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<mods:title id="47F309A4EA56573316A84A5554185DBE">Taxonomic revision of the jumping spider genus Tullgrenella Mello-Leitão, 1941 (Araneae: Salticidae: Freyina)</mods:title>
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; gender feminine, by original designation); 
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Males of 
<taxonomicName id="47B24D53FFCB4C016FB059D9FDF10481" authorityName="Mello-Leitao" authorityYear="1941" box="[393,529,418,444]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016FB059D9FDF10481" box="[393,529,418,444]" italics="true" pageId="5" pageNumber="6">Tullgrenella</emphasis>
</taxonomicName>
resemble those of other related 
<taxonomicName id="47B24D53FFCB4C016DBC59D9FC3B0481" authorityName=", Edwards" authorityYear="2015" box="[901,987,418,444]" class="Arachnida" family="Hahniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="65" phylum="Arthropoda" rank="subTribe" subTribe="Freyina">Freyina</taxonomicName>
genera, such as: 
<taxonomicName id="47B24D53FFCB4C016A9A59D9FF2F04DC" authority="C. L. Kock, 1850" authorityName="C. L. Kock" authorityYear="1850" class="Arachnida" family="Salticidae" genus="Aphirape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016A9A59D9FAEB0481" box="[1187,1291,418,444]" italics="true" pageId="5" pageNumber="6">Aphirape</emphasis>
C. L. Kock, 1850
</taxonomicName>
by having a hook-shaped RTA retro-dorsally (see 
<bibRefCitation id="E4234B21FFCB4C016D1A59BDFC0F04DD" author="Galiano, M. E." box="[803,1007,454,480]" pageId="5" pageNumber="6" pagination="93 - 111" refId="ref42229" refString="Galiano, M. E. (1981 b) Revision del genero Aphirape C. L. Koch, 1851 (Araneae, Salticidae). Comunicaciones del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, Entomologia, 1, 93 - 111." type="journal article" year="1981">Galiano 1981b: 6</bibRefCitation>
, figs 21, 23); 
<taxonomicName id="47B24D53FFCB4C016AA259BDFF2F0739" authority="Galiano, 1983" authorityName="Galiano" authorityYear="1983" class="Arachnida" family="Salticidae" genus="Sumampattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016AA259BDFAD204DD" box="[1179,1330,454,480]" italics="true" pageId="5" pageNumber="6">Sumampattus</emphasis>
Galiano, 1983
</taxonomicName>
by the presence of endites with distal projections (see 
<bibRefCitation id="E4234B21FFCB4C016D6A5991FC120739" author="Galiano, M. E." box="[851,1010,490,516]" pageId="5" pageNumber="6" pagination="323 - 355" refId="ref42099" refString="Galiano, M. E. (1970) Revision del genero Tullgrenella Mello-Leitao, 1941 (Araneae, Salticidae). Physis, Revista de la Sociedad Argentina de Ciencias Naturales, Seccio´n C, 29, 323 - 355." type="journal article" year="1970">Galiano 1970</bibRefCitation>
: fig. 15; 
<bibRefCitation id="E4234B21FFCB4C016A675991FAE90739" author="Galiano, M. E." box="[1118,1289,490,516]" pageId="5" pageNumber="6" pagination="216 - 218" refId="ref42195" refString="Galiano, M. E. (1981 a) Three new species of Salticidae (Araneae). Bulletin of the American Museum of Natural History, 170, 216 - 218." type="journal article" year="1981">Galiano 1981a</bibRefCitation>
: figs 1–2, 9; Galiano 1983: fig. 8), tegular membranous process (
<figureCitation id="18892A55FFCB4C016CE05A75FC800715" box="[729,864,526,552]" captionStart="FIGURE 2" captionStartId="7.[151,250,1490,1514]" captionTargetBox="[151,1372,181,1459]" captionTargetId="figure-17@7.[151,1435,181,1465]" captionTargetPageId="7" captionText="FIGURE 2. Schematic digital illustrations of male left palp of species of the tribe Aelurillini.A Aelurillus v-insignitus (Clerck, 1757) (modified from Metzner 1999, fig. 42B, p. 196), ventral view (Aelurillina). B Pochyta spinosa Simon, 1901 (modified from Wesolowska &amp; Sztüts 2021, fig. 124, p. 31), ventral view (Thiratoscirtina). C Aphirape uncifera (MCTP 26426), ventral view (Freyina). D Sumampattus verae Marta, Figueiredo &amp; Rodrigues, 2023 (modified from Marta et al. 2023, fig. 2A, p. 4), ventral view (Freyina). E Hisukattus transversalis Galiano, 1987 (MCTP 39091), ventral view (Freyina). F Tullgrenella morenensis (Tullgren, 1905), ventral view (Freyina). Scale bars: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675792" httpUri="https://zenodo.org/record/10675792/files/figure.png" pageId="5" pageNumber="6">Figs 2D–2F</figureCitation>
) and pars pendula (see Galiano 1983: figs 6–7; 
<figureCitation id="18892A55FFCB4C016B495A75FF590771" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">Fig. 4F</figureCitation>
); 
<taxonomicName id="47B24D53FFCB4C016EF75A49FE0E0771" authority="Galiano, 1987" authorityName="Galiano" authorityYear="1987" box="[206,494,562,588]" class="Arachnida" family="Salticidae" genus="Hisukattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016EF75A49FEA60771" box="[206,326,562,588]" italics="true" pageId="5" pageNumber="6">Hisukattus</emphasis>
<bibRefCitation id="E4234B21FFCB4C016F775A49FE0E0771" author="Galiano, M. E." box="[334,494,562,588]" pageId="5" pageNumber="6" pagination="137 - 148" refId="ref42276" refString="Galiano, M. E. (1987) Descripcion de Hisukattus nuevo genero (Araneae, Salticidae). Revista de la Sociedad Entomologica Argentina, 44, 137 - 148." type="journal article" year="1987">Galiano, 1987</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="47B24D53FFCB4C016C115A49FC4B0771" authority="Peckham &amp; Peckham, 1896" authorityName="Peckham &amp; Peckham" authorityYear="1896" box="[552,939,562,588]" class="Arachnida" family="Salticidae" genus="Akela" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016C115A49FD880771" box="[552,616,562,588]" italics="true" pageId="5" pageNumber="6">Akela</emphasis>
Peckham &amp; Peckham, 1896
</taxonomicName>
by having proximal retrolateral lobe on the tegulum (see 
<bibRefCitation id="E4234B21FFCB4C016F085A2DFE2E074D" author="Galiano, M. E." box="[305,462,598,624]" pageId="5" pageNumber="6" pagination="1 - 4" refId="ref42309" refString="Galiano, M. E. (1999) Descripcion de Akela ruricola, nueva especies (Araneae, Salticidae). Physis, Revista de la Sociedad Argentina de Ciencias Naturales, Seccion C, 55, 1 - 4." type="journal article" year="1999">Galiano 1999</bibRefCitation>
: fig. 2E). Females of 
<taxonomicName id="47B24D53FFCB4C016CFC5A2DFCAD074D" authorityName="Mello-Leitao" authorityYear="1941" box="[709,845,598,624]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016CFC5A2DFCAD074D" box="[709,845,598,624]" italics="true" pageId="5" pageNumber="6">Tullgrenella</emphasis>
</taxonomicName>
resemble those of other freyines, such as: 
<taxonomicName id="47B24D53FFCB4C016B0D5A2DFA7C074D" authorityName="C. L. Kock" authorityYear="1850" box="[1332,1436,598,624]" class="Arachnida" family="Salticidae" genus="Aphirape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016B0D5A2DFA7C074D" box="[1332,1436,598,624]" italics="true" pageId="5" pageNumber="6">Aphirape</emphasis>
</taxonomicName>
and 
<taxonomicName id="47B24D53FFCB4C016EFE5A01FEBE07A9" authorityName="Galiano" authorityYear="1983" box="[199,350,634,660]" class="Arachnida" family="Salticidae" genus="Sumampattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016EFE5A01FEBE07A9" box="[199,350,634,660]" italics="true" pageId="5" pageNumber="6">Sumampattus</emphasis>
</taxonomicName>
by the window in ventral epigyne and the presence of posteriorly-opening coupling pocket (
<figureCitation id="18892A55FFCB4C016B535A01FEEC0785" captionStart="FIGURE 3" captionStartId="8.[151,250,1714,1738]" captionTargetBox="[170,1419,433,1683]" captionTargetId="figure-110@8.[151,1436,417,1691]" captionTargetPageId="8" captionText="FIGURE 3. Illustrations of the epigynum of species of the tribe Aelurillini. A Aelurillus v-insignitus (Clerck, 1757) (modified from Azarkina 2020, fig. 2, p. 3), dorsal view (Aelurillina). B Pochyta spinosa Simon, 1901 (adapted from Wesolowska &amp; Szűts 2021, fig. 136, p. 32), dorsal view (Thiratoscirtina). C Aphirape uncifera (MCTP 10891), dorsal view (Freyina). D Sumampattus verae Marta, Figueiredo &amp; Rodrigues, 2023 (modified from Marta et al. 2023, fig. 6B, p. 9], dorsal view (Freyina). E Hisukattus transversalis Galiano, 1987 (MCTP 39248), dorsal view (Freyina). F Tullgrenella morenensis (Tullgren, 1905), dorsal view (Freyina).Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675794" httpUri="https://zenodo.org/record/10675794/files/figure.png" pageId="5" pageNumber="6">Figs 3C–D, 3F</figureCitation>
); 
<taxonomicName id="47B24D53FFCB4C016F1A5AE5FE7B0785" authorityName="Galiano" authorityYear="1987" box="[291,411,670,696]" class="Arachnida" family="Salticidae" genus="Hisukattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016F1A5AE5FE7B0785" box="[291,411,670,696]" italics="true" pageId="5" pageNumber="6">Hisukattus</emphasis>
</taxonomicName>
and 
<taxonomicName id="47B24D53FFCB4C016FEE5AE5FDF70785" authorityName="Peckham &amp; Peckham" authorityYear="1896" box="[471,535,670,696]" class="Arachnida" family="Salticidae" genus="Akela" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016FEE5AE5FDF70785" box="[471,535,670,696]" italics="true" pageId="5" pageNumber="6">Akela</emphasis>
</taxonomicName>
by the copulatory ducts in concentric loops (
<figureCitation id="18892A55FFCB4C016A1F5AE5FB420785" box="[1062,1186,670,696]" captionStart="FIGURE 3" captionStartId="8.[151,250,1714,1738]" captionTargetBox="[170,1419,433,1683]" captionTargetId="figure-110@8.[151,1436,417,1691]" captionTargetPageId="8" captionText="FIGURE 3. Illustrations of the epigynum of species of the tribe Aelurillini. A Aelurillus v-insignitus (Clerck, 1757) (modified from Azarkina 2020, fig. 2, p. 3), dorsal view (Aelurillina). B Pochyta spinosa Simon, 1901 (adapted from Wesolowska &amp; Szűts 2021, fig. 136, p. 32), dorsal view (Thiratoscirtina). C Aphirape uncifera (MCTP 10891), dorsal view (Freyina). D Sumampattus verae Marta, Figueiredo &amp; Rodrigues, 2023 (modified from Marta et al. 2023, fig. 6B, p. 9], dorsal view (Freyina). E Hisukattus transversalis Galiano, 1987 (MCTP 39248), dorsal view (Freyina). F Tullgrenella morenensis (Tullgren, 1905), dorsal view (Freyina).Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675794" httpUri="https://zenodo.org/record/10675794/files/figure.png" pageId="5" pageNumber="6">Figs 3E–F</figureCitation>
). However, the genus 
<taxonomicName id="47B24D53FFCB4C016EAE5AB9FEFF07E1" authorityName="Mello-Leitao" authorityYear="1941" box="[151,287,706,732]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016EAE5AB9FEFF07E1" box="[151,287,706,732]" italics="true" pageId="5" pageNumber="6">Tullgrenella</emphasis>
</taxonomicName>
can be distinguished from the aforementioned genera by males having a cradle-shaped conductor (
<figureCitation id="18892A55FFCB4C016B535AB9FF55063D" captionStart="FIGURE 2" captionStartId="7.[151,250,1490,1514]" captionTargetBox="[151,1372,181,1459]" captionTargetId="figure-17@7.[151,1435,181,1465]" captionTargetPageId="7" captionText="FIGURE 2. Schematic digital illustrations of male left palp of species of the tribe Aelurillini.A Aelurillus v-insignitus (Clerck, 1757) (modified from Metzner 1999, fig. 42B, p. 196), ventral view (Aelurillina). B Pochyta spinosa Simon, 1901 (modified from Wesolowska &amp; Sztüts 2021, fig. 124, p. 31), ventral view (Thiratoscirtina). C Aphirape uncifera (MCTP 26426), ventral view (Freyina). D Sumampattus verae Marta, Figueiredo &amp; Rodrigues, 2023 (modified from Marta et al. 2023, fig. 2A, p. 4), ventral view (Freyina). E Hisukattus transversalis Galiano, 1987 (MCTP 39091), ventral view (Freyina). F Tullgrenella morenensis (Tullgren, 1905), ventral view (Freyina). Scale bars: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675792" httpUri="https://zenodo.org/record/10675792/files/figure.png" pageId="5" pageNumber="6">Figs 2F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016EF95A9DFEE2063D" box="[192,258,742,768]" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">4A–F</figureCitation>
) and a tegular membranous process upside to the embolar base (
<figureCitation id="18892A55FFCB4C016DE85A9DFBB7063D" box="[977,1111,742,768]" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">Figs 4A, 4F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A5B5A9DFB65063D" box="[1122,1157,742,768]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">8C</figureCitation>
), and females having the window on the ventral epigyne (
<figureCitation id="18892A55FFCB4C016C385B71FD600618" box="[513,640,778,805]" captionStart="FIGURE 5" captionStartId="10.[151,250,1750,1774]" captionTargetBox="[151,1436,417,1727]" captionTargetId="figure-109@10.[151,1436,417,1727]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electronic microscopy of epigynum, ventral view. A T. musica (MCN 55519), white arrow: septum. B T. aisenbergae Marta, sp. nov. (MCTP 12575), white arrow: septum. C T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135), white arrow: septum. D T. galianoae Marta, sp. nov. (MCTP 39711); red dashed line: window, red arrow: fundus membranous groove of the window. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675798" httpUri="https://zenodo.org/record/10675798/files/figure.png" pageId="5" pageNumber="6">Figs 5A–D</figureCitation>
) and monoplanar concentric copulatory ducts (
<figureCitation id="18892A55FFCB4C016AB65B71FAF00619" box="[1167,1296,778,804]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="5" pageNumber="6">Figs 7A–Q</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8A8655BFFCB4C016EFE5B56FEB20298" pageId="5" pageNumber="6" type="description">
<paragraph id="800D36D0FFCB4C016EFE5B56FDDA0229" blockId="5.[151,1437,417,2028]" pageId="5" pageNumber="6">
<emphasis id="B2C6EAC2FFCB4C016EFE5B56FEB9067A" bold="true" box="[199,345,813,839]" pageId="5" pageNumber="6">Description.</emphasis>
Length 4.0–9.0 mm. Dark, reddish–brown salticids (
<figureCitation id="18892A55FFCB4C016DA25B55FBF20675" box="[923,1042,814,840]" captionStart="FIGURE 1" captionStartId="6.[151,250,1833,1857]" captionTargetBox="[151,1435,522,1806]" captionTargetId="figure-150@6.[151,1435,522,1806]" captionTargetPageId="6" captionText="FIGURE 1. Photographs in vivo of Tullgrenella (morenensis group). A–C T. guayapae Galiano, 1970 (FCE-Ar 11212): A–B female, C male. D–F T. musica (Mello-Leitão, 1945) (FCE-Ar 9637): D–E female, F male. Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675790" httpUri="https://zenodo.org/record/10675790/files/figure.png" pageId="5" pageNumber="6">Figs 1A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A225B55FB910675" box="[1051,1137,814,840]" captionStart="FIGURE 20" captionStartId="28.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@28.[151,1435,181,1465]" captionTargetPageId="28" captionText="FIGURE 20. Photographs in vivo. A–D T. quadripunctata (Mello-Leitão, 1944):A–B. female (FCE-Ar 9395), C–D male (FCE- Ar 2284). E–H T. melanica (Mello-Leitão, 1941): E–F female (FCE-Ar 12259), G–H male (FCE-Ar 10505). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675844" httpUri="https://zenodo.org/record/10675844/files/figure.png" pageId="5" pageNumber="6">20A–H</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A435B55FB280675" box="[1146,1224,814,840]" captionStart="FIGURE 32" captionStartId="42.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@42.[151,1435,181,1465]" captionTargetPageId="42" captionText="FIGURE 32. Photographs in vivo. A–D T. serrana Galiano, 1970:A–B female (FCE-Ar 11149), C–D male (FCE-Ar 9538). E–F female of T. legalissima Marta &amp; Hagopián, sp. nov. (FCE-Ar 12155). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675884" httpUri="https://zenodo.org/record/10675884/files/figure.png" pageId="5" pageNumber="6">32A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016AE85B55FAC40674" box="[1233,1316,814,841]" captionStart="FIGURE 45" captionStartId="58.[151,250,1689,1713]" captionTargetBox="[151,1435,381,1665]" captionTargetId="figure-102@58.[151,1435,381,1665]" captionTargetPageId="58" captionText="FIGURE 45. Photographs in vivo of T. peniaflorensis Galiano, 1970. A–F male. Photographs by Asiel Olivares." figureDoi="http://doi.org/10.5281/zenodo.10675916" httpUri="https://zenodo.org/record/10675916/files/figure.png" pageId="5" pageNumber="6">45A–F</figureCitation>
). Carapace reddish-brown, cephalic region smooth and dark, thoracic region with slope and two perpendicular bands of white setae to each other on thoracic slope (
<figureCitation id="18892A55FFCB4C016C7C5B0DFD4406AD" box="[581,676,886,912]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8A</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016C885B0DFD3706AD" box="[689,727,886,912]" captionStart="FIGURE 9" captionStartId="15.[151,250,1490,1514]" captionTargetBox="[158,1431,181,1460]" captionTargetId="figure-17@15.[151,1435,181,1465]" captionTargetPageId="15" captionText="FIGURE 9. Female of T. morenensis (Tullgren, 1905) (MCN 19940). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675810" httpUri="https://zenodo.org/record/10675810/files/figure.png" pageId="5" pageNumber="6">9A</figureCitation>
). Anterior half of thoracic region at the same level as cephalic region. Chelicerae robust and vertical in both sexes (not dimorphic), promargin with two teeth (proximal reduced), and retromargin with one tooth (
<figureCitation id="18892A55FFCB4C016FC45BC5FD6A06E5" box="[509,650,958,984]" captionStart="FIGURE 44" captionStartId="57.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@57.[151,1435,181,1465]" captionTargetPageId="57" captionText="FIGURE 44. Scanning electronic microscopy. A right chelicera of T. yungae Galiano, 1970 (MCN 44133), prolateral view, arrow: distal and proximal teeth. B left chelicera of T. musica (Mello-Leitão, 1945) (MCTP 55370), retrolateral view, arrow: tooth. C claw of T. quadripunctata (Mello-Leitão, 1944) (MCN 26658), arrow: claw teeth and subungueal tufts. D claw of T. yungae Galiano, 1970 (MCTP 39711), arrow: claw teeth and subungueal tufts." figureDoi="http://doi.org/10.5281/zenodo.10675914" httpUri="https://zenodo.org/record/10675914/files/figure.png" pageId="5" pageNumber="6">Figs 44A–B</figureCitation>
). Legs brown and yellow, with brown spots on distal portion and tarsus always lighter than other articles of legs. Legs formula: 4312/ 
<quantity id="474A9B35FFCB4C016D5C5B99FC5D06C1" box="[869,957,994,1020]" metricMagnitude="2" metricUnit="m" metricValue="1.049528" pageId="5" pageNumber="6" unit="in" value="4132.0">4132 in</quantity>
both sexes. Tarsal claws small, with five inferior teeth, not surpassing subungueal sulci (
<figureCitation id="18892A55FFCB4C016C9F5C7DFCD4011D" box="[678,820,1030,1056]" captionStart="FIGURE 44" captionStartId="57.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@57.[151,1435,181,1465]" captionTargetPageId="57" captionText="FIGURE 44. Scanning electronic microscopy. A right chelicera of T. yungae Galiano, 1970 (MCN 44133), prolateral view, arrow: distal and proximal teeth. B left chelicera of T. musica (Mello-Leitão, 1945) (MCTP 55370), retrolateral view, arrow: tooth. C claw of T. quadripunctata (Mello-Leitão, 1944) (MCN 26658), arrow: claw teeth and subungueal tufts. D claw of T. yungae Galiano, 1970 (MCTP 39711), arrow: claw teeth and subungueal tufts." figureDoi="http://doi.org/10.5281/zenodo.10675914" httpUri="https://zenodo.org/record/10675914/files/figure.png" pageId="5" pageNumber="6">Figs 44C–D</figureCitation>
), and the distal one larger in females. Claw tufts poorly developed (
<figureCitation id="18892A55FFCB4C016F245C51FE4D0179" box="[285,429,1066,1092]" captionStart="FIGURE 44" captionStartId="57.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@57.[151,1435,181,1465]" captionTargetPageId="57" captionText="FIGURE 44. Scanning electronic microscopy. A right chelicera of T. yungae Galiano, 1970 (MCN 44133), prolateral view, arrow: distal and proximal teeth. B left chelicera of T. musica (Mello-Leitão, 1945) (MCTP 55370), retrolateral view, arrow: tooth. C claw of T. quadripunctata (Mello-Leitão, 1944) (MCN 26658), arrow: claw teeth and subungueal tufts. D claw of T. yungae Galiano, 1970 (MCTP 39711), arrow: claw teeth and subungueal tufts." figureDoi="http://doi.org/10.5281/zenodo.10675914" httpUri="https://zenodo.org/record/10675914/files/figure.png" pageId="5" pageNumber="6">Figs 44C–D</figureCitation>
, see arrow). Abdomen: dark or brownish–yellow, with white spots on dorsal posterior abdomen (
<figureCitation id="18892A55FFCB4C016F295C35FE900155" box="[272,368,1102,1128]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8A</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016F465C35FE510155" box="[383,433,1102,1128]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10A</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016FF95C35FDEE0155" box="[448,526,1102,1128]" captionStart="FIGURE 13" captionStartId="19.[151,250,1761,1785]" captionTargetBox="[152,1416,453,1730]" captionTargetId="figure-110@19.[151,1435,453,1737]" captionTargetPageId="19" captionText="FIGURE 13. Female of T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675822" httpUri="https://zenodo.org/record/10675822/files/figure.png" pageId="5" pageNumber="6">13A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016C245C35FD920155" box="[541,626,1102,1128]" captionStart="FIGURE 20" captionStartId="28.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@28.[151,1435,181,1465]" captionTargetPageId="28" captionText="FIGURE 20. Photographs in vivo. A–D T. quadripunctata (Mello-Leitão, 1944):A–B. female (FCE-Ar 9395), C–D male (FCE- Ar 2284). E–H T. melanica (Mello-Leitão, 1941): E–F female (FCE-Ar 12259), G–H male (FCE-Ar 10505). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675844" httpUri="https://zenodo.org/record/10675844/files/figure.png" pageId="5" pageNumber="6">20A–H</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016CB95C35FD530155" box="[640,691,1102,1128]" captionStart="FIGURE 23" captionStartId="31.[151,250,1979,2003]" captionTargetBox="[151,1435,1060,1948]" captionTargetId="figure-311@31.[151,1435,1049,1956]" captionTargetPageId="31" captionText="FIGURE 23. Male of T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view. B same, ventral view. C left palp, ventral view. D same, retrolateral view, red arrow: ventral projection on tibia. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675852" httpUri="https://zenodo.org/record/10675852/files/figure.png" pageId="5" pageNumber="6">23A</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016CFB5C35FCF00155" box="[706,784,1102,1128]" captionStart="FIGURE 32" captionStartId="42.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@42.[151,1435,181,1465]" captionTargetPageId="42" captionText="FIGURE 32. Photographs in vivo. A–D T. serrana Galiano, 1970:A–B female (FCE-Ar 11149), C–D male (FCE-Ar 9538). E–F female of T. legalissima Marta &amp; Hagopián, sp. nov. (FCE-Ar 12155). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675884" httpUri="https://zenodo.org/record/10675884/files/figure.png" pageId="5" pageNumber="6">32A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D275C35FCB30155" box="[798,851,1102,1128]" captionStart="FIGURE 33" captionStartId="43.[151,250,1779,1803]" captionTargetBox="[155,1435,855,1756]" captionTargetId="figure-271@43.[151,1435,849,1756]" captionTargetPageId="43" captionText="FIGURE 33. Male of T. serrana Galiano, 1970 (MCN 46708). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675888" httpUri="https://zenodo.org/record/10675888/files/figure.png" pageId="5" pageNumber="6">33A</figureCitation>
); venter usually with a median longitudinal brown stripe of irregular contours (
<figureCitation id="18892A55FFCB4C016FF55C09FDC401B1" box="[460,548,1138,1164]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8B</figureCitation>
– 
<figureCitation id="18892A55FFCB4C016C0C5C09FD8401B1" box="[565,612,1138,1164]" captionStart="FIGURE 17" captionStartId="24.[151,250,1689,1713]" captionTargetBox="[156,1435,381,1656]" captionTargetId="figure-100@24.[151,1435,381,1665]" captionTargetPageId="24" captionText="FIGURE 17. Female of T. corrugata Galiano, 1981 (MACN-Ar 7206). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675834" httpUri="https://zenodo.org/record/10675834/files/figure.png" pageId="5" pageNumber="6">17B</figureCitation>
); sides with white longitudinal band at tip of anterior abdomen. Coloration of spinnerets as in legs. Male palp: femur macrosetae d1–1–1. Femur with ventral projection (
<figureCitation id="18892A55FFCB4C016AE25CEDFA8A018D" box="[1243,1386,1174,1200]" captionStart="FIGURE 43" captionStartId="56.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@56.[151,1435,181,1465]" captionTargetPageId="56" captionText="FIGURE 43. Scanning electronic microscopy of male palp, ventral view. A zoom of RTA of T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531), white arrow: hook between RTA and prolatero-dorsal tibial apophysis, red arrow: lyriform organ. B zoom of RTA of T. serrana Galiano, 1970 (MCTP 31126), white arrow: hook between RTA and prolatero-dorsal tibial apophysis. C femur ventral projection of T. yungae Galiano, 1970 (MCN 44133), white arrow: projection. D femur ventral projection of T. aisenbergae Marta sp. nov. (IBSP 118477), white arrow: projection." figureDoi="http://doi.org/10.5281/zenodo.10675912" httpUri="https://zenodo.org/record/10675912/files/figure.png" pageId="5" pageNumber="6">Figs 43C–D</figureCitation>
, see arrow). Tibia with modest RTA with ventral sinuose or acute prong (
<figureCitation id="18892A55FFCB4C016DAF5CC1FC1301E9" box="[918,1011,1210,1236]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8D</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016DC65CC1FBD101E9" box="[1023,1073,1210,1236]" captionStart="FIGURE 23" captionStartId="31.[151,250,1979,2003]" captionTargetBox="[151,1435,1060,1948]" captionTargetId="figure-311@31.[151,1435,1049,1956]" captionTargetPageId="31" captionText="FIGURE 23. Male of T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view. B same, ventral view. C left palp, ventral view. D same, retrolateral view, red arrow: ventral projection on tibia. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675852" httpUri="https://zenodo.org/record/10675852/files/figure.png" pageId="5" pageNumber="6">23D</figureCitation>
) and usually incipient hook-like projection between the RTA and prolatero-dorsal tibial apophysis (
<figureCitation id="18892A55FFCB4C016D455CA5FC3901C5" box="[892,985,1246,1272]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8D</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016DDD5CA5FBF701C5" box="[996,1047,1246,1272]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10D</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A1B5CA5FBB501C5" box="[1058,1109,1246,1272]" captionStart="FIGURE 14" captionStartId="21.[151,250,1419,1443]" captionTargetBox="[155,1427,501,1391]" captionTargetId="figure-120@21.[151,1435,489,1396]" captionTargetPageId="21" captionText="FIGURE 14. Male of T. musica (Mello-Leitão, 1945) (MCN 53371). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675826" httpUri="https://zenodo.org/record/10675826/files/figure.png" pageId="5" pageNumber="6">14D</figureCitation>
, see red arrow). Usually with a small prolatero-dorsal tibial apophysis (
<figureCitation id="18892A55FFCB4C016C4C5D79FD340021" box="[629,724,1282,1308]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8D</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016CDA5D79FCF70021" box="[739,791,1282,1308]" captionStart="FIGURE 14" captionStartId="21.[151,250,1419,1443]" captionTargetBox="[155,1427,501,1391]" captionTargetId="figure-120@21.[151,1435,489,1396]" captionTargetPageId="21" captionText="FIGURE 14. Male of T. musica (Mello-Leitão, 1945) (MCN 53371). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675826" httpUri="https://zenodo.org/record/10675826/files/figure.png" pageId="5" pageNumber="6">14D</figureCitation>
). Cymbium apex suboval or sickle-shaped (
<figureCitation id="18892A55FFCB4C016B285D79FA710021" box="[1297,1425,1282,1308]" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">Figs 4A–F</figureCitation>
). Tegulum oblique in relation to cymbium apex and central depression (
<figureCitation id="18892A55FFCB4C016D8E5D5DFBC2007D" box="[951,1058,1318,1344]" captionStart="FIGURE 14" captionStartId="21.[151,250,1419,1443]" captionTargetBox="[155,1427,501,1391]" captionTargetId="figure-120@21.[151,1435,489,1396]" captionTargetPageId="21" captionText="FIGURE 14. Male of T. musica (Mello-Leitão, 1945) (MCN 53371). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675826" httpUri="https://zenodo.org/record/10675826/files/figure.png" pageId="5" pageNumber="6">Figs 14C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A095D5DFB80007D" box="[1072,1120,1318,1344]" captionStart="FIGURE 21" captionStartId="29.[151,250,1112,1136]" captionTargetBox="[156,1406,197,1084]" captionTargetId="figure-17@29.[151,1435,181,1088]" captionTargetPageId="29" captionText="FIGURE 21. Male of T. quadripunctata (Mello-Leitão, 1944). A habitus (MCN 14882), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp (IBSP 123127), ventral view, red arrow: tegulum central depression. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675846" httpUri="https://zenodo.org/record/10675846/files/figure.png" pageId="5" pageNumber="6">21C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A575D5DFB7F007C" box="[1134,1183,1318,1345]" captionStart="FIGURE 25" captionStartId="33.[151,250,1921,1945]" captionTargetBox="[176,1406,994,1898]" captionTargetId="figure-342@33.[172,1416,991,1898]" captionTargetPageId="33" captionText="FIGURE 25. Male of T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675858" httpUri="https://zenodo.org/record/10675858/files/figure.png" pageId="5" pageNumber="6">25C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A955D5DFB3D007D" box="[1196,1245,1318,1344]" captionStart="FIGURE 29" captionStartId="38.[151,250,1548,1572]" captionTargetBox="[156,1420,625,1518]" captionTargetId="figure-166@38.[151,1435,617,1524]" captionTargetPageId="38" captionText="FIGURE 29. Male of T. yungae Galiano, 1970 (MCN 44133). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis, white dashed line: RTA. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675874" httpUri="https://zenodo.org/record/10675874/files/figure.png" pageId="5" pageNumber="6">29C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016AD35D5DFAFB007D" box="[1258,1307,1318,1344]" captionStart="FIGURE 33" captionStartId="43.[151,250,1779,1803]" captionTargetBox="[155,1435,855,1756]" captionTargetId="figure-271@43.[151,1435,849,1756]" captionTargetPageId="43" captionText="FIGURE 33. Male of T. serrana Galiano, 1970 (MCN 46708). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675888" httpUri="https://zenodo.org/record/10675888/files/figure.png" pageId="5" pageNumber="6">33C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B115D5DFABA007D" box="[1320,1370,1318,1344]" captionStart="FIGURE 40" captionStartId="52.[151,250,1112,1136]" captionTargetBox="[155,1418,203,1083]" captionTargetId="figure-20@52.[151,1435,181,1088]" captionTargetPageId="52" captionText="FIGURE 40. Male of T. galianoae Marta, sp. nov. (IBSP 118546). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolaterodorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675904" httpUri="https://zenodo.org/record/10675904/files/figure.png" pageId="5" pageNumber="6">40C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B5E5D5DFA77007D" box="[1383,1431,1318,1344]" captionStart="FIGURE 48" captionStartId="61.[151,250,1203,1227]" captionTargetBox="[155,1435,279,1175]" captionTargetId="figure-41@61.[151,1435,273,1180]" captionTargetPageId="61" captionText="FIGURE 48. Male of T. selenita Galiano, 1970 (MACN-Ar 6250). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675924" httpUri="https://zenodo.org/record/10675924/files/figure.png" pageId="5" pageNumber="6">48C</figureCitation>
, see red arrow) with inverted U-shaped spermophore visible (
<figureCitation id="18892A55FFCB4C016D035D31FC750059" box="[826,917,1354,1380]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D995D31FC310059" box="[928,977,1354,1380]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016DE55D31FBED0059" box="[988,1037,1354,1380]" captionStart="FIGURE 12" captionStartId="18.[151,250,1869,1893]" captionTargetBox="[156,1435,953,1842]" captionTargetId="figure-286@18.[151,1435,939,1846]" captionTargetPageId="18" captionText="FIGURE 12. Male of T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675816" httpUri="https://zenodo.org/record/10675816/files/figure.png" pageId="5" pageNumber="6">12C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A215D31FBAA0059" box="[1048,1098,1354,1380]" captionStart="FIGURE 23" captionStartId="31.[151,250,1979,2003]" captionTargetBox="[151,1435,1060,1948]" captionTargetId="figure-311@31.[151,1435,1049,1956]" captionTargetPageId="31" captionText="FIGURE 23. Male of T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view. B same, ventral view. C left palp, ventral view. D same, retrolateral view, red arrow: ventral projection on tibia. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675852" httpUri="https://zenodo.org/record/10675852/files/figure.png" pageId="5" pageNumber="6">23C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A6D5D31FB650059" box="[1108,1157,1354,1380]" captionStart="FIGURE 33" captionStartId="43.[151,250,1779,1803]" captionTargetBox="[155,1435,855,1756]" captionTargetId="figure-271@43.[151,1435,849,1756]" captionTargetPageId="43" captionText="FIGURE 33. Male of T. serrana Galiano, 1970 (MCN 46708). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675888" httpUri="https://zenodo.org/record/10675888/files/figure.png" pageId="5" pageNumber="6">33C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016AA95D31FB220059" box="[1168,1218,1354,1380]" captionStart="FIGURE 40" captionStartId="52.[151,250,1112,1136]" captionTargetBox="[155,1418,203,1083]" captionTargetId="figure-20@52.[151,1435,181,1088]" captionTargetPageId="52" captionText="FIGURE 40. Male of T. galianoae Marta, sp. nov. (IBSP 118546). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolaterodorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675904" httpUri="https://zenodo.org/record/10675904/files/figure.png" pageId="5" pageNumber="6">40C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016AF55D31FB1D0059" box="[1228,1277,1354,1380]" captionStart="FIGURE 46" captionStartId="59.[151,250,1563,1587]" captionTargetBox="[151,1435,633,1540]" captionTargetId="figure-211@59.[151,1435,633,1540]" captionTargetPageId="59" captionText="FIGURE 46. Male of T. peniaflorensis Galiano, 1970 (FNAG36 1463). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: ventral hook between RTA and prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675918" httpUri="https://zenodo.org/record/10675918/files/figure.png" pageId="5" pageNumber="6">46C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B315D31FADB0059" box="[1288,1339,1354,1380]" captionStart="FIGURE 48" captionStartId="61.[151,250,1203,1227]" captionTargetBox="[155,1435,279,1175]" captionTargetId="figure-41@61.[151,1435,273,1180]" captionTargetPageId="61" captionText="FIGURE 48. Male of T. selenita Galiano, 1970 (MACN-Ar 6250). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675924" httpUri="https://zenodo.org/record/10675924/files/figure.png" pageId="5" pageNumber="6">48C</figureCitation>
); tegular distal portion with developed proximal retrolateral lobe of tegulum (
<figureCitation id="18892A55FFCB4C016D995D15FC1C00B5" box="[928,1020,1390,1416]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">Figs 8C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A335D15FBDB00B5" box="[1034,1083,1390,1416]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A715D15FB9900B5" box="[1096,1145,1390,1416]" captionStart="FIGURE 12" captionStartId="18.[151,250,1869,1893]" captionTargetBox="[156,1435,953,1842]" captionTargetId="figure-286@18.[151,1435,939,1846]" captionTargetPageId="18" captionText="FIGURE 12. Male of T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675816" httpUri="https://zenodo.org/record/10675816/files/figure.png" pageId="5" pageNumber="6">12C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016ABF5D15FB5800B5" box="[1158,1208,1390,1416]" captionStart="FIGURE 14" captionStartId="21.[151,250,1419,1443]" captionTargetBox="[155,1427,501,1391]" captionTargetId="figure-120@21.[151,1435,489,1396]" captionTargetPageId="21" captionText="FIGURE 14. Male of T. musica (Mello-Leitão, 1945) (MCN 53371). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675826" httpUri="https://zenodo.org/record/10675826/files/figure.png" pageId="5" pageNumber="6">14C</figureCitation>
). Tegulum with two projections: membranous process of tegulum (globose or elongated) upside to embolar base (
<figureCitation id="18892A55FFCB4C016A9B5DE9FACD0091" box="[1186,1325,1426,1452]" captionStart="FIGURE 6" captionStartId="11.[151,250,1490,1514]" captionTargetBox="[158,1420,182,1465]" captionTargetId="figure-17@11.[151,1435,181,1465]" captionTargetPageId="11" captionText="FIGURE 6. Expanded male left palp of Tullgrenella.A–C T. musica (Mello-Leitão, 1945) (MCN 53371), ventral (A), retrolateral (B) and prolateral (C) view. D–F T. lunata (Mello-Leitão, 1944) (MACN-Ar 42300), ventral view (D), retrolateral (E) and prolateral (F) view. G–I T. serrana Galiano, 1970 (MACN-Ar 6262), ventral (G), retrolateral (H) and prolateral (I) view. Scale bars: 0.50 mm. Black arrow: rotation direction; arrows in other colors refer to the structures with the same annotated colors." figureDoi="http://doi.org/10.5281/zenodo.10675802" httpUri="https://zenodo.org/record/10675802/files/figure.png" pageId="5" pageNumber="6">Figs 6A, 6C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B015DE9FABB0091" box="[1336,1371,1426,1452]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">8C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B5E5DE9FA770091" box="[1383,1431,1426,1452]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016EAE5DCDFF2900ED" box="[151,201,1462,1488]" captionStart="FIGURE 38" captionStartId="49.[151,250,1888,1912]" captionTargetBox="[155,1413,977,1859]" captionTargetId="figure-286@49.[151,1435,957,1864]" captionTargetPageId="49" captionText="FIGURE 38. Male of T. sciosciae Marta &amp; Bustamante, sp. nov. A habitus (MCTP 10652), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp (MCTP 31075), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675900" httpUri="https://zenodo.org/record/10675900/files/figure.png" pageId="5" pageNumber="6">38C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016EED5DCDFEE500ED" box="[212,261,1462,1488]" captionStart="FIGURE 46" captionStartId="59.[151,250,1563,1587]" captionTargetBox="[151,1435,633,1540]" captionTargetId="figure-211@59.[151,1435,633,1540]" captionTargetPageId="59" captionText="FIGURE 46. Male of T. peniaflorensis Galiano, 1970 (FNAG36 1463). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: ventral hook between RTA and prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675918" httpUri="https://zenodo.org/record/10675918/files/figure.png" pageId="5" pageNumber="6">46C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016F285DCDFEA200ED" box="[273,322,1462,1488]" captionStart="FIGURE 48" captionStartId="61.[151,250,1203,1227]" captionTargetBox="[155,1435,279,1175]" captionTargetId="figure-41@61.[151,1435,273,1180]" captionTargetPageId="61" captionText="FIGURE 48. Male of T. selenita Galiano, 1970 (MACN-Ar 6250). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675924" httpUri="https://zenodo.org/record/10675924/files/figure.png" pageId="5" pageNumber="6">48C</figureCitation>
, see purple arrow) and one conductor (
<figureCitation id="18892A55FFCB4C016CC05DCDFCAF00ED" box="[761,847,1462,1488]" captionStart="FIGURE 2" captionStartId="7.[151,250,1490,1514]" captionTargetBox="[151,1372,181,1459]" captionTargetId="figure-17@7.[151,1435,181,1465]" captionTargetPageId="7" captionText="FIGURE 2. Schematic digital illustrations of male left palp of species of the tribe Aelurillini.A Aelurillus v-insignitus (Clerck, 1757) (modified from Metzner 1999, fig. 42B, p. 196), ventral view (Aelurillina). B Pochyta spinosa Simon, 1901 (modified from Wesolowska &amp; Sztüts 2021, fig. 124, p. 31), ventral view (Thiratoscirtina). C Aphirape uncifera (MCTP 26426), ventral view (Freyina). D Sumampattus verae Marta, Figueiredo &amp; Rodrigues, 2023 (modified from Marta et al. 2023, fig. 2A, p. 4), ventral view (Freyina). E Hisukattus transversalis Galiano, 1987 (MCTP 39091), ventral view (Freyina). F Tullgrenella morenensis (Tullgren, 1905), ventral view (Freyina). Scale bars: 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675792" httpUri="https://zenodo.org/record/10675792/files/figure.png" pageId="5" pageNumber="6">Figs 2F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D625DCDFC7C00ED" box="[859,924,1462,1488]" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">4A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D9E5DCDFC1A00ED" box="[935,1018,1462,1488]" captionStart="FIGURE 6" captionStartId="11.[151,250,1490,1514]" captionTargetBox="[158,1420,182,1465]" captionTargetId="figure-17@11.[151,1435,181,1465]" captionTargetPageId="11" captionText="FIGURE 6. Expanded male left palp of Tullgrenella.A–C T. musica (Mello-Leitão, 1945) (MCN 53371), ventral (A), retrolateral (B) and prolateral (C) view. D–F T. lunata (Mello-Leitão, 1944) (MACN-Ar 42300), ventral view (D), retrolateral (E) and prolateral (F) view. G–I T. serrana Galiano, 1970 (MACN-Ar 6262), ventral (G), retrolateral (H) and prolateral (I) view. Scale bars: 0.50 mm. Black arrow: rotation direction; arrows in other colors refer to the structures with the same annotated colors." figureDoi="http://doi.org/10.5281/zenodo.10675802" httpUri="https://zenodo.org/record/10675802/files/figure.png" pageId="5" pageNumber="6">6A, 6C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A3E5DCDFBC900ED" box="[1031,1065,1462,1488]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">8C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A0F5DCDFB8600ED" box="[1078,1126,1462,1488]" captionStart="FIGURE 10" captionStartId="16.[151,250,1203,1227]" captionTargetBox="[155,1435,303,1175]" captionTargetId="figure-60@16.[151,1435,273,1180]" captionTargetPageId="16" captionText="FIGURE 10. Male of T. guayapae Galiano, 1970.A habitus (MCN 21128), dorsal view. B same, ventral view. C left palp (MCN 55543), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675812" httpUri="https://zenodo.org/record/10675812/files/figure.png" pageId="5" pageNumber="6">10C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A4A5DCDFB4300ED" box="[1139,1187,1462,1488]" captionStart="FIGURE 21" captionStartId="29.[151,250,1112,1136]" captionTargetBox="[156,1406,197,1084]" captionTargetId="figure-17@29.[151,1435,181,1088]" captionTargetPageId="29" captionText="FIGURE 21. Male of T. quadripunctata (Mello-Leitão, 1944). A habitus (MCN 14882), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp (IBSP 123127), ventral view, red arrow: tegulum central depression. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675846" httpUri="https://zenodo.org/record/10675846/files/figure.png" pageId="5" pageNumber="6">21C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A895DCDFB0100ED" box="[1200,1249,1462,1488]" captionStart="FIGURE 23" captionStartId="31.[151,250,1979,2003]" captionTargetBox="[151,1435,1060,1948]" captionTargetId="figure-311@31.[151,1435,1049,1956]" captionTargetPageId="31" captionText="FIGURE 23. Male of T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 23782). A habitus, dorsal view. B same, ventral view. C left palp, ventral view. D same, retrolateral view, red arrow: ventral projection on tibia. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675852" httpUri="https://zenodo.org/record/10675852/files/figure.png" pageId="5" pageNumber="6">23C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016AD45DCDFAFE00ED" box="[1261,1310,1462,1488]" captionStart="FIGURE 27" captionStartId="36.[151,250,1112,1136]" captionTargetBox="[155,1421,196,1084]" captionTargetId="figure-20@36.[151,1435,181,1088]" captionTargetPageId="36" captionText="FIGURE 27. Male of T. melanica (Mello-Leitão, 1941). A habitus (MACN-Ar 6243), dorsal view. B same, ventral view. C left palp (MCN 23782), ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, white dashed line: RTA. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675866" httpUri="https://zenodo.org/record/10675866/files/figure.png" pageId="5" pageNumber="6">27C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B135DCDFABB00ED" box="[1322,1371,1462,1488]" captionStart="FIGURE 33" captionStartId="43.[151,250,1779,1803]" captionTargetBox="[155,1435,855,1756]" captionTargetId="figure-271@43.[151,1435,849,1756]" captionTargetPageId="43" captionText="FIGURE 33. Male of T. serrana Galiano, 1970 (MCN 46708). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675888" httpUri="https://zenodo.org/record/10675888/files/figure.png" pageId="5" pageNumber="6">33C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016B5E5DCDFA7800ED" box="[1383,1432,1462,1488]" captionStart="FIGURE 38" captionStartId="49.[151,250,1888,1912]" captionTargetBox="[155,1413,977,1859]" captionTargetId="figure-286@49.[151,1435,957,1864]" captionTargetPageId="49" captionText="FIGURE 38. Male of T. sciosciae Marta &amp; Bustamante, sp. nov. A habitus (MCTP 10652), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp (MCTP 31075), ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675900" httpUri="https://zenodo.org/record/10675900/files/figure.png" pageId="5" pageNumber="6">38C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016EAE5DA1FF2800C9" box="[151,200,1498,1524]" captionStart="FIGURE 46" captionStartId="59.[151,250,1563,1587]" captionTargetBox="[151,1435,633,1540]" captionTargetId="figure-211@59.[151,1435,633,1540]" captionTargetPageId="59" captionText="FIGURE 46. Male of T. peniaflorensis Galiano, 1970 (FNAG36 1463). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: ventral hook between RTA and prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675918" httpUri="https://zenodo.org/record/10675918/files/figure.png" pageId="5" pageNumber="6">46C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016EED5DA1FEE600C9" box="[212,262,1498,1524]" captionStart="FIGURE 48" captionStartId="61.[151,250,1203,1227]" captionTargetBox="[155,1435,279,1175]" captionTargetId="figure-41@61.[151,1435,273,1180]" captionTargetPageId="61" captionText="FIGURE 48. Male of T. selenita Galiano, 1970 (MACN-Ar 6250). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675924" httpUri="https://zenodo.org/record/10675924/files/figure.png" pageId="5" pageNumber="6">48C</figureCitation>
). Embolar base projecting dorso-prolaterally (
<figureCitation id="18892A55FFCB4C016D315DA1FC5300C9" box="[776,947,1498,1524]" captionStart="FIGURE 6" captionStartId="11.[151,250,1490,1514]" captionTargetBox="[158,1420,182,1465]" captionTargetId="figure-17@11.[151,1435,181,1465]" captionTargetPageId="11" captionText="FIGURE 6. Expanded male left palp of Tullgrenella.A–C T. musica (Mello-Leitão, 1945) (MCN 53371), ventral (A), retrolateral (B) and prolateral (C) view. D–F T. lunata (Mello-Leitão, 1944) (MACN-Ar 42300), ventral view (D), retrolateral (E) and prolateral (F) view. G–I T. serrana Galiano, 1970 (MACN-Ar 6262), ventral (G), retrolateral (H) and prolateral (I) view. Scale bars: 0.50 mm. Black arrow: rotation direction; arrows in other colors refer to the structures with the same annotated colors." figureDoi="http://doi.org/10.5281/zenodo.10675802" httpUri="https://zenodo.org/record/10675802/files/figure.png" pageId="5" pageNumber="6">Figs 6C, 6F, 6I</figureCitation>
). Embolus sclerotized and slender, embolus way starting at 8–9 o’ clock (left palp; 
<figureCitation id="18892A55FFCB4C016C5D5D85FD3F0325" box="[612,735,1534,1560]" captionStart="FIGURE 4" captionStartId="9.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-17@9.[151,1435,181,1465]" captionTargetPageId="9" captionText="FIGURE 4. Scanning electronic microscopy of male left palp, ventral view.A T. musica (Mello-Leitão, 1945) (MCN 53370). B T. quadripunctata (Mello-Leitão, 1944) (MCTP 16531). C T. yungae Galiano, 1970 (MCN 44133). D T. serrana Galiano, 1970 (MCTP 31126). E T. aisenbergae Marta, sp. nov. (IBSP 118477). F T. peniaflorensis Galiano, 1970 (FNAG36 1464). Scale bars: 0.50 mm. Colored homology: purple and purple arrow (tegular membranous process), blue (embolar base), green (conductor), magenta (tegulum), orange (embolus), red (pars pendula)." figureDoi="http://doi.org/10.5281/zenodo.10675796" httpUri="https://zenodo.org/record/10675796/files/figure.png" pageId="5" pageNumber="6">Figs 4A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016CD45D85FCC90325" box="[749,809,1534,1560]" captionStart="FIGURE 6" captionStartId="11.[151,250,1490,1514]" captionTargetBox="[158,1420,182,1465]" captionTargetId="figure-17@11.[151,1435,181,1465]" captionTargetPageId="11" captionText="FIGURE 6. Expanded male left palp of Tullgrenella.A–C T. musica (Mello-Leitão, 1945) (MCN 53371), ventral (A), retrolateral (B) and prolateral (C) view. D–F T. lunata (Mello-Leitão, 1944) (MACN-Ar 42300), ventral view (D), retrolateral (E) and prolateral (F) view. G–I T. serrana Galiano, 1970 (MACN-Ar 6262), ventral (G), retrolateral (H) and prolateral (I) view. Scale bars: 0.50 mm. Black arrow: rotation direction; arrows in other colors refer to the structures with the same annotated colors." figureDoi="http://doi.org/10.5281/zenodo.10675802" httpUri="https://zenodo.org/record/10675802/files/figure.png" pageId="5" pageNumber="6">6A–I</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D015D85FCBB0325" box="[824,859,1534,1560]" captionStart="FIGURE 8" captionStartId="14.[151,250,1851,1875]" captionTargetBox="[155,1423,929,1827]" captionTargetId="figure-290@14.[151,1435,921,1828]" captionTargetPageId="14" captionText="FIGURE 8. Male of T. morenensis (Tullgren, 1905) (MACN-Ar 42165). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line:prolaterodorsal tibial apophysis. Scale bars: A–B 2 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675806" httpUri="https://zenodo.org/record/10675806/files/figure.png" pageId="5" pageNumber="6">8C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D535D85FC7B0325" box="[874,923,1534,1560]" captionStart="FIGURE 21" captionStartId="29.[151,250,1112,1136]" captionTargetBox="[156,1406,197,1084]" captionTargetId="figure-17@29.[151,1435,181,1088]" captionTargetPageId="29" captionText="FIGURE 21. Male of T. quadripunctata (Mello-Leitão, 1944). A habitus (MCN 14882), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp (IBSP 123127), ventral view, red arrow: tegulum central depression. D same, retrolateral view. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675846" httpUri="https://zenodo.org/record/10675846/files/figure.png" pageId="5" pageNumber="6">21C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D925D85FC3C0325" box="[939,988,1534,1560]" captionStart="FIGURE 27" captionStartId="36.[151,250,1112,1136]" captionTargetBox="[155,1421,196,1084]" captionTargetId="figure-20@36.[151,1435,181,1088]" captionTargetPageId="36" captionText="FIGURE 27. Male of T. melanica (Mello-Leitão, 1941). A habitus (MACN-Ar 6243), dorsal view. B same, ventral view. C left palp (MCN 23782), ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, white dashed line: RTA. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675866" httpUri="https://zenodo.org/record/10675866/files/figure.png" pageId="5" pageNumber="6">27C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016DD25D85FBFC0325" box="[1003,1052,1534,1560]" captionStart="FIGURE 33" captionStartId="43.[151,250,1779,1803]" captionTargetBox="[155,1435,855,1756]" captionTargetId="figure-271@43.[151,1435,849,1756]" captionTargetPageId="43" captionText="FIGURE 33. Male of T. serrana Galiano, 1970 (MCN 46708). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression. D same, retrolateral view, red arrow: hook between RTA and prolatero-dorsal tibial apophysis, red dashed line: prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675888" httpUri="https://zenodo.org/record/10675888/files/figure.png" pageId="5" pageNumber="6">33C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016A125D85FBBE0325" box="[1067,1118,1534,1560]" captionStart="FIGURE 46" captionStartId="59.[151,250,1563,1587]" captionTargetBox="[151,1435,633,1540]" captionTargetId="figure-211@59.[151,1435,633,1540]" captionTargetPageId="59" captionText="FIGURE 46. Male of T. peniaflorensis Galiano, 1970 (FNAG36 1463). A habitus, dorsal view. B same, ventral view. C left palp, ventral view, red arrow: tegulum central depression, purple arrow: tegular membranous process. D same, retrolateral view, red arrow: ventral hook between RTA and prolatero-dorsal tibial apophysis. Scale bars: A–B 1 mm, C–D 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.10675918" httpUri="https://zenodo.org/record/10675918/files/figure.png" pageId="5" pageNumber="6">46C</figureCitation>
). Epigynum: epigynal plate with well-developed posteriorly-opening coupling pocket (
<figureCitation id="18892A55FFCB4C016D145E59FC4D0300" box="[813,941,1570,1597]" captionStart="FIGURE 5" captionStartId="10.[151,250,1750,1774]" captionTargetBox="[151,1436,417,1727]" captionTargetId="figure-109@10.[151,1436,417,1727]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electronic microscopy of epigynum, ventral view. A T. musica (MCN 55519), white arrow: septum. B T. aisenbergae Marta, sp. nov. (MCTP 12575), white arrow: septum. C T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135), white arrow: septum. D T. galianoae Marta, sp. nov. (MCTP 39711); red dashed line: window, red arrow: fundus membranous groove of the window. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675798" httpUri="https://zenodo.org/record/10675798/files/figure.png" pageId="5" pageNumber="6">Figs 5A–D</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D805E59FC1F0301" box="[953,1023,1570,1596]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="5" pageNumber="6">7A–Q</figureCitation>
) and usually with a window circling around the copulatory opening, where the membranous copulatory ducts below could inflate during the movement of copulation (
<figureCitation id="18892A55FFCB4C016F005E11FE5703B8" box="[313,439,1642,1669]" captionStart="FIGURE 5" captionStartId="10.[151,250,1750,1774]" captionTargetBox="[151,1436,417,1727]" captionTargetId="figure-109@10.[151,1436,417,1727]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electronic microscopy of epigynum, ventral view. A T. musica (MCN 55519), white arrow: septum. B T. aisenbergae Marta, sp. nov. (MCTP 12575), white arrow: septum. C T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135), white arrow: septum. D T. galianoae Marta, sp. nov. (MCTP 39711); red dashed line: window, red arrow: fundus membranous groove of the window. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675798" httpUri="https://zenodo.org/record/10675798/files/figure.png" pageId="5" pageNumber="6">Figs 5A–C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016FFB5E11FDEB03B9" box="[450,523,1642,1668]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="5" pageNumber="6">7A–Q</figureCitation>
). Copulatory opening median-posteriorly located, sometimes forming a septum or semi-septum, normally facing up (
<figureCitation id="18892A55FFCB4C016C2E5EF5FD750394" box="[535,661,1678,1705]" captionStart="FIGURE 5" captionStartId="10.[151,250,1750,1774]" captionTargetBox="[151,1436,417,1727]" captionTargetId="figure-109@10.[151,1436,417,1727]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electronic microscopy of epigynum, ventral view. A T. musica (MCN 55519), white arrow: septum. B T. aisenbergae Marta, sp. nov. (MCTP 12575), white arrow: septum. C T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135), white arrow: septum. D T. galianoae Marta, sp. nov. (MCTP 39711); red dashed line: window, red arrow: fundus membranous groove of the window. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675798" httpUri="https://zenodo.org/record/10675798/files/figure.png" pageId="5" pageNumber="6">Figs 5A–C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016C995EF5FD230395" box="[672,707,1678,1704]" captionStart="FIGURE 9" captionStartId="15.[151,250,1490,1514]" captionTargetBox="[158,1431,181,1460]" captionTargetId="figure-17@15.[151,1435,181,1465]" captionTargetPageId="15" captionText="FIGURE 9. Female of T. morenensis (Tullgren, 1905) (MCN 19940). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675810" httpUri="https://zenodo.org/record/10675810/files/figure.png" pageId="5" pageNumber="6">9C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016CF65EF5FD1F0395" box="[719,767,1678,1704]" captionStart="FIGURE 11" captionStartId="17.[151,250,1798,1822]" captionTargetBox="[158,1435,539,1756]" captionTargetId="figure-136@17.[151,1435,518,1774]" captionTargetPageId="17" captionText="FIGURE 11. Female of T. guayapae Galiano, 1970. A habitus (MCN 21128), dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum (MCN 55520), ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675814" httpUri="https://zenodo.org/record/10675814/files/figure.png" pageId="5" pageNumber="6">11C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D335EF5FCDC0394" box="[778,828,1678,1705]" captionStart="FIGURE 15" captionStartId="22.[151,250,1490,1514]" captionTargetBox="[159,1432,181,1458]" captionTargetId="figure-20@22.[151,1435,181,1465]" captionTargetPageId="22" captionText="FIGURE 15. Female of T. musica (Mello-Leitão, 1945) (MCN 53372). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 2 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675828" httpUri="https://zenodo.org/record/10675828/files/figure.png" pageId="5" pageNumber="6">15C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D7E5EF5FC980395" box="[839,888,1678,1704]" captionStart="FIGURE 47" captionStartId="60.[151,250,1509,1533]" captionTargetBox="[157,1435,207,1478]" captionTargetId="figure-20@60.[151,1435,201,1485]" captionTargetPageId="60" captionText="FIGURE 47. Female of T. peniaflorensis Galiano, 1970 (MACN-Ar 6242).A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675922" httpUri="https://zenodo.org/record/10675922/files/figure.png" pageId="5" pageNumber="6">47C</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016DBA5EF5FC560395" box="[899,950,1678,1704]" captionStart="FIGURE 49" captionStartId="62.[151,248,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@62.[151,1435,181,1465]" captionTargetPageId="62" captionText="FIGURE 49. Female of T. selenita Galiano, 1970 (FNAG36 1456). A habitus, dorsal view, red arrow: spotted coloration pattern. B same, ventral view. C epigynum, ventral view, red arrow: subseptum. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675926" httpUri="https://zenodo.org/record/10675926/files/figure.png" pageId="5" pageNumber="6">49C</figureCitation>
). In dorsal view, the spiral copulatory ducts with one, two, or three loops, and usually dorso-posteriorly situated from spermathecae (
<figureCitation id="18892A55FFCB4C016A545EC9FB0D03F1" box="[1133,1261,1714,1740]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="5" pageNumber="6">Figs 7A–Q</figureCitation>
). Spermathecae circular to ovoid, with the base of the fertilization ducts placed above the spermathecae at the median-posterior region of the epigynum (
<figureCitation id="18892A55FFCB4C016F955E81FDCE0229" box="[428,558,1786,1812]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="5" pageNumber="6">Figs 7A–Q</figureCitation>
).
</paragraph>
<paragraph id="800D36D0FFCB4C016EFE5F65FEB20298" blockId="5.[151,1437,417,2028]" pageId="5" pageNumber="6">
<emphasis id="B2C6EAC2FFCB4C016EFE5F65FEE60205" bold="true" box="[199,262,1822,1848]" pageId="5" pageNumber="6">Note.</emphasis>
Species of 
<taxonomicName id="47B24D53FFCB4C016FA85F65FDF90205" authorityName="Mello-Leitao" authorityYear="1941" box="[401,537,1822,1848]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016FA85F65FDF90205" box="[401,537,1822,1848]" italics="true" pageId="5" pageNumber="6">Tullgrenella</emphasis>
</taxonomicName>
(
<figureCitation id="18892A55FFCB4C016C135F65FD460205" box="[554,678,1822,1848]" captionStart="FIGURE 1" captionStartId="6.[151,250,1833,1857]" captionTargetBox="[151,1435,522,1806]" captionTargetId="figure-150@6.[151,1435,522,1806]" captionTargetPageId="6" captionText="FIGURE 1. Photographs in vivo of Tullgrenella (morenensis group). A–C T. guayapae Galiano, 1970 (FCE-Ar 11212): A–B female, C male. D–F T. musica (Mello-Leitão, 1945) (FCE-Ar 9637): D–E female, F male. Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675790" httpUri="https://zenodo.org/record/10675790/files/figure.png" pageId="5" pageNumber="6">Figs 1A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016C8D5F65FCEA0205" box="[692,778,1822,1848]" captionStart="FIGURE 20" captionStartId="28.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@28.[151,1435,181,1465]" captionTargetPageId="28" captionText="FIGURE 20. Photographs in vivo. A–D T. quadripunctata (Mello-Leitão, 1944):A–B. female (FCE-Ar 9395), C–D male (FCE- Ar 2284). E–H T. melanica (Mello-Leitão, 1941): E–F female (FCE-Ar 12259), G–H male (FCE-Ar 10505). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675844" httpUri="https://zenodo.org/record/10675844/files/figure.png" pageId="5" pageNumber="6">20A–H</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D2E5F65FC860205" box="[791,870,1822,1848]" captionStart="FIGURE 32" captionStartId="42.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@42.[151,1435,181,1465]" captionTargetPageId="42" captionText="FIGURE 32. Photographs in vivo. A–D T. serrana Galiano, 1970:A–B female (FCE-Ar 11149), C–D male (FCE-Ar 9538). E–F female of T. legalissima Marta &amp; Hagopián, sp. nov. (FCE-Ar 12155). Photographs by D. Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675884" httpUri="https://zenodo.org/record/10675884/files/figure.png" pageId="5" pageNumber="6">32A–F</figureCitation>
, 
<figureCitation id="18892A55FFCB4C016D4D5F65FC230204" box="[884,963,1822,1849]" captionStart="FIGURE 45" captionStartId="58.[151,250,1689,1713]" captionTargetBox="[151,1435,381,1665]" captionTargetId="figure-102@58.[151,1435,381,1665]" captionTargetPageId="58" captionText="FIGURE 45. Photographs in vivo of T. peniaflorensis Galiano, 1970. A–F male. Photographs by Asiel Olivares." figureDoi="http://doi.org/10.5281/zenodo.10675916" httpUri="https://zenodo.org/record/10675916/files/figure.png" pageId="5" pageNumber="6">45A–F</figureCitation>
) have bright colors, as well as a slightly elongated body, as in most genera of the subtribe e.g., 
<taxonomicName id="47B24D53FFCB4C016CCC5F39FBE90261" authority="C. L. Koch, 1846" authorityName="C. L. Koch" authorityYear="1846" box="[757,1033,1858,1884]" class="Arachnida" family="Salticidae" genus="Phiale" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016CCC5F39FCDE0261" box="[757,830,1858,1884]" italics="true" pageId="5" pageNumber="6">Phiale</emphasis>
C. L. Koch, 1846
</taxonomicName>
, 
<taxonomicName id="47B24D53FFCB4C016A2A5F39FB9B0261" authorityName="C. L. Kock" authorityYear="1850" box="[1043,1147,1858,1884]" class="Arachnida" family="Salticidae" genus="Aphirape" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016A2A5F39FB9B0261" box="[1043,1147,1858,1884]" italics="true" pageId="5" pageNumber="6">Aphirape</emphasis>
</taxonomicName>
, 
<taxonomicName id="47B24D53FFCB4C016AB05F39FAC00261" authorityName="Galiano" authorityYear="1983" box="[1161,1312,1858,1884]" class="Arachnida" family="Salticidae" genus="Sumampattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016AB05F39FAC00261" box="[1161,1312,1858,1884]" italics="true" pageId="5" pageNumber="6">Sumampattus</emphasis>
</taxonomicName>
, 
<taxonomicName id="47B24D53FFCB4C016B145F39FA8D0261" authorityName="Peckham &amp; Peckham" authorityYear="1896" box="[1325,1389,1858,1884]" class="Arachnida" family="Salticidae" genus="Akela" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016B145F39FA8D0261" box="[1325,1389,1858,1884]" italics="true" pageId="5" pageNumber="6">Akela</emphasis>
</taxonomicName>
and 
<taxonomicName id="47B24D53FFCB4C016EAE5F1DFEEF02BD" authorityName="Galiano" authorityYear="1987" box="[151,271,1894,1920]" class="Arachnida" family="Salticidae" genus="Hisukattus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFCB4C016EAE5F1DFEEF02BD" box="[151,271,1894,1920]" italics="true" pageId="5" pageNumber="6">Hisukattus</emphasis>
</taxonomicName>
. These features suggest possible mimetic relationships with other aposematically colored arthropods (
<bibRefCitation id="E4234B21FFCB4C016EA65FF1FEA60298" author="Edwards, G. B." box="[159,326,1930,1957]" pageId="5" pageNumber="6" pagination="1 - 87" refId="ref41883" refString="Edwards, G. B. (2015) Freyinae, a major new subfamily of Neotropical jumping spiders (Araneae: Salticidae). Zootaxa, 4036 (1), 1 - 87. https: // doi. org / 10.11646 / zootaxa. 4036.1.1" type="journal article" year="2015">Edwards 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8A8655BFFCB4C026EFE5FD6FB6904EC" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="distribution">
<paragraph id="800D36D0FFCB4C016EFE5FD6FB6302F4" blockId="5.[151,1437,417,2028]" box="[199,1155,1965,1993]" pageId="5" pageNumber="6">
<emphasis id="B2C6EAC2FFCB4C016EFE5FD6FE8102FA" bold="true" box="[199,353,1965,1991]" pageId="5" pageNumber="6">Distribution.</emphasis>
<collectingCountry id="F8A57640FFCB4C016F5E5FD5FE5C02F5" box="[359,444,1966,1992]" name="Bolivia" pageId="5" pageNumber="6">Bolivia</collectingCountry>
, 
<collectingCountry id="F8A57640FFCB4C016FF15FD5FDE702F5" box="[456,519,1966,1992]" name="Chile" pageId="5" pageNumber="6">Chile</collectingCountry>
, 
<collectingCountry id="F8A57640FFCB4C016C2A5FD5FDBA02F5" box="[531,602,1966,1992]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
, 
<collectingCountry id="F8A57640FFCB4C016C5C5FD5FD2E02F5" box="[613,718,1966,1992]" name="Paraguay" pageId="5" pageNumber="6">Paraguay</collectingCountry>
, 
<collectingCountry id="F8A57640FFCB4C016CE05FD5FCDC02F5" box="[729,828,1966,1992]" name="Uruguay" pageId="5" pageNumber="6">Uruguay</collectingCountry>
, and 
<collectingCountry id="F8A57640FFCB4C016D4C5FD5FC0402F5" box="[885,996,1966,1992]" name="Argentina" pageId="5" pageNumber="6">Argentina</collectingCountry>
(
<figureCitation id="18892A55FFCB4C016DCD5FD5FB9702F4" box="[1012,1143,1966,1993]" captionStart-0="FIGURE 50" captionStart-1="FIGURE51" captionStart-2="FIGURE 52" captionStart-3="FIGURE 53" captionStart-4="FIGURE 54" captionStartId-0="63.[151,250,1689,1713]" captionStartId-1="64.[151,250,1490,1514]" captionStartId-2="65.[151,250,1797,1821]" captionStartId-3="66.[151,250,1490,1514]" captionStartId-4="67.[151,250,1451,1475]" captionTargetBox-0="[151,1435,381,1665]" captionTargetBox-1="[151,1435,181,1465]" captionTargetBox-2="[151,1435,489,1773]" captionTargetBox-3="[151,1435,181,1465]" captionTargetBox-4="[151,1435,181,1339]" captionTargetId-0="figure-82@63.[151,1435,381,1665]" captionTargetId-1="figure-20@64.[151,1435,181,1465]" captionTargetId-2="figure-142@65.[151,1435,489,1773]" captionTargetId-3="figure-20@66.[151,1435,181,1465]" captionTargetId-4="figure-17@67.[151,1435,181,1428]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="65" captionTargetPageId-3="66" captionTargetPageId-4="67" captionText-0="FIGURE 50. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the morenensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Argentina: La Pampa, Lihuel Calel, photographed by collector Ivan Magalhães." captionText-1="FIGURE51. NeotropicalandAndean regionalization(Morrone 2014) anddistributional recordsfor speciesof the quadripunctata group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Montevideo, Melilla, photographed by collector Damián Hagopián." captionText-2="FIGURE 52. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the melanica group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Río Negro, Estancia Las Cadenas, photographed by collector Damián Hagopián." captionText-3="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." captionText-4="FIGURE 54. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the peniaflorensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence." figureDoi-0="http://doi.org/10.5281/zenodo.10675928" figureDoi-1="http://doi.org/10.5281/zenodo.10675930" figureDoi-2="http://doi.org/10.5281/zenodo.10675932" figureDoi-3="http://doi.org/10.5281/zenodo.10675934" figureDoi-4="http://doi.org/10.5281/zenodo.10675936" httpUri-0="https://zenodo.org/record/10675928/files/figure.png" httpUri-1="https://zenodo.org/record/10675930/files/figure.png" httpUri-2="https://zenodo.org/record/10675932/files/figure.png" httpUri-3="https://zenodo.org/record/10675934/files/figure.png" httpUri-4="https://zenodo.org/record/10675936/files/figure.png" pageId="5" pageNumber="6">Figs 50–54</figureCitation>
).
</paragraph>
<paragraph id="800D36D0FFCB4C026EFE5FA9FDA30420" blockId="5.[151,1437,417,2028]" lastBlockId="6.[151,1437,151,465]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B2C6EAC2FFCB4C016EFE5FA9FE6902D6" bold="true" box="[199,393,2001,2028]" pageId="5" pageNumber="6">Natural history.</emphasis>
Most individuals were collected on outcrops. Their habitat can be considered the soil or epigeous shrubs that occur in southern South America (grasslands and rocks), such as Chacoan subregion (
<collectingCountry id="F8A57640FFC84C026B7658ECFA78058C" box="[1359,1432,151,177]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
, 
<collectingCountry id="F8A57640FFC84C026EAE58C0FF1A05E8" box="[151,250,187,213]" name="Uruguay" pageId="6" pageNumber="7">Uruguay</collectingCountry>
, and 
<collectingCountry id="F8A57640FFC84C026F0D58C0FE4805E8" box="[308,424,187,213]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
), Brazilian subregion (Northern 
<collectingCountry id="F8A57640FFC84C026D2C58C0FCBE05E8" box="[789,862,187,213]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
), and Andean region (
<collectingCountry id="F8A57640FFC84C026A6C58C0FB7805E8" box="[1109,1176,187,213]" name="Chile" pageId="6" pageNumber="7">Chile</collectingCountry>
). Few individuals were recorded in the South American Transition Zone (
<figureCitation id="18892A55FFC84C026CE858A4FCB605C4" box="[721,854,222,249]" captionStart-0="FIGURE 50" captionStart-1="FIGURE51" captionStart-2="FIGURE 52" captionStart-3="FIGURE 53" captionStart-4="FIGURE 54" captionStartId-0="63.[151,250,1689,1713]" captionStartId-1="64.[151,250,1490,1514]" captionStartId-2="65.[151,250,1797,1821]" captionStartId-3="66.[151,250,1490,1514]" captionStartId-4="67.[151,250,1451,1475]" captionTargetBox-0="[151,1435,381,1665]" captionTargetBox-1="[151,1435,181,1465]" captionTargetBox-2="[151,1435,489,1773]" captionTargetBox-3="[151,1435,181,1465]" captionTargetBox-4="[151,1435,181,1339]" captionTargetId-0="figure-82@63.[151,1435,381,1665]" captionTargetId-1="figure-20@64.[151,1435,181,1465]" captionTargetId-2="figure-142@65.[151,1435,489,1773]" captionTargetId-3="figure-20@66.[151,1435,181,1465]" captionTargetId-4="figure-17@67.[151,1435,181,1428]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="65" captionTargetPageId-3="66" captionTargetPageId-4="67" captionText-0="FIGURE 50. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the morenensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Argentina: La Pampa, Lihuel Calel, photographed by collector Ivan Magalhães." captionText-1="FIGURE51. NeotropicalandAndean regionalization(Morrone 2014) anddistributional recordsfor speciesof the quadripunctata group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Montevideo, Melilla, photographed by collector Damián Hagopián." captionText-2="FIGURE 52. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the melanica group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Río Negro, Estancia Las Cadenas, photographed by collector Damián Hagopián." captionText-3="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." captionText-4="FIGURE 54. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the peniaflorensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence." figureDoi-0="http://doi.org/10.5281/zenodo.10675928" figureDoi-1="http://doi.org/10.5281/zenodo.10675930" figureDoi-2="http://doi.org/10.5281/zenodo.10675932" figureDoi-3="http://doi.org/10.5281/zenodo.10675934" figureDoi-4="http://doi.org/10.5281/zenodo.10675936" httpUri-0="https://zenodo.org/record/10675928/files/figure.png" httpUri-1="https://zenodo.org/record/10675930/files/figure.png" httpUri-2="https://zenodo.org/record/10675932/files/figure.png" httpUri-3="https://zenodo.org/record/10675934/files/figure.png" httpUri-4="https://zenodo.org/record/10675936/files/figure.png" pageId="6" pageNumber="7">Figs 50–54</figureCitation>
). The altitudinal occurrence of these species varies from 
<quantity id="474A9B35FFC84C026EEC5978FEBB0420" box="[213,347,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" metricValueMax="4.776" metricValueMin="0.004" pageId="6" pageNumber="7" unit="m" value="2390.0" valueMax="4776.0" valueMin="4.0">4 to 4776 m</quantity>
a. s. l. (
<figureCitation id="18892A55FFC84C026F8A5978FDD60420" box="[435,566,258,285]" captionStart-0="FIGURE 50" captionStart-1="FIGURE51" captionStart-2="FIGURE 52" captionStart-3="FIGURE 53" captionStart-4="FIGURE 54" captionStartId-0="63.[151,250,1689,1713]" captionStartId-1="64.[151,250,1490,1514]" captionStartId-2="65.[151,250,1797,1821]" captionStartId-3="66.[151,250,1490,1514]" captionStartId-4="67.[151,250,1451,1475]" captionTargetBox-0="[151,1435,381,1665]" captionTargetBox-1="[151,1435,181,1465]" captionTargetBox-2="[151,1435,489,1773]" captionTargetBox-3="[151,1435,181,1465]" captionTargetBox-4="[151,1435,181,1339]" captionTargetId-0="figure-82@63.[151,1435,381,1665]" captionTargetId-1="figure-20@64.[151,1435,181,1465]" captionTargetId-2="figure-142@65.[151,1435,489,1773]" captionTargetId-3="figure-20@66.[151,1435,181,1465]" captionTargetId-4="figure-17@67.[151,1435,181,1428]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="65" captionTargetPageId-3="66" captionTargetPageId-4="67" captionText-0="FIGURE 50. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the morenensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Argentina: La Pampa, Lihuel Calel, photographed by collector Ivan Magalhães." captionText-1="FIGURE51. NeotropicalandAndean regionalization(Morrone 2014) anddistributional recordsfor speciesof the quadripunctata group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Montevideo, Melilla, photographed by collector Damián Hagopián." captionText-2="FIGURE 52. Neotropical and Andean regionalization (Morrone 2014) and distributional records for species of the melanica group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, Uruguay: Río Negro, Estancia Las Cadenas, photographed by collector Damián Hagopián." captionText-3="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." captionText-4="FIGURE 54. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the peniaflorensis group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence." figureDoi-0="http://doi.org/10.5281/zenodo.10675928" figureDoi-1="http://doi.org/10.5281/zenodo.10675930" figureDoi-2="http://doi.org/10.5281/zenodo.10675932" figureDoi-3="http://doi.org/10.5281/zenodo.10675934" figureDoi-4="http://doi.org/10.5281/zenodo.10675936" httpUri-0="https://zenodo.org/record/10675928/files/figure.png" httpUri-1="https://zenodo.org/record/10675930/files/figure.png" httpUri-2="https://zenodo.org/record/10675932/files/figure.png" httpUri-3="https://zenodo.org/record/10675934/files/figure.png" httpUri-4="https://zenodo.org/record/10675936/files/figure.png" pageId="6" pageNumber="7">Figs 50–54</figureCitation>
).
</paragraph>
<paragraph id="800D36D0FFC84C026EFE595DFB6904EC" blockId="6.[151,1437,151,465]" pageId="6" pageNumber="7">
<emphasis id="B2C6EAC2FFC84C026EFE595DFE9E047C" bold="true" box="[199,382,294,321]" pageId="6" pageNumber="7">Species groups.</emphasis>
Based on the morphological variation of genital structures (palp and epigynum), as well as the geographic regionalization and altitudinal occurrence, we propose five species groups for the genus 
<taxonomicName id="47B24D53FFC84C026B345930FA750458" authorityName="Mello-Leitao" authorityYear="1941" box="[1293,1429,331,357]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B2C6EAC2FFC84C026B345930FA750458" box="[1293,1429,331,357]" italics="true" pageId="6" pageNumber="7">Tullgrenella</emphasis>
</taxonomicName>
: 
<taxonomicName id="47B24D53FFC84C026EAE5914FEF604B5" authorityName="Tullgren" authorityYear="1905" box="[151,278,367,392]" class="Arachnida" family="Salticidae" genus="Euophrys" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="morenensis">
<emphasis id="B2C6EAC2FFC84C026EAE5914FEF604B5" box="[151,278,367,392]" italics="true" pageId="6" pageNumber="7">morenensis</emphasis>
</taxonomicName>
group (with six species, including the 
<typeStatus id="5F098872FFC84C026CE05914FCE904B4" box="[729,777,367,393]" pageId="6" pageNumber="7">type</typeStatus>
species); 
<taxonomicName id="47B24D53FFC84C026D475914FBCB04B4" authorityName="MCTP" authorityYear="2239" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1944" box="[894,1067,367,393]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadripunctata">
<emphasis id="B2C6EAC2FFC84C026D475914FBCB04B4" box="[894,1067,367,393]" italics="true" pageId="6" pageNumber="7">quadripunctata</emphasis>
</taxonomicName>
group (three species); 
<taxonomicName id="47B24D53FFC84C026B0F5914FA7D04B4" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[1334,1437,367,393]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="melanica">
<emphasis id="B2C6EAC2FFC84C026B0F5914FA7D04B4" box="[1334,1437,367,393]" italics="true" pageId="6" pageNumber="7">melanica</emphasis>
</taxonomicName>
group (two species); 
<taxonomicName id="47B24D53FFC84C026FB859E8FE380491" authorityName="Galiano" authorityYear="1970" box="[385,472,403,428]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="serrana">
<emphasis id="B2C6EAC2FFC84C026FB859E8FE380491" box="[385,472,403,428]" italics="true" pageId="6" pageNumber="7">serrana</emphasis>
</taxonomicName>
group (six species, included four new species) and 
<taxonomicName id="47B24D53FFC84C026A2C59E8FB530490" authorityName="Galiano" authorityYear="1970" box="[1045,1203,403,429]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="peniaflorensis">
<emphasis id="B2C6EAC2FFC84C026A2C59E8FB530490" box="[1045,1203,403,429]" italics="true" pageId="6" pageNumber="7">peniaflorensis</emphasis>
</taxonomicName>
group (two species). Putative synapomorphies of genital structures for each species group are suggested below.
</paragraph>
</subSubSection>
</treatment>
</document>