treatments-xml/data/08/12/4C/08124C2AFFF7FF84FDA8FDBAFD895369.xml

<document id="646BD0BF5C793EA8217196C5554AE6C5" ID-CLB-Dataset="31128" ID-DOI="10.5852/ejt.2017.381" ID-GBIF-Dataset="4bedeb19-2f4c-4325-9ab4-22eace75b174" ID-Zenodo-Dep="1118110" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513662381649" checkinUser="plazi" docAuthor="Moritz, Leif &amp; Wesener, Thomas" docDate="2017" docId="08124C2AFFF7FF84FDA8FDBAFD895369" docLanguage="en" docName="ejt.381.pdf" docOrigin="European Journal of Taxonomy 381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sphaeromimus kalambatritra Moritz &amp; Wesener 2017, sp. nov." docType="treatment" docUuid="620CAD6B-DAB4-4FFC-8556-61357C75AF40" docUuidSource="ZooBank" docVersion="8" lastPageNumber="13" masterDocId="F42B3452FFF1FF88FF9FFFAFFFD5557C" masterDocTitle="Integrative description of two new species of Malagasy chirping giant pill-millipedes, genus Sphaeromimus (Diplopoda: Sphaerotheriida: Arthrosphaeridae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1698510406634" updateUser="ExternalLinkService">
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<mods:title id="0E9E90CD0479E40401EABE82659705BD">Integrative description of two new species of Malagasy chirping giant pill-millipedes, genus Sphaeromimus (Diplopoda: Sphaerotheriida: Arthrosphaeridae)</mods:title>
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<mods:namePart id="5682420B238D6AFA7C94F196AB2EF5F2">Moritz, Leif</mods:namePart>
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<mods:date id="F4BF691A7EB2F448ACAEDC24BB67FA15">2017</mods:date>
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<taxonomicName id="47BB86BFFFF7FF8EFDA8FDBAFC4E574C" authority="Moritz &amp; Wesener, 2017" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[567,923,533,560]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kalambatritra" status="sp. nov.">
<emphasis id="B2CF212EFFF7FF8EFDA8FDBAFC4E574C" bold="true" box="[567,923,533,560]" italics="true" pageId="6" pageNumber="7">Sphaeromimus kalambatritra</emphasis>
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<paragraph id="8004FD3CFFF7FF8EFEF5FD96FCB7570B" blockId="6.[362,1225,533,631]" pageId="6" pageNumber="7">
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6B-DAB4-4FFC-8556-61357C75AF40 
<figureCitation id="1880E1B9FFF7FF8EFD4EFDF2FCEC570B" box="[721,825,605,631]" captionStart-0="Fig. 1" captionStart-1="Fig.2" captionStart-2="Fig. 3" captionStart-3="Fig.4" captionStart-4="Fig. 5" captionStart-5="Fig. 6" captionStartId-0="7.[189,232,1686,1712]" captionStartId-1="8.[189,232,1930,1956]" captionStartId-2="9.[189,232,1825,1851]" captionStartId-3="10.[189,232,1928,1954]" captionStartId-4="11.[189,232,1788,1814]" captionStartId-5="12.[189,232,758,784]" captionTargetBox-0="[189,1398,711,1646]" captionTargetBox-1="[189,1398,1079,1886]" captionTargetBox-2="[189,1399,438,1783]" captionTargetBox-3="[189,1401,1078,1882]" captionTargetBox-4="[193,1396,571,1748]" captionTargetBox-5="[189,1396,263,730]" captionTargetId-0="figure@7.[189,1398,711,1646]" captionTargetId-1="figure@8.[74,1398,1079,1887]" captionTargetId-2="figure@9.[189,1399,438,1783]" captionTargetId-3="figure@10.[189,1401,1078,1883]" captionTargetId-4="figure@11.[192,1396,569,1748]" captionTargetId-5="figure@12.[189,1396,263,730]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." captionText-1="Fig.2.Sphaeromimus kalambatritrasp.nov.,SEM ofleft antenna,♂, holotype (CASENT9068297-A). A. Overview. B.Antennal disc, detail. C. Margin of antennal disc, detail.Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensorycone; I–VI = antennomeres." captionText-2="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." captionText-3="Fig.4.Sphaeromimuskalambatritra sp. nov., SEM of endotergum, ♂, holotype (CASENT9068297-A). A.Overview,endotergum.B. Marginofendotergum,detail.Abbreviations:(1)=innerpart; (2)=middle part; (3) =outer part." captionText-4="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." captionText-5="Fig. 6. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♀, paratype (CASENT 9058301). A. Left vulva (2 nd coxa). B. Subanal plate. Abbreviations: cx = coxa; ep = exterior plate; ip = interior plate; op = operculum; pref =prefemur; sr = stridulation ribs. Scale bars =1 mm." httpUri-0="https://zenodo.org/record/1118112/files/figure.png" httpUri-1="https://zenodo.org/record/1118114/files/figure.png" httpUri-2="https://zenodo.org/record/1118116/files/figure.png" httpUri-3="https://zenodo.org/record/1118118/files/figure.png" httpUri-4="https://zenodo.org/record/1118120/files/figure.png" httpUri-5="https://zenodo.org/record/1118122/files/figure.png" pageId="6" pageNumber="7">Figs 1–6</figureCitation>
, 
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</heading>
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<subSubSection id="C8A1AEB7FFF7FF8EFF22FD30FBA6562E" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8004FD3CFFF7FF8EFF22FD30FEE357C5" blockId="6.[189,310,671,697]" box="[189,310,671,697]" pageId="6" pageNumber="7">
<heading id="DB4C4A50FFF7FF8EFF22FD30FEE357C5" bold="true" box="[189,310,671,697]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B2CF212EFFF7FF8EFF22FD30FEE357C5" bold="true" box="[189,310,671,697]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
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<paragraph id="8004FD3CFFF7FF8EFF22FD61FBA6562E" blockId="6.[189,1399,717,851]" pageId="6" pageNumber="7">
Large, massive, brown 
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<emphasis id="B2CF212EFFF7FF8EFE53FD61FDAC5794" box="[460,633,718,744]" italics="true" pageId="6" pageNumber="7">Sphaeromimus</emphasis>
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,&gt; 
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long. Differing from all other species of 
<taxonomicName id="47BB86BFFFF7FF8EFB56FD61FAA35794" authorityName="de Saussure &amp; Zehntner" authorityYear="1902" box="[1225,1398,718,744]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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in having five stridulation ribs on the male harp (as in 
<taxonomicName id="47BB86BFFFF7FF8EFCC3FD5DFA945677" authority="(Saussure &amp; Zehntner, 1897)" authorityName="" authorityYear="1897" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[860,1345,753,780]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="musicus">
<emphasis id="B2CF212EFFF7FF8EFCC3FD5DFC0E5677" box="[860,987,754,780]" italics="true" pageId="6" pageNumber="7">S. musicus</emphasis>
(Saussure &amp; Zehntner, 1897)
</taxonomicName>
and 
<taxonomicName id="47BB86BFFFF7FF8EFF22FCBAFE295653" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[189,508,789,815]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vatovavy">
<emphasis id="B2CF212EFFF7FF8EFF22FCBAFE915653" box="[189,324,789,815]" italics="true" pageId="6" pageNumber="7">S. vatovavy</emphasis>
<bibRefCitation id="E42A80CDFFF7FF8EFED2FCBAFE295653" author="Blanke A. &amp; Wesener T." box="[333,508,789,815]" journalOrPublisher="Arthropod Structure &amp; Development" pageId="6" pageNumber="7" pagination="63 - 75" part="43" publicationUrl="https://doi.org/10.1016/j.asd.2013.10.003" refString="Blanke A. &amp; Wesener T. 2014. Revival of forgotten characters and modern imaging techniques help to produce a robust phylogeny of the Diplopoda (Arthropoda, Myriapoda). Arthropod Structure &amp; Development 43: 63 - 75. https: // doi. org / 10.1016 / j. asd. 2013.10.003" title="Revival of forgotten characters and modern imaging techniques help to produce a robust phylogeny of the Diplopoda (Arthropoda, Myriapoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
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) in the following characters: tarsus of leg 3 without an apical spine, coxal process almost absent, posterior telopod with two membranous lobes.&gt;80 ocelli.
</paragraph>
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<subSubSection id="C8A1AEB7FFF7FF8EFF22FCD5FE22569B" pageId="6" pageNumber="7" type="etymology">
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<emphasis id="B2CF212EFFF7FF8EFF22FCD5FE9156E8" bold="true" box="[189,324,890,916]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8004FD3CFFF7FF8EFF22FC06FE22569B" blockId="6.[189,1406,937,999]" pageId="6" pageNumber="7">
The specific epithet ‘kalambatritra’, noun in apposition, refers to the type-locality, the Réserve Spéciale de Kalambatritra (
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).
</paragraph>
</subSubSection>
<subSubSection id="C8A1AEB7FFF7FF8EFF22FBA1FCD950EF" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8004FD3CFFF7FF8EFF22FBA1FD975155" blockId="6.[189,578,1038,1065]" box="[189,578,1038,1065]" pageId="6" pageNumber="7">
<emphasis id="B2CF212EFFF7FF8EFF22FBA1FE7F5154" bold="true" box="[189,426,1038,1064]" pageId="6" pageNumber="7">Material examined</emphasis>
(3 ♂♂, 1 ♀)
</paragraph>
<materialsCitation id="30D3F761FFF7FF8EFF75FB91FE0251BF" ID-GBIF-Occurrence="1795859480" collectedFrom=" montane  rainforest" collectingDate="2009-02" collectingMethod="general collecting" collectionCode="CASENT" collectorName="B. L.  Fisher " country="Madagascar" county="Befarara" elevation="1390" latitude="-23.41733" location="Toliaria,   Réserve  Spéciale  Kalambatritra,   Befarara" longitude="46.442" municipality="Kalambatritra" pageId="6" pageNumber="7" specimenCode="CASENT 9068297-A" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectingCountry id="F8ACBDACFFF7FF8EFF22FBCEFE475107" box="[189,402,1121,1147]" name="Madagascar" pageId="6" pageNumber="7">MADAGASCAR</collectingCountry>
: 
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, 
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, BLF21330, 
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, 
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,alt. 
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</elevation>
, 
<collectedFrom id="33A7BEC1FFF7FF8EFE6DFB2AFD0B51E3" box="[498,734,1157,1183]" pageId="6" pageNumber="7">montane rainforest</collectedFrom>
, 
<collectingDate id="E4412214FFF7FF8EFD73FB2AFCAA51E3" box="[748,895,1157,1183]" pageId="6" pageNumber="7">
7 
<date id="F405DBFCFFF7FF8EFC9AFB2AFCAA51E3" box="[773,895,1157,1183]" pageId="6" pageNumber="7" value="2009-02">Feb.2009</date>
</collectingDate>
, 
<collectorName id="2D4E98EAFFF7FF8EFC11FB2AFBC851E3" box="[910,1053,1157,1183]" pageId="6" pageNumber="7">B.L. Fisher</collectorName>
<emphasis id="B2CF212EFFF7FF8EFB80FB29FB8951E3" box="[1055,1116,1157,1183]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
leg., 
<collectingMethod id="59FA852BFFF7FF8EFB3CFB2AFAA251E3" box="[1187,1399,1157,1183]" pageId="6" pageNumber="7">general collecting</collectingMethod>
(
<specimenCode id="D01D5547FFF7FF8EFF5AFB07FE1E51BF" box="[197,459,1192,1219]" pageId="6" pageNumber="7">CASENT 9068297-A</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8004FD3CFFF7FF8EFF75FB45FEB25078" blockId="6.[189,1406,1258,1427]" box="[234,359,1258,1284]" pageId="6" pageNumber="7">
<emphasis id="B2CF212EFFF7FF8EFF75FB45FEB25078" bold="true" box="[234,359,1258,1284]" pageId="6" pageNumber="7">Paratypes</emphasis>
</paragraph>
<paragraph id="8004FD3CFFF7FF8EFF22FAA1FCD950EF" blockId="6.[189,1406,1258,1427]" pageId="6" pageNumber="7">
<materialsCitation id="30D3F761FFF7FF8EFF22FAA1FB525054" ID-GBIF-Occurrence="1795859477" box="[189,1159,1293,1320]" collectionCode="ZFMK" country="Madagascar" location="MADAGASCAR" pageId="6" pageNumber="7" specimenCode="MYR06125" specimenCount="2" specimenCount-male="1">
<collectingCountry id="F8ACBDACFFF7FF8EFF22FAA1FE475054" box="[189,402,1294,1320]" name="Madagascar" pageId="6" pageNumber="7">MADAGASCAR</collectingCountry>
: 
<specimenCount id="96BD36B5FFF7FF8EFE03FAA1FE7E5054" box="[412,427,1294,1320]" pageId="6" pageNumber="7">1</specimenCount>
<specimenCount id="96BD36B5FFF7FF8EFE2DFAA1FE1F5054" box="[434,458,1294,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
, same collection data as for holotype (
<collectionCode id="E6AA65F9FFF7FF8EFC17FAA1FC0E5054" box="[904,987,1294,1320]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="7">ZFMK</collectionCode>
<specimenCode id="D01D5547FFF7FF8EFC7DFAA2FBA15054" box="[994,1140,1293,1320]" pageId="6" pageNumber="7">MYR06125</specimenCode>
);
</materialsCitation>
<materialsCitation id="30D3F761FFF7FF8EFB19FAA1FD125030" ID-GBIF-Occurrence="1795859476" collectionCode="CASENT" country="Madagascar" location="MADAGASCAR" pageId="6" pageNumber="7" specimenCode="CASENT  9068297-C" specimenCount="1" specimenCount-male="1">
1 
<specimenCount id="96BD36B5FFF7FF8EFB03FAA1FB615054" box="[1180,1204,1294,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
, same collection data as for holotype (
<specimenCode id="D01D5547FFF7FF8EFE29FA9EFD6F5030" box="[438,698,1329,1356]" pageId="6" pageNumber="7">CASENT 9068297-C</specimenCode>
);
</materialsCitation>
<materialsCitation id="30D3F761FFF7FF8EFD50FA9EFCD950EF" ID-GBIF-Occurrence="1795859479" collectedFrom="montane  rainforest" collectingDate="9 - 10   Feb. 2009" collectingMethod="general collecting" collectionCode="BLF, CASENT" collectorName="B. L.   Fisher " elevation="1270" latitude="-23.463612" location="Toliaria,  Reserve  Speciale  Kalambatritra,   Ampanihy" pageId="6" pageNumber="7" specimenCode="BLF21565, CASENT 9058301" specimenCount="1" specimenCount-female="1">
1 
<specimenCount id="96BD36B5FFF7FF8EFD7AFA9DFD2E5030" box="[741,763,1330,1356]" pageId="6" pageNumber="7" type="female">♀</specimenCount>
, 
<location id="8564ABE7FFF7FF8EFC95FA9EFAA75037" LSID="urn:lsid:plazi:treatment:08124C2AFFF7FF84FDA8FDBAFD895369:8564ABE7FFF7FF8EFC95FA9EFAA75037" box="[778,1394,1329,1356]" latitude="-23.463612" name="Toliaria,  Reserve  Speciale  Kalambatritra,   Ampanihy" pageId="6" pageNumber="7">Toliaria, Réserve Spéciale Kalambatritra, Ampanihy</location>
, 
<specimenCode id="D01D5547FFF7FF8EFF22FAFAFEEA500C" box="[189,319,1365,1392]" pageId="6" pageNumber="7">BLF21565</specimenCode>
, 
<geoCoordinate id="E58F9BFBFFF7FF8EFED1FAFCFE345013" box="[334,481,1363,1391]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="-23.463612">23°27′49″S</geoCoordinate>
, 
<geoCoordinate id="E58F9BFBFFF7FF8EFE6FFAFCFD5D5013" box="[496,648,1363,1391]" pageId="6" pageNumber="7">46°27′47″ E</geoCoordinate>
,alt. 
<elevation id="0B961A0FFFF7FF8EFD5DFAFAFCF55013" box="[706,800,1365,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="m" value="1270.0">
<quantity id="474350D9FFF7FF8EFD5DFAFAFCF55013" box="[706,800,1365,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="m" value="1270.0">1270m</quantity>
</elevation>
, 
<collectedFrom id="33A7BEC1FFF7FF8EFCB0FAFAFBC65013" box="[815,1043,1365,1391]" pageId="6" pageNumber="7">montane rainforest</collectedFrom>
, 
<collectingDate id="E4412214FFF7FF8EFBBFFAFAFB0A5013" box="[1056,1247,1365,1391]" pageId="6" pageNumber="7">
9–10 
<date id="F405DBFCFFF7FF8EFBFAFAFAFB0A5013" box="[1125,1247,1365,1391]" pageId="6" pageNumber="7" value="2009-02">Feb.2009</date>
</collectingDate>
, 
<collectorName id="2D4E98EAFFF7FF8EFB70FAFAFAAB5013" box="[1263,1406,1365,1391]" pageId="6" pageNumber="7">B.L. Fisher</collectorName>
<emphasis id="B2CF212EFFF7FF8EFF22FAD6FF2D50EE" box="[189,248,1400,1426]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
leg., 
<collectingMethod id="59FA852BFFF7FF8EFEA6FAD7FDDF50EE" box="[313,522,1400,1426]" pageId="6" pageNumber="7">general collecting</collectingMethod>
(
<specimenCode id="D01D5547FFF7FF8EFD86FAD6FCD550EF" box="[537,768,1400,1427]" pageId="6" pageNumber="7">CASENT 9058301</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8A1AEB7FFF7FF84FF22FA15FE87502A" lastPageId="12" lastPageNumber="13" pageId="6" pageNumber="7" type="description">
<paragraph id="8004FD3CFFF7FF8EFF22FA15FE9B50A8" blockId="6.[189,334,1466,1491]" box="[189,334,1466,1492]" pageId="6" pageNumber="7">
<emphasis id="B2CF212EFFF7FF8EFF22FA15FE9B50A8" bold="true" box="[189,334,1466,1492]" pageId="6" pageNumber="7">Description</emphasis>
</paragraph>
<paragraph id="8004FD3CFFF7FF8EFF22FA45FDD85336" blockId="6.[189,1399,1511,1611]" pageId="6" pageNumber="7">
MEASUREMENTS. Male 
<typeStatus id="5F00439EFFF7FF8EFE21FA46FDF0537F" box="[446,549,1513,1539]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
ca 
<quantity id="474350D9FFF7FF8EFDD1FA46FD635378" box="[590,694,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" pageId="6" pageNumber="7" unit="mm" value="51.5">51.5 mm</quantity>
long, 
<quantity id="474350D9FFF7FF8EFC9FFA46FCBD5378" box="[768,872,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="2.44" pageId="6" pageNumber="7" unit="mm" value="24.4">24.4 mm</quantity>
(2nd segment) up to 
<quantity id="474350D9FFF7FF8EFBCDFA46FB6F5378" box="[1106,1210,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="6" pageNumber="7" unit="mm" value="28.2">28.2 mm</quantity>
(10th segment = widest) wide, 
<quantity id="474350D9FFF7FF8EFEFDF9A2FE61535B" box="[354,436,1549,1575]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="7" unit="mm" value="14.0">14 mm</quantity>
(2nd segment) high. Female ca 
<quantity id="474350D9FFF7FF8EFC83F9A2FCBB535B" box="[796,878,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="7" unit="mm" value="52.0">52 mm</quantity>
long, 
<quantity id="474350D9FFF7FF8EFC27F9A2FBF5535B" box="[952,1056,1549,1575]" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="6" pageNumber="7" unit="mm" value="24.7">24.7 mm</quantity>
(2nd segment) wide, 
<quantity id="474350D9FFF7FF8EFA91F9A2FAA2535B" box="[1294,1399,1549,1576]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="6" pageNumber="7" unit="mm" value="15.1">15.1 mm</quantity>
(2nd segment) high (
<figureCitation id="1880E1B9FFF7FF8EFE3DF99FFE2A5336" box="[418,511,1584,1610]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8004FD3CFFF7FF8EFF22F9DDFD045267" blockId="6.[189,1399,1650,1819]" pageId="6" pageNumber="7">
COLORATION OF PRESERVED SPECIMEN (
<figureCitation id="1880E1B9FFF7FF8EFDF7F9DDFD1353F0" box="[616,710,1650,1677]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
). Tergites in anterior half dark brown, posterior half light brown, posterior margin with thin dark brown band. Paratergites light brown with dark brown to blackish tips. Paratergite impressions and groove of thoracic shield light brown. Antennae brown, and legs and pleurites light brown to grey. Head laterally around eyes and at posterior margin dark brown, frontally light brown. Collum dark brown. Eyes green.
</paragraph>
<paragraph id="8004FD3CFFF7FF8FFF22F8ECFC9E5410" blockId="6.[189,1399,1858,2027]" lastBlockId="7.[189,1399,267,364]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
HEAD. Eyes with&gt;80 ocelli, median ocelli small and increasing in size towards lateral and posterior part of eyes (
<figureCitation id="1880E1B9FFF7FF8EFEFDF8C9FE1652FC" box="[354,451,1894,1920]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
), several larger ocelli on lateral margin clearly separated from eye. Organ of Tömösváry positioned in antennal-groove (
<figureCitation id="1880E1B9FFF7FF8EFD29F826FCC552D8" box="[694,784,1929,1956]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="6" pageNumber="7">Fig. 1B</figureCitation>
). Antennae short, protruding to coxa of third leg, not reaching margin of thoracic shield. Antennomere 1 as long as 2+3; antennomeres 2–5 of similar length; antennomere 6 as long as 4+5, but shorter than 1. Antennomeres 1–6 and antennal groove densely pubescent (
<figureCitation id="1880E1B9FFF6FF8FFEDBFEA4FE715459" box="[324,420,267,293]" captionStart="Fig.2" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,1079,1886]" captionTargetId="figure@8.[74,1398,1079,1887]" captionTargetPageId="8" captionText="Fig.2.Sphaeromimus kalambatritrasp.nov.,SEM ofleft antenna,♂, holotype (CASENT9068297-A). A. Overview. B.Antennal disc, detail. C. Margin of antennal disc, detail.Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensorycone; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118114/files/figure.png" pageId="7" pageNumber="8">Fig. 2A</figureCitation>
). Antennomere 6 towards disc with single row of sensilla basiconica (
<figureCitation id="1880E1B9FFF6FF8FFB77FEA4FABF5459" box="[1256,1386,267,293]" captionStart="Fig.2" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,1079,1886]" captionTargetId="figure@8.[74,1398,1079,1887]" captionTargetPageId="8" captionText="Fig.2.Sphaeromimus kalambatritrasp.nov.,SEM ofleft antenna,♂, holotype (CASENT9068297-A). A. Overview. B.Antennal disc, detail. C. Margin of antennal disc, detail.Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensorycone; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118114/files/figure.png" pageId="7" pageNumber="8">Fig. 2B–C</figureCitation>
). Disc in female with 33/31, in male with 53/55 apical cones, as well as several setae shorter, or as long as, apical cones (
<figureCitation id="1880E1B9FFF6FF8FFE1BFEFDFE345410" box="[388,481,338,364]" captionStart="Fig.2" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,1079,1886]" captionTargetId="figure@8.[74,1398,1079,1887]" captionTargetPageId="8" captionText="Fig.2.Sphaeromimus kalambatritrasp.nov.,SEM ofleft antenna,♂, holotype (CASENT9068297-A). A. Overview. B.Antennal disc, detail. C. Margin of antennal disc, detail.Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensorycone; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118114/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
). Margin of labrum with setae.
</paragraph>
<paragraph id="8004FD3CFFF6FF8FFF22FE3CFAA25721" blockId="7.[189,1405,403,606]" pageId="7" pageNumber="8">
GNATHOCHILARIUM. Gnathochilarium typical for members of the order 
<taxonomicName id="47BB86BFFFF6FF8FFC46FE3CFB4354D1" authorityName="Brandt" authorityYear="1833" box="[985,1174,403,429]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="order">Sphaerotheriida</taxonomicName>
, stipes and lamellamentum densely pubescent (
<figureCitation id="1880E1B9FFF6FF8FFD90FE19FDA554AC" box="[527,624,438,464]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="7" pageNumber="8">Figs 1B</figureCitation>
, 
<figureCitation id="1880E1B9FFF6FF8FFDE0FE19FD1F54AD" box="[639,714,438,465]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="7" pageNumber="8">3A, E</figureCitation>
). Lateral palpi rudimentary, not distinctly projecting over level of surrounding cuticle, consisting of four sensillae (
<figureCitation id="1880E1B9FFF6FF8FFCCBFE76FC7A5488" box="[852,943,473,500]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="7" pageNumber="8">Fig.3C</figureCitation>
). Inner palpi well developed, with field of sensory-cones and scale-like structures (
<figureCitation id="1880E1B9FFF6FF8FFD2CFE52FCC4576B" box="[691,785,509,535]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
). 
<collectingRegion id="427F33DEFFF6FF8FFCBAFE52FCAE576B" box="[805,891,509,535]" country="Fiji" name="Central" pageId="7" pageNumber="8">Central</collectingRegion>
pads (protuberance of endochilarium) with field of sensory-cones and smaller scale-like structures (
<figureCitation id="1880E1B9FFF6FF8FFCC2FD8FFC625747" box="[861,951,544,571]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="7" pageNumber="8">Fig.3E</figureCitation>
). Endochilarium with deep triangular incision between central pads, central pads projecting lamella-mentum nearly to base of inner palpi.
</paragraph>
<caption id="D4C4ADB4FFF6FF8FFF22F939FC515291" httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="7" pageNumber="8" targetBox="[189,1398,711,1646]" targetPageId="7">
<paragraph id="8004FD3CFFF6FF8FFF22F939FC515291" blockId="7.[189,1399,1686,2029]" pageId="7" pageNumber="8">
<emphasis id="B2CF212EFFF6FF8FFF22F939FEDF53CC" bold="true" box="[189,266,1686,1712]" pageId="7" pageNumber="8">Fig. 1.</emphasis>
<taxonomicName id="47BB86BFFFF6FF8FFE8EF939FDBD53CC" authority="Moritz &amp; Wesener, 2017" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[273,616,1686,1712]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kalambatritra" status="sp. nov.">
<emphasis id="B2CF212EFFF6FF8FFE8EF939FDBD53CC" box="[273,616,1686,1712]" italics="true" pageId="7" pageNumber="8">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9FC9C55FFF6FF8FFDF0F938FD1F53CD" box="[623,714,1687,1713]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B2CF212EFFF6FF8FFD4FF939FD3053CC" bold="true" box="[720,741,1686,1712]" pageId="7" pageNumber="8">A</emphasis>
. Lateral view. 
<emphasis id="B2CF212EFFF6FF8FFC0CF939FC1E53CC" bold="true" box="[915,971,1686,1712]" pageId="7" pageNumber="8">B–D</emphasis>
. Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). 
<emphasis id="B2CF212EFFF6FF8FFCB7F916FCE953AF" bold="true" box="[808,828,1721,1747]" pageId="7" pageNumber="8">B</emphasis>
. Head, outer fronto lateral view. 
<emphasis id="B2CF212EFFF6FF8FFB58F916FB0953AF" bold="true" box="[1223,1244,1721,1747]" pageId="7" pageNumber="8">C</emphasis>
. Head, inner posterior view of sclerotized structures only. 
<emphasis id="B2CF212EFFF6FF8FFD54F972FD35538B" bold="true" box="[715,736,1757,1783]" pageId="7" pageNumber="8">D</emphasis>
. Head, inner posterior view of sclerotized structures and soft structures. 
<emphasis id="B2CF212EFFF6FF8FFEEDF8AFFE535266" bold="true" box="[370,390,1792,1818]" pageId="7" pageNumber="8">E</emphasis>
. Segmentation of the left tentorium and nebententorium, frontal view. Abbreviations: aD = antennal disc; atm = antennal muscle; C = condylus of mandible gnathal lobe; eb = epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td = tendon; tm = tentorial muscle; to = Tömösváry organ; tt = tentorium; I–VI = antennomeres.
</paragraph>
</caption>
<paragraph id="8004FD3CFFF9FF80FF22FEA4FBB75435" blockId="8.[189,1398,267,329]" pageId="8" pageNumber="9">
Lateral endochilarium with densely packed median-pointing setae. Hypopharynx with single row of teeth lateral on both sides, anterior distinctly separated row of 8 teeth (
<figureCitation id="1880E1B9FFF9FF80FC67FE81FB815435" box="[1016,1108,302,329]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8004FD3CFFF9FF80FF22FEDFFDCA57D8" blockId="8.[189,1399,368,676]" pageId="8" pageNumber="9">
MANDIBLE. Mandible with typical shape of 
<taxonomicName id="47BB86BFFFF9FF80FD52FEDFFC5254F6" authorityName="Brandt" authorityYear="1833" box="[717,903,368,394]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">Sphaerotheriida</taxonomicName>
(
<figureCitation id="1880E1B9FFF9FF80FC05FEDFFC2254F6" box="[922,1015,368,394]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="8" pageNumber="9">Fig. 3F</figureCitation>
), inner tooth 3-combed, with 6 long pectinate lamellae (
<figureCitation id="1880E1B9FFF9FF80FE7AFE3CFD9354D1" box="[485,582,403,429]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="8" pageNumber="9">Fig. 3G</figureCitation>
), condylus with a single, lower step at its apex (
<figureCitation id="1880E1B9FFF9FF80FB0DFE3CFB3B54D1" box="[1170,1262,403,429]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="8" pageNumber="9">Fig. 3F</figureCitation>
). Condylus of mandible mounted against cuticular thickening on lateral margin of labrum, anteriorly of antennae (
<figureCitation id="1880E1B9FFF9FF80FF59FE76FEF45488" box="[198,289,473,500]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="8" pageNumber="9">Fig. 1C</figureCitation>
). Tentorium lacking connection to head capsule via transverse bar. Posterior process triangular and plate-like, laying parallel to plate-like gnathal lobe sclerite. Epipharyngeal bar broad and laminar, running within epipahryngeal wall in direction of mandible condylus, broadening distally. Epipharyngeal bar with short lateral offshoot. Hypopharyngeal bar rod-like, curved within hypopharyngeal wall. Distal tip of hypopharyngeal bar reaches plate-like nebententorium. Nebententorium oriented at right angle to hypopharyngeal bar (
<figureCitation id="1880E1B9FFF9FF80FE28FD26FDC457D8" box="[439,529,649,676]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="8" pageNumber="9">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8004FD3CFFF9FF80FF22FD64FCBC5799" blockId="8.[189,873,715,742]" box="[189,873,715,742]" pageId="8" pageNumber="9">COLLUM. Collum glabrous except for few setae at margins.</paragraph>
<paragraph id="8004FD3CFFF9FF80FF22FCA2FD935636" blockId="8.[189,1399,780,842]" pageId="8" pageNumber="9">
THORACIC SHIELD. Thoracic shield glabrous, with a chagrinated (leather-like) surface, few setae in grooves. Grooves deep (
<figureCitation id="1880E1B9FFF9FF80FE44FC9FFDED5636" box="[475,568,816,842]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8004FD3CFFF9FF80FF22FCDDFDD556AD" blockId="8.[189,1398,881,977]" pageId="8" pageNumber="9">
BODY RINGS. Tergites 3–12 with a chagrinated surface, small hair only present at posterior margin and in grooves, paratergite tips of mid-body tergites strongly projecting posteriorly (
<figureCitation id="1880E1B9FFF9FF80FBF5FC3BFB1256D2" box="[1130,1223,916,942]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
). Tergites with single black locking carina.
</paragraph>
<caption id="D4C4ADB4FFF9FF80FF22F825FBC35297" httpUri="https://zenodo.org/record/1118114/files/figure.png" pageId="8" pageNumber="9" targetBox="[189,1398,1079,1886]" targetPageId="8">
<paragraph id="8004FD3CFFF9FF80FF22F825FBC35297" blockId="8.[189,1398,1930,2027]" pageId="8" pageNumber="9">
<emphasis id="B2CF212EFFF9FF80FF22F825FEDC52D8" bold="true" box="[189,265,1930,1956]" pageId="8" pageNumber="9">Fig. 2.</emphasis>
<taxonomicName id="47BB86BFFFF9FF80FE90F825FDB352D8" authority="Moritz &amp; Wesener, 2017" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[271,614,1930,1956]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kalambatritra" status="sp. nov.">
<emphasis id="B2CF212EFFF9FF80FE90F825FDB352D8" box="[271,614,1930,1956]" italics="true" pageId="8" pageNumber="9">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9FC9C55FFF9FF80FDF3F824FD1D52D9" box="[620,712,1931,1957]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, SEM of left antenna, ♂, holotype (CASENT 9068297- A). 
<emphasis id="B2CF212EFFF9FF80FF22F802FF0752BB" bold="true" box="[189,210,1965,1991]" pageId="8" pageNumber="9">A</emphasis>
. Overview. 
<emphasis id="B2CF212EFFF9FF80FEFDF801FEA352B4" bold="true" box="[354,374,1966,1992]" pageId="8" pageNumber="9">B</emphasis>
. Antennal disc, detail. 
<emphasis id="B2CF212EFFF9FF80FD1CF802FD4D52BB" bold="true" box="[643,664,1965,1991]" pageId="8" pageNumber="9">C</emphasis>
. Margin of antennal disc, detail. Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensory cone; I–VI = antennomeres.
</paragraph>
</caption>
<caption id="D4C4ADB4FFF8FF81FF22F88EFC9F5290" httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="9" pageNumber="10" targetBox="[189,1399,438,1783]" targetPageId="9">
<paragraph id="8004FD3CFFF8FF81FF22F88EFC9F5290" blockId="9.[189,1398,1825,2028]" pageId="9" pageNumber="10">
<emphasis id="B2CF212EFFF8FF81FF22F88EFEDF5240" bold="true" box="[189,266,1825,1852]" pageId="9" pageNumber="10">Fig. 3.</emphasis>
<taxonomicName id="47BB86BFFFF8FF81FE8FF88DFDB25240" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[272,615,1826,1852]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalambatritra">
<emphasis id="B2CF212EFFF8FF81FE8FF88DFDB25240" box="[272,615,1826,1852]" italics="true" pageId="9" pageNumber="10">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
sp. nov., SEM of mouthparts, ♂, holotype (CASENT 9068297- A). 
<emphasis id="B2CF212EFFF8FF81FF22F8EAFF075223" bold="true" box="[189,210,1861,1887]" pageId="9" pageNumber="10">A</emphasis>
. Gnathochilarium overview, ventral. 
<emphasis id="B2CF212EFFF8FF81FD0EF8EAFD705223" bold="true" box="[657,677,1861,1887]" pageId="9" pageNumber="10">B</emphasis>
. Gnathochilarium overview, frontodorsal. 
<emphasis id="B2CF212EFFF8FF81FB3FF8EAFB605223" bold="true" box="[1184,1205,1861,1887]" pageId="9" pageNumber="10">C</emphasis>
. Lateral palpus, detail. 
<emphasis id="B2CF212EFFF8FF81FE94F8C7FEF552FE" bold="true" box="[267,288,1896,1922]" pageId="9" pageNumber="10">D</emphasis>
. Inner palpus, detail. 
<emphasis id="B2CF212EFFF8FF81FD85F8C7FDFB52FE" bold="true" box="[538,558,1896,1922]" pageId="9" pageNumber="10">E</emphasis>
. Central pad, detail. 
<emphasis id="B2CF212EFFF8FF81FC82F8C7FCFA52FE" bold="true" box="[797,815,1896,1922]" pageId="9" pageNumber="10">F</emphasis>
. Right mandible, overview. 
<emphasis id="B2CF212EFFF8FF81FBE9F8C7FB5852FE" bold="true" box="[1142,1165,1896,1922]" pageId="9" pageNumber="10">G</emphasis>
. Pectinate lamellae, detail. Abbreviations: C = condyles; cP = central pad; eT = exterior teeth; Hyp = hypopharynx; iP = inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium.
</paragraph>
</caption>
<paragraph id="8004FD3CFFFBFF82FF22FEA3FAA35434" blockId="10.[189,1398,267,329]" pageId="10" pageNumber="11">
ANAL SHIELD. Anal shield large, with a steep edge, entirely glabrous, with lighter and darker patches (
<figureCitation id="1880E1B9FFFBFF82FF59FE81FEF65435" box="[198,291,302,329]" captionStart="Fig. 1" captionStartId="7.[189,232,1686,1712]" captionTargetBox="[189,1398,711,1646]" captionTargetId="figure@7.[189,1398,711,1646]" captionTargetPageId="7" captionText="Fig. 1. Sphaeromimus kalambatritra sp. nov.A. Lateral view.B–D.Volume rendering of head based on µCT-images, female paratype (CASENT 9058301). B. Head, outer fronto lateral view. C. Head, inner posterior view of sclerotized structures only.D. Head, inner posterior view of sclerotized structures and soft structures. E. Segmentation of the left tentorium and nebententorium, frontal view.Abbreviations: aD = antennal disc; atm = antennal muscle; C =condylus of mandible gnathal lobe; eb =epipharyngeal bar; G = gula; gls = gnathal lobe sclerite; hb = hypopharyngeal bar; il = incisura lateralis; iP = inner palpi; lab = labium; LL = lamella lingualis; m = mandibular muscle; mdb = mandibular base; Me = mentum; mgl = mandible gnathal lobe; mp = molar plate; nt = nebententorium; oc = ocelli; PM = paramentum; pp = posterior process; St = stipes of gnathochilarium; td= tendon; tm =tentorial muscle; to =Tömösváry organ; tt = tentorium; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118112/files/figure.png" pageId="10" pageNumber="11">Fig. 1A</figureCitation>
). Underside with single black locking carina, located closer to tergite margins than to pleurite.
</paragraph>
<paragraph id="8004FD3CFFFBFF82FF22FEDFFAD254AC" blockId="10.[189,1406,368,464]" pageId="10" pageNumber="11">
ENDOTERGUM. Endotergum inner section with loose field of short, cone-shaped spines and long setae (
<figureCitation id="1880E1B9FFFBFF82FF59FE3CFEFD54D1" box="[198,296,403,429]" captionStart="Fig.4" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1401,1078,1882]" captionTargetId="figure@10.[189,1401,1078,1883]" captionTargetPageId="10" captionText="Fig.4.Sphaeromimuskalambatritra sp. nov., SEM of endotergum, ♂, holotype (CASENT9068297-A). A.Overview,endotergum.B. Marginofendotergum,detail.Abbreviations:(1)=innerpart; (2)=middle part; (3) =outer part." httpUri="https://zenodo.org/record/1118118/files/figure.png" pageId="10" pageNumber="11">Fig. 4A</figureCitation>
). Externally 2–3 dense rows of long marginal bristles, slightly protruding above margin of tergite. Bristles covered with small, triangular spines, apically increasing in density (
<figureCitation id="1880E1B9FFFBFF82FB02FE19FB2F54AC" box="[1181,1274,438,464]" captionStart="Fig.4" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1401,1078,1882]" captionTargetId="figure@10.[189,1401,1078,1883]" captionTargetPageId="10" captionText="Fig.4.Sphaeromimuskalambatritra sp. nov., SEM of endotergum, ♂, holotype (CASENT9068297-A). A.Overview,endotergum.B. Marginofendotergum,detail.Abbreviations:(1)=innerpart; (2)=middle part; (3) =outer part." httpUri="https://zenodo.org/record/1118118/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="8004FD3CFFFBFF82FF22FE57FBB35749" blockId="10.[189,1398,504,566]" pageId="10" pageNumber="11">
STIGMATIC PLATES. First stigma-carrying plate with a well-rounded projecting apex, apex covered with tiny spines and setae (
<figureCitation id="1880E1B9FFFBFF82FE20FDB4FDCB574A" box="[447,542,539,566]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Second plate (
<figureCitation id="1880E1B9FFFBFF82FD46FDB4FCE7574A" box="[729,818,539,566]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="10" pageNumber="11">Fig. 5B</figureCitation>
) without apex and spines.
</paragraph>
<paragraph id="8004FD3CFFFBFF82FF22FDF2FBD3570A" blockId="10.[189,1030,604,631]" box="[189,1030,604,631]" pageId="10" pageNumber="11">FIRST PLEURITE. First pleurite laterally well-rounded, without extensions.</paragraph>
<paragraph id="8004FD3CFFFBFF82FF22FD31FEC55614" blockId="10.[189,1405,670,872]" pageId="10" pageNumber="11">
LEGS. Leg 1 with 2, 2 with 4, 3 with 10 ventral spines and no apical spine. Leg pairs 4–21 with 12–16 ventral spines and an apical spine. Small coxa process developed, covered with field of spines (
<figureCitation id="1880E1B9FFFBFF82FA91FD6EFABC57A0" box="[1294,1385,705,732]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="10" pageNumber="11">Fig.5C</figureCitation>
). Femur 2.0, tarsus 3.3 times as long as wide. All podomeres with few setae (
<figureCitation id="1880E1B9FFFBFF82FBC5FD4BFB6C5783" box="[1114,1209,740,767]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
). Toothed ridge (cleaning comb?) of femur with&gt;40 teeth, reaching ca. 0.8 times length of femur. Coxa in anterior aspect basally with a row of teeth, similar to “cleaning comb” on femur. Pronounced coxal process absent.
</paragraph>
<paragraph id="8004FD3CFFFBFF82FF22FC3FFC1C56B1" blockId="10.[189,1398,911,973]" pageId="10" pageNumber="11">
MALE GONOPORE. Male gonopore typical for genus, plate glabrous, surrounded by relative large membranous area, few small spines basally of gonopore (
<figureCitation id="1880E1B9FFFBFF82FCC0FC1DFC6956B1" box="[863,956,946,973]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="10" pageNumber="11">Fig. 5B</figureCitation>
).
</paragraph>
<caption id="D4C4ADB4FFFBFF82FF22F827FE605295" httpUri="https://zenodo.org/record/1118118/files/figure.png" pageId="10" pageNumber="11" targetBox="[189,1401,1078,1882]" targetPageId="10">
<paragraph id="8004FD3CFFFBFF82FF22F827FE605295" blockId="10.[189,1398,1928,2025]" pageId="10" pageNumber="11">
<emphasis id="B2CF212EFFFBFF82FF22F827FEDC52DE" bold="true" box="[189,265,1928,1954]" pageId="10" pageNumber="11">Fig. 4.</emphasis>
<taxonomicName id="47BB86BFFFFBFF82FE90F827FDB052DE" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[271,613,1928,1954]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kalambatritra">
<emphasis id="B2CF212EFFFBFF82FE90F827FDB052DE" box="[271,613,1928,1954]" italics="true" pageId="10" pageNumber="11">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
sp. nov., SEM of endotergum, ♂, holotype (CASENT 9068297- A). 
<emphasis id="B2CF212EFFFBFF82FF22F804FF0752B9" bold="true" box="[189,210,1963,1989]" pageId="10" pageNumber="11">A</emphasis>
. Overview, endotergum. 
<emphasis id="B2CF212EFFFBFF82FE68F804FDDE52B9" bold="true" box="[503,523,1963,1989]" pageId="10" pageNumber="11">B</emphasis>
. Margin of endotergum, detail. Abbreviations: (1) = inner part; (2) = middle part; (3) = outer part.
</paragraph>
</caption>
<paragraph id="8004FD3CFFFAFF83FF22FEA3FCF454CF" blockId="11.[189,1406,267,435]" pageId="11" pageNumber="12">
ANTERIOR TELOPOD. Harp carrying five well-developed stridulation ribs (
<figureCitation id="1880E1B9FFFAFF83FBAAFEA4FB4C545A" box="[1077,1177,267,294]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="11" pageNumber="12">Fig. 5D</figureCitation>
). Shape usual for genus, with setae and tiny teeth at apical margin. Telopoditomere 4 with one large, triangular, apically weakly sclerotized spine and 2 smaller ones (
<figureCitation id="1880E1B9FFFAFF83FD45FEFDFC825411" box="[730,855,338,365]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="11" pageNumber="12">Fig. 5E–F</figureCitation>
); basally on its anterior side with several tiny teeth. Telopoditomere 3 with triangular hump laterally, located close to border to telopoditomere 4 and juxtaposed to process of telopoditomere 2 (
<figureCitation id="1880E1B9FFFAFF83FD27FE37FCC654CF" box="[696,787,408,435]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="11" pageNumber="12">Fig. 5E</figureCitation>
).
</paragraph>
<caption id="D4C4ADB4FFFAFF83FF22F953FB805296" httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="11" pageNumber="12" targetBox="[193,1396,571,1748]" targetPageId="11">
<paragraph id="8004FD3CFFFAFF83FF22F953FB805296" blockId="11.[189,1399,1788,2026]" pageId="11" pageNumber="12">
<emphasis id="B2CF212EFFFAFF83FF22F953FEDE526B" bold="true" box="[189,267,1788,1815]" pageId="11" pageNumber="12">Fig. 5.</emphasis>
<taxonomicName id="47BB86BFFFFAFF83FE8CF952FDBE526B" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[275,619,1789,1815]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kalambatritra">
<emphasis id="B2CF212EFFFAFF83FE8CF952FDBE526B" box="[275,619,1789,1815]" italics="true" pageId="11" pageNumber="12">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). 
<emphasis id="B2CF212EFFFAFF83FF22F88FFF075246" bold="true" box="[189,210,1824,1850]" pageId="11" pageNumber="12">A</emphasis>
. 1 
<taxonomicName id="47BB86BFFFFAFF83FF71F8B1FEF05246" box="[238,293,1822,1850]" pageId="11" pageNumber="12" rank="subSpecies" subSpecies="left">st left</taxonomicName>
coxae and stigma carrying plate. 
<emphasis id="B2CF212EFFFAFF83FD3AF88FFD6C5246" bold="true" box="[677,697,1824,1850]" pageId="11" pageNumber="12">B</emphasis>
. 2nd left coxa and stigma carrying plate. 
<emphasis id="B2CF212EFFFAFF83FB1CF88FFB4D5246" bold="true" box="[1155,1176,1824,1850]" pageId="11" pageNumber="12">C</emphasis>
. 9th left leg. 
<emphasis id="B2CF212EFFFAFF83FABBF88FFAEC5246" bold="true" box="[1316,1337,1824,1850]" pageId="11" pageNumber="12">D</emphasis>
. Left anterior telopod, anterior view. 
<emphasis id="B2CF212EFFFAFF83FDAFF8ECFD915221" bold="true" box="[560,580,1859,1885]" pageId="11" pageNumber="12">E</emphasis>
. Left anterior telopod, posterior view. 
<emphasis id="B2CF212EFFFAFF83FB95F8ECFBC95221" bold="true" box="[1034,1052,1859,1885]" pageId="11" pageNumber="12">F</emphasis>
. Left anterior telopod, lateral view. 
<emphasis id="B2CF212EFFFAFF83FE9CF8C9FECF52FC" bold="true" box="[259,282,1894,1920]" pageId="11" pageNumber="12">G</emphasis>
. Left posterior telopod, anterior view. 
<emphasis id="B2CF212EFFFAFF83FD7FF8C9FD2252FC" bold="true" box="[736,759,1894,1920]" pageId="11" pageNumber="12">H</emphasis>
. Left posterior telopod, posterior view. Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref = prefemur; sp = spine; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="D4C4ADB4FFFDFF84FF22FD59FBA25624" httpUri="https://zenodo.org/record/1118122/files/figure.png" pageId="12" pageNumber="13" targetBox="[189,1396,263,730]" targetPageId="12">
<paragraph id="8004FD3CFFFDFF84FF22FD59FBA25624" blockId="12.[189,1399,758,856]" pageId="12" pageNumber="13">
<emphasis id="B2CF212EFFFDFF84FF22FD59FEDB566D" bold="true" box="[189,270,758,785]" pageId="12" pageNumber="13">Fig. 6.</emphasis>
<taxonomicName id="47BB86BFFFFDFF84FE86FD58FDA1566D" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[281,628,759,785]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kalambatritra">
<emphasis id="B2CF212EFFFDFF84FE86FD58FDA1566D" box="[281,628,759,785]" italics="true" pageId="12" pageNumber="13">Sphaeromimus kalambatritra</emphasis>
</taxonomicName>
sp. nov., schematic drawing, ♀, paratype (CASENT 9058301). 
<emphasis id="B2CF212EFFFDFF84FF22FCB5FF075648" bold="true" box="[189,210,794,820]" pageId="12" pageNumber="13">A</emphasis>
. Left vulva (2nd coxa). 
<emphasis id="B2CF212EFFFDFF84FE75FCB5FE2B5648" bold="true" box="[490,510,794,820]" pageId="12" pageNumber="13">B</emphasis>
. Subanal plate. Abbreviations: cx = coxa; ep = exterior plate; ip = interior plate; op = operculum; pref = prefemur; sr = stridulation ribs. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8004FD3CFFFDFF84FF22FC08FEBB5100" blockId="12.[189,1405,934,1148]" pageId="12" pageNumber="13">
POSTERIOR TELOPOD. Podomere 3 slightly curved, 3.3 times as long as wide, slightly longer than immovable finger (
<figureCitation id="1880E1B9FFFDFF84FE8BFC63FE4C569B" box="[276,409,972,999]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="12" pageNumber="13">Fig.5G–H</figureCitation>
). Hollowed-out inner margin with two lobes and two spines, posterior aspect with ca 30 small crenulated teeth. Immovable finger straight, basally wide, apically tapering, only apical tip strongly curved towards podomere 3. Podomere 1 with few setae on lateral margin (
<figureCitation id="1880E1B9FFFDFF84FB10FBB8FB39514E" box="[1167,1260,1047,1074]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="12" pageNumber="13">Fig. 5H</figureCitation>
), podomere 2 only with few setae at anterior side, posterior side glabrous (
<figureCitation id="1880E1B9FFFDFF84FC3DFB93FBF2512B" box="[930,1063,1084,1111]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="12" pageNumber="13">Fig. 5G–H</figureCitation>
). Podomere 3 with only few marginal setae.
</paragraph>
<paragraph id="8004FD3CFFFDFF84FF22FB09FE87502A" blockId="12.[189,1406,1190,1366]" pageId="12" pageNumber="13">
FEMALE SEXUAL CHARACTERS. Vulva massive. Operculum well-rounded, protruding up to apical third of prefemur, with few marginal setae (
<figureCitation id="1880E1B9FFFDFF84FDF7FB64FD1C519A" box="[616,713,1227,1254]" captionStart="Fig. 6" captionStartId="12.[189,232,758,784]" captionTargetBox="[189,1396,263,730]" captionTargetId="figure@12.[189,1396,263,730]" captionTargetPageId="12" captionText="Fig. 6. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♀, paratype (CASENT 9058301). A. Left vulva (2 nd coxa). B. Subanal plate. Abbreviations: cx = coxa; ep = exterior plate; ip = interior plate; op = operculum; pref =prefemur; sr = stridulation ribs. Scale bars =1 mm." httpUri="https://zenodo.org/record/1118122/files/figure.png" pageId="12" pageNumber="13">Fig. 6A</figureCitation>
). Subanal plate large, with shallow invagination at apical margin (shape typical for genus). Washboard with ten stridulation ribs on each side, median in anterior half with black triangular field (
<figureCitation id="1880E1B9FFFDFF84FDA4FAB9FD42504D" box="[571,663,1302,1329]" captionStart="Fig. 6" captionStartId="12.[189,232,758,784]" captionTargetBox="[189,1396,263,730]" captionTargetId="figure@12.[189,1396,263,730]" captionTargetPageId="12" captionText="Fig. 6. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♀, paratype (CASENT 9058301). A. Left vulva (2 nd coxa). B. Subanal plate. Abbreviations: cx = coxa; ep = exterior plate; ip = interior plate; op = operculum; pref =prefemur; sr = stridulation ribs. Scale bars =1 mm." httpUri="https://zenodo.org/record/1118122/files/figure.png" pageId="12" pageNumber="13">Fig.6B</figureCitation>
). The female carried several hundred eggs with a diameter of 
<quantity id="474350D9FFFDFF84FF22FA94FE98502A" box="[189,333,1339,1366]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="12" pageNumber="13" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
.
</paragraph>
</subSubSection>
</subSection>
<subSubSection id="C8A1AEB7FFFDFF84FF22FAD0FD895369" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8004FD3CFFFDFF84FF22FAD0FE8350E5" blockId="12.[189,342,1407,1433]" box="[189,342,1407,1433]" pageId="12" pageNumber="13">
<heading id="DB4C4A50FFFDFF84FF22FAD0FE8350E5" bold="true" box="[189,342,1407,1433]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B2CF212EFFFDFF84FF22FAD0FE8350E5" bold="true" box="[189,342,1407,1433]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8004FD3CFFFDFF84FF22FA1FFD895369" blockId="12.[189,1399,1456,1558]" pageId="12" pageNumber="13">
Only known from the type-locality, the Réserve Spéciale de Kalambatritra, which is a mountainous rainforest (
<figureCitation id="1880E1B9FFFDFF84FEDEFA79FE4F508C" box="[321,410,1494,1520]" captionStart="Fig. 11" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[199,1399,624,1923]" captionTargetId="figure@19.[189,1399,624,1923]" captionTargetPageId="19" captionText="Fig. 11. Distribution of the genus Sphaeromimus de Saussure &amp; Zehntner, 1902 in south-eastern Madagascar. Map modified after Moat &amp; Smith 2007." httpUri="https://zenodo.org/record/1118132/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
). In the same habitat, two undetermined giant pill-millipede species of the genus 
<emphasis id="B2CF212EFFFDFF84FF22FA54FEB35369" box="[189,358,1531,1557]" italics="true" pageId="12" pageNumber="13">Zoosphaerium</emphasis>
occur sympatrically.
</paragraph>
</subSubSection>
</treatment>
</document>