9AD2911B46B3350FD9A07E10BFBAAFE" box="[512,686,1940,1965]" pageId="5" pageNumber="591" rank="subSpecies">new subspecies</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C8F01BF3B46B3350FE1C07C10B5CAA9E" box="[390,521,1972,1997]" pageId="5" pageNumber="591" type="description">
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<paragraph id="80554878B46B3350FE1C07C10B5CAA9E" blockId="5.[226,686,1940,1997]" box="[390,521,1972,1997]" pageId="5" pageNumber="591">
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(
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C8F01BF3B46B3350FF1207810DFAAF04" pageId="5" pageNumber="591" type="materials_examined">
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<paragraph id="80554878B46B3350FF1207810A5DA57F" blockId="5.[136,776,2036,2093]" pageId="5" pageNumber="591">
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<emphasis id="B29E946AB46B3350FF1207810861A55E" bold="true" box="[136,308,2036,2061]" pageId="5" pageNumber="591">Type material.</emphasis>
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||
<materialsCitation id="30824225B46B3350FEA407810A5DA57F" collectionCode="HNHM" country="Laos" location="Xianghouang prov." pageId="5" pageNumber="591" specimenCode="HNHM 104398" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5F51F6DAB46B3350FEA4078108F7A55F" box="[318,418,2037,2061]" pageId="5" pageNumber="591" type="holotype">Holotype</typeStatus>
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(H:
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<quantity id="4712E59DB46B3350FE4007810B6AA55E" box="[474,575,2037,2061]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="5" pageNumber="591" unit="mm" value="10.2">10.2 mm</quantity>
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, D:
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<quantity id="4712E59DB46B3350FDE907810B9EA55E" box="[627,715,2037,2061]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="591" unit="mm" value="3.6">3.6 mm</quantity>
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,
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<figureCitation id="18D154FDB46B3350FD4307810993A57E" captionStart="Fig" captionStartId="6.[113,148,1970,1992]" captionTargetBox="[115,1449,249,1954]" captionTargetId="figure-66@6.[115,1449,249,1955]" captionTargetPageId="6" captionText="Fig. 5. A–F, Vargapupa biheli viridis, new subspecies (holotype, HNHM 104398); G–L, Vargapupa biheli viridis, new subspecies (paratype, HNHM 104399); M–Q, Vargapupa biheli biheli Páll-Gergely, 2015 (holotype, MNHN-IM-2012–27020). Scale ‘a’ refers to images of the whole shell, scale ‘b’ refers to images of the body whorl, both scales represent 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343701" httpUri="https://zenodo.org/record/5343701/files/figure.png" pageId="5" pageNumber="591">Figs 5A–F</figureCitation>
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||
) (
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||
<specimenCode id="D04CE003B46B3350FF40086108D3A57F" box="[218,390,2069,2093]" collectionCode="HNHM" country="Hungary" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="591">HNHM 104398</specimenCode>
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),
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||
<collectingCountry id="F8FD08E8B46B3350FE0D08610898A57E" box="[407,461,2069,2092]" name="Laos" pageId="5" pageNumber="591">Laos</collectingCountry>
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,
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<location id="85351EA3B46B3350FE4D08610BF2A57E" LSID="urn:lsid:plazi:treatment:0843F96EB46B3350FF7807E10AB8AA44:85351EA3B46B3350FE4D08610BF2A57E" box="[471,679,2069,2093]" country="Laos" name="Xianghouang prov." pageId="5" pageNumber="591">Xianghouang prov.</location>
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||
, ca.
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<quantity id="4712E59DB46B3350FD4908610A5DA57F" box="[723,776,2069,2093]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="591" unit="km" value="4.0">4 km</quantity>
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</materialsCitation>
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</paragraph>
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<paragraph id="80554878B46B3350FCD900CA0DFAAF04" blockId="5.[835,1474,190,598]" pageId="5" pageNumber="591">
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<materialsCitation id="30824225B46B3350FCD900CA0D5FAC44" collectingDate="2017-02-23" collectorName="J. Grego" elevation="1230" latitude="19.377832" location="Ban Khong" longLatPrecision="1" longitude="102.979034" pageId="5" pageNumber="591" specimenCount="1">
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SW of
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||
<location id="85351EA3B46B3350FC0C00CB0D42AD84" LSID="urn:lsid:plazi:treatment:0843F96EB46B3350FF7807E10AB8AA44:85351EA3B46B3350FC0C00CB0D42AD84" box="[918,1047,190,214]" latitude="19.377832" longLatPrecision="1" longitude="102.979034" name="Ban Khong" pageId="5" pageNumber="591">Ban Khong</location>
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, cliffs foot above larger vertical cave,
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<elevation id="0BC7AF4BB46B3350FCD900AA0A8FADA4" box="[835,986,222,246]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="591" unit="m" value="1230.0">
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<quantity id="4712E59DB46B3350FCD900AA0ACBADA4" box="[835,926,222,246]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="591" unit="m" value="1230.0">1,230 m</quantity>
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a.s.l.
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||
</elevation>
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,
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<geoCoordinate id="E5DE2EBFB46B3350FC7000A90D21ADA4" box="[1002,1140,221,246]" degrees="19" direction="north" minutes="22.670" orientation="latitude" pageId="5" pageNumber="591" precision="1" value="19.377832">19°22.670′N</geoCoordinate>
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||
,
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<geoCoordinate id="E5DE2EBFB46B3350FB1800A90C42ADA4" box="[1154,1303,221,246]" degrees="102" direction="east" minutes="58.742" orientation="longitude" pageId="5" pageNumber="591" precision="1" value="102.979034">102°58.742′E</geoCoordinate>
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||
, coll.
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||
<collectorName id="2D1F2DAEB46B3350FAC400AB0CEBADA4" box="[1374,1470,222,246]" pageId="5" pageNumber="591">J. Grego</collectorName>
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,
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||
<date id="F4546EB8B46B3350FCD9008A0D5FAC44" box="[835,1034,254,278]" pageId="5" pageNumber="591" value="2017-02-23">
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||
<collectingDate id="E4109750B46B3350FCD9008A0D5FAC44" box="[835,1034,254,278]" pageId="5" pageNumber="591" value="2017-02-23">23 February 2017</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="30824225B46B3350FB8D008A0DF9AF04" collectingDate="2017-02-23" collectionCode="SMF, NHMUK" collectorName="J. Grego" country="Laos" elevation="1060" latitude="19.387516" location="Xianghouang prov." longLatPrecision="1" longitude="102.97497" municipality="Caverns" pageId="5" pageNumber="591" specimenCode="SMF 326927, NHMUK 20190575" specimenCount="55" specimenCount-juvenile="2" typeStatus="paratype">
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||
figured
|
||
<typeStatus id="5F51F6DAB46B3350FBF1008B0D92AC45" box="[1131,1223,255,279]" pageId="5" pageNumber="591" type="paratype">paratype</typeStatus>
|
||
(HNHM 104399,
|
||
<figureCitation id="18D154FDB46B3350FA0E008A0AD1AC64" captionStart="Fig" captionStartId="6.[113,148,1970,1992]" captionTargetBox="[115,1449,249,1954]" captionTargetId="figure-66@6.[115,1449,249,1955]" captionTargetPageId="6" captionText="Fig. 5. A–F, Vargapupa biheli viridis, new subspecies (holotype, HNHM 104398); G–L, Vargapupa biheli viridis, new subspecies (paratype, HNHM 104399); M–Q, Vargapupa biheli biheli Páll-Gergely, 2015 (holotype, MNHN-IM-2012–27020). Scale ‘a’ refers to images of the whole shell, scale ‘b’ refers to images of the body whorl, both scales represent 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343701" httpUri="https://zenodo.org/record/5343701/files/figure.png" pageId="5" pageNumber="591">Figs 5G–L</figureCitation>
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||
), same data as for holotype;
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||
<specimenCount id="96EC83F1B46B3350FB3C016A0C49AC65" box="[1190,1308,286,311]" pageId="5" pageNumber="591" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
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(
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<specimenCode id="D04CE003B46B3350FAB3016A0CE2AC64" box="[1321,1463,286,310]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="591">SMF 326927</specimenCode>
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||
), same data as for
|
||
<typeStatus id="5F51F6DAB46B3350FC6A014A0D05AC04" box="[1008,1104,318,342]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
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;
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<specimenCount id="96EC83F1B46B3350FBC3014A0D87AC05" box="[1113,1234,318,343]" pageId="5" pageNumber="591" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
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(
|
||
<specimenCode id="D04CE003B46B3350FB47014A0CE2AC04" box="[1245,1463,318,342]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="591" type="Museum">NHMUK 20190575</specimenCode>
|
||
), same data as for
|
||
<typeStatus id="5F51F6DAB46B3350FC63012A0D0FAC24" box="[1017,1114,350,374]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="96EC83F1B46B3350FBFF012A0D83AC25" box="[1125,1238,350,375]" pageId="5" pageNumber="591" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
(ZRC.MOL.015633), same data as for
|
||
<typeStatus id="5F51F6DAB46B3350FC61010A0D09ACC4" box="[1019,1116,382,406]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="96EC83F1B46B3350FBFD010A0DBAACC5" box="[1127,1263,382,407]" pageId="5" pageNumber="591" type="generic" typeStatus="paratypes">47 paratypes</specimenCount>
|
||
(JG/47), same data as for
|
||
<typeStatus id="5F51F6DAB46B3350FC1E01EA0AB0ACE4" box="[900,997,414,438]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="96EC83F1B46B3350FC7501EA0D0DACE4" box="[1007,1112,414,438]" pageId="5" pageNumber="591" type="juvenile">2 juvenile</specimenCount>
|
||
shells (not
|
||
<typeStatus id="5F51F6DAB46B3350FB4B01EB0C6FACE5" box="[1233,1338,415,439]" pageId="5" pageNumber="591" type="paratype">paratypes</typeStatus>
|
||
, JG/2), same data as for
|
||
<typeStatus id="5F51F6DAB46B3350FC2101CA0D49AC84" box="[955,1052,446,470]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="96EC83F1B46B3350FBBD01CA0DCDAC85" box="[1063,1176,446,471]" pageId="5" pageNumber="591" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
broken to two parts (JG/1),
|
||
<collectingCountry id="F8FD08E8B46B3350FCD901AB0A2CACA4" box="[835,889,479,502]" name="Laos" pageId="5" pageNumber="591">Laos</collectingCountry>
|
||
,
|
||
<location id="85351EA3B46B3350FC1F01AA0D03ACA4" LSID="urn:lsid:plazi:treatment:0843F96EB46B3350FF7807E10AB8AA44:85351EA3B46B3350FC1F01AA0D03ACA4" box="[901,1110,478,502]" country="Laos" latitude="19.387516" longLatPrecision="1" longitude="102.97497" municipality="Caverns" name="Xianghouang prov." pageId="5" pageNumber="591">Xianghouang prov.</location>
|
||
,
|
||
<collectingMunicipality id="6031D202B46B3350FBFB01AA0DEDACA4" box="[1121,1208,478,502]" pageId="5" pageNumber="591">Caverns</collectingMunicipality>
|
||
among boulders at large sinkhole below small settlements (5 houses about
|
||
<locationDeviation id="91E7515AB46B3350FAC6018A0ABFAF64" location="Bang Knong" pageId="5" pageNumber="591">
|
||
<quantity id="4712E59DB46B3350FAC6018A0CC6AF44" box="[1372,1427,510,534]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="591" unit="km" value="6.0">6 km</quantity>
|
||
SW of Bang Knong
|
||
</locationDeviation>
|
||
),
|
||
<elevation id="0BC7AF4BB46B3350FC61026A0DDCAF64" box="[1019,1161,542,566]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="5" pageNumber="591" unit="m" value="1060.0">
|
||
<quantity id="4712E59DB46B3350FC61026A0D07AF64" box="[1019,1106,542,566]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="5" pageNumber="591" unit="m" value="1060.0">1,060 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="E5DE2EBFB46B3350FB0F02690C75AF64" box="[1173,1312,541,566]" degrees="19" direction="north" minutes="23.251" orientation="latitude" pageId="5" pageNumber="591" precision="1" value="19.387516">19°23.251′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E5DE2EBFB46B3350FAB302690CEBAF64" box="[1321,1470,541,566]" degrees="102" direction="east" minutes="58.498" orientation="longitude" pageId="5" pageNumber="591" precision="1" value="102.97497">102°58.498′E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="2D1F2DAEB46B3350FCE1024B0A8CAF04" box="[891,985,574,598]" pageId="5" pageNumber="591">J. Grego</collectorName>
|
||
,
|
||
<date id="F4546EB8B46B3350FC7C024A0DF9AF04" box="[998,1196,574,598]" pageId="5" pageNumber="591" value="2017-02-23">
|
||
<collectingDate id="E4109750B46B3350FC7C024A0DF9AF04" box="[998,1196,574,598]" pageId="5" pageNumber="591" value="2017-02-23">23 February 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD9020A0C4EAFE4" pageId="5" pageNumber="591" type="diagnosis">
|
||
<paragraph id="80554878B46B3350FCD9020A0C4EAFE4" blockId="5.[835,1474,638,694]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD9020A0AEDAFC4" bold="true" box="[835,952,638,662]" pageId="5" pageNumber="591">Diagnosis.</emphasis>
|
||
A subspecies of
|
||
<taxonomicName id="47EA33FBB46B3350FBF5020B0C7EAFC4" authorityName="Pall-Gergely" authorityYear="2015" box="[1135,1323,638,662]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="biheli">
|
||
<emphasis id="B29E946AB46B3350FBF5020B0C7EAFC4" box="[1135,1323,638,662]" italics="true" pageId="5" pageNumber="591">Vargapupa biheli</emphasis>
|
||
</taxonomicName>
|
||
with greenish shell colour and weaker periumbilical keel.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD902AA0C97AFA4" box="[835,1474,734,758]" pageId="5" pageNumber="591" type="description">
|
||
<paragraph id="80554878B46B3350FCD902AA0C97AFA4" blockId="5.[835,1474,734,758]" box="[835,1474,734,758]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD902AA0D00AFA4" bold="true" box="[835,1109,734,758]" pageId="5" pageNumber="591">Measurements (in mm).</emphasis>
|
||
H = 8.4–11, D = 3.4–3.7 (n = 7).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD9036A0DC6A8C4" pageId="5" pageNumber="591" type="diagnosis">
|
||
<paragraph id="80554878B46B3350FCD9036A0D9CA9C4" blockId="5.[835,1475,798,1174]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD9036A0D01AE64" bold="true" box="[835,1108,798,822]" pageId="5" pageNumber="591">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="47EA33FBB46B3350FBFA036B0CD5AE64" authority="Páll-Gergely & Grego, 2019" authorityName="Páll-Gergely & Grego" authorityYear="2019" box="[1120,1408,798,822]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="subSpecies" species="biheli" status="subsp. nov." subSpecies="viridis">
|
||
<emphasis id="B29E946AB46B3350FBFA036B0CD5AE64" box="[1120,1408,798,822]" italics="true" pageId="5" pageNumber="591">Vargapupa biheli viridis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A9AD2911B46B3350FA0E036B0AE0AE04" pageId="5" pageNumber="591" rank="subSpecies">new subspecies</taxonomicNameLabel>
|
||
differs from the nominotypical subspecies (
|
||
<figureCitation id="18D154FDB46B3350FA0D034A0ADEAE24" captionStart="Fig" captionStartId="6.[113,148,1970,1992]" captionTargetBox="[115,1449,249,1954]" captionTargetId="figure-66@6.[115,1449,249,1955]" captionTargetPageId="6" captionText="Fig. 5. A–F, Vargapupa biheli viridis, new subspecies (holotype, HNHM 104398); G–L, Vargapupa biheli viridis, new subspecies (paratype, HNHM 104399); M–Q, Vargapupa biheli biheli Páll-Gergely, 2015 (holotype, MNHN-IM-2012–27020). Scale ‘a’ refers to images of the whole shell, scale ‘b’ refers to images of the body whorl, both scales represent 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5343701" httpUri="https://zenodo.org/record/5343701/files/figure.png" pageId="5" pageNumber="591">Fig. 5M–Q</figureCitation>
|
||
) by the lower, slightly denser ribs and weaker keel. The keel of
|
||
<taxonomicName id="47EA33FBB46B3350FC50030B0D80AEC4" authorityName="Páll-Gergely & Grego" authorityYear="2019" box="[970,1237,894,918]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="subSpecies" species="biheli" subSpecies="viridis">
|
||
<emphasis id="B29E946AB46B3350FC50030B0D80AEC4" box="[970,1237,894,918]" italics="true" pageId="5" pageNumber="591">Vargapupa biheli viridis</emphasis>
|
||
</taxonomicName>
|
||
is less clearly visible from standard apertural view (i.e., more visible in
|
||
<taxonomicName id="47EA33FBB46B3350FAD103EB0A96AE84" authority="biheli" authorityName="biheli Pall-Gergely" authorityYear="2015" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="subSpecies" species="biheli" subSpecies="biheli">
|
||
<emphasis id="B29E946AB46B3350FAD103EB0A96AE84" italics="true" pageId="5" pageNumber="591">Vargapupa biheli biheli</emphasis>
|
||
</taxonomicName>
|
||
), and it is overall less elevated, and less marked by deep sutures on both sides. Fresh shells of the new subspecies are semitransparent greenish. Although we had no possibility to examine fresh shells of the nominotypical subspecies, the reddish colouration of the apical whorls of some shells, such as the
|
||
<typeStatus id="5F51F6DAB46B3350FBD3042A0DFCA924" box="[1097,1193,1118,1142]" pageId="5" pageNumber="591" type="holotype">holotype</typeStatus>
|
||
, indicates that fresh shells might have a reddish brown colour.
|
||
</paragraph>
|
||
<paragraph id="80554878B46B3350FCD904CA0DC6A8C4" blockId="5.[835,1474,1214,1430]" pageId="5" pageNumber="591">
|
||
The differences mentioned between
|
||
<taxonomicName id="47EA33FBB46B3350FB5A04CB0CDEA984" authorityName="Pall-Gergely" authorityYear="2016" box="[1216,1419,1214,1238]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="humilis">
|
||
<emphasis id="B29E946AB46B3350FB5A04CB0CDEA984" box="[1216,1419,1214,1238]" italics="true" pageId="5" pageNumber="591">Vargapupa humilis</emphasis>
|
||
</taxonomicName>
|
||
Páll- Gergely, 2016 and
|
||
<taxonomicName id="47EA33FBB46B3350FB8F04AB0D27A9A4" authorityName="Pall-Gergely" authorityYear="2015" box="[1045,1138,1246,1270]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="biheli">
|
||
<emphasis id="B29E946AB46B3350FB8F04AB0D27A9A4" box="[1045,1138,1246,1270]" italics="true" pageId="5" pageNumber="591">V. biheli</emphasis>
|
||
</taxonomicName>
|
||
are true to the new subspecies as well. These are the following:
|
||
<taxonomicName id="47EA33FBB46B3350FB24048B0C2BA844" authorityName="Pall-Gergely" authorityYear="2015" box="[1214,1406,1278,1302]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="biheli">
|
||
<emphasis id="B29E946AB46B3350FB24048B0C2BA844" box="[1214,1406,1278,1302]" italics="true" pageId="5" pageNumber="591">Vargapupa biheli</emphasis>
|
||
</taxonomicName>
|
||
has a much stronger keel, a slender triangular shell (rather spindleshaped in
|
||
<taxonomicName id="47EA33FBB46B3350FC29054B0D77A804" authorityName="Pall-Gergely" authorityYear="2016" box="[947,1058,1342,1366]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="humilis">
|
||
<emphasis id="B29E946AB46B3350FC29054B0D77A804" box="[947,1058,1342,1366]" italics="true" pageId="5" pageNumber="591">V. humilis</emphasis>
|
||
</taxonomicName>
|
||
), a less oblique peristome to the shell axis and a somewhat rectangular aperture (nearly rounded, teardrop-shaped in
|
||
<taxonomicName id="47EA33FBB46B3350FB8F050B0DD1A8C4" authorityName="Pall-Gergely" authorityYear="2016" box="[1045,1156,1406,1430]" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="species" species="humilis">
|
||
<emphasis id="B29E946AB46B3350FB8F050B0DD1A8C4" box="[1045,1156,1406,1430]" italics="true" pageId="5" pageNumber="591">V. humilis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD905CA0C96AB64" pageId="5" pageNumber="591" type="discussion">
|
||
<paragraph id="80554878B46B3350FCD905CA0C96AB64" blockId="5.[835,1475,1470,1590]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD905CA0AE2A884" bold="true" box="[835,951,1470,1494]" pageId="5" pageNumber="591">Remarks.</emphasis>
|
||
We initially considered to handling the two subspecies as species in their own right. However, due to the similar size, shape and the lack of fresh specimens of
|
||
<taxonomicName id="47EA33FBB46B3350FAF0058B0AD5AB64" authority="biheli" authorityName="biheli Pall-Gergely" authorityYear="2015" class="Gastropoda" family="Pupinidae" genus="Vargapupa" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="591" phylum="Mollusca" rank="subSpecies" species="biheli" subSpecies="biheli">
|
||
<emphasis id="B29E946AB46B3350FAF0058B0AD5AB64" italics="true" pageId="5" pageNumber="591">V. biheli biheli</emphasis>
|
||
</taxonomicName>
|
||
, we rather keep the new taxon on a subspecific level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD9062A0D89ABC4" pageId="5" pageNumber="591" type="etymology">
|
||
<paragraph id="80554878B46B3350FCD9062A0D89ABC4" blockId="5.[835,1474,1630,1686]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD9062A0A90AB24" bold="true" box="[835,965,1630,1654]" pageId="5" pageNumber="591">Etymology.</emphasis>
|
||
The name “viridis” refers to the greenish colour of fresh shells of the new subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8F01BF3B46B3350FCD906CA0AB8AA44" pageId="5" pageNumber="591" type="distribution">
|
||
<paragraph id="80554878B46B3350FCD906CA0AB8AA44" blockId="5.[835,1474,1726,1814]" pageId="5" pageNumber="591">
|
||
<emphasis id="B29E946AB46B3350FCD906CA0A8AAB84" bold="true" box="[835,991,1726,1750]" pageId="5" pageNumber="591">Distribution.</emphasis>
|
||
The new subspecies is known from two nearby sites, practically a single population in Xianghouang Province,
|
||
<collectingCountry id="F8FD08E8B46B3350FC2B068B0ABDAA44" box="[945,1000,1791,1814]" name="Laos" pageId="5" pageNumber="591">Laos</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |