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<document id="CDAE4BEB7DE2A5E4DF21CA082A374DC4" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D64242FFF482AA39F14B7BF99B" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liparocephalus litoralis Kirschenblatt 1938" docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="256" updateTime="1702375545460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B83F14DEBD48A4D6FA59976B5D0946ED">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:name id="E322F60F64BF70AC40807670C8106FDE" type="personal">
<mods:role id="D6134F30410FE8F573A4733E4DA5E98F">
<mods:roleTerm id="E994198562A3C5ABB60F315818BA85A9">Author</mods:roleTerm>
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<mods:namePart id="291BA4BC935E48DFED3FF04333066CC7">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="9A445B8D01631D39F7E6594C89B81BEE">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="1859E8E6A91AF727491BD6850D9C16AA">Ono, Hiroki</mods:namePart>
<mods:affiliation id="42DD39E7ABD3CC28821EA7282B3A6C53">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
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<mods:namePart id="5CC8685D8F89CE4DACBADCB7C59AAD23">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="07DD99AD0317B20D3881F79B991CD54A">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:title id="F97D10909ED4F8B6A2F611D7075E67F0">Zootaxa</mods:title>
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<mods:part id="F3B59FBAE5F690D0FE34C3315B81728B">
<mods:date id="3A4E13DF488801D6B75FE542E70E14B2">2023</mods:date>
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<mods:number id="53BFDED702C4E39CEBD4E32B4E9049E2">2023-12-12</mods:number>
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<mods:number id="42579A062D8A8902F9744F1334EF1D40">3</mods:number>
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<treatment id="03FE87D64242FFF482AA39F14B7BF99B" ID-DOI="http://doi.org/10.5281/zenodo.10361600" ID-Zenodo-Dep="10361600" LSID="urn:lsid:plazi:treatment:03FE87D64242FFF482AA39F14B7BF99B" httpUri="http://treatment.plazi.org/id/03FE87D64242FFF482AA39F14B7BF99B" lastPageNumber="256" pageId="5" pageNumber="256">
<subSubSection id="C34D654B4242FFF482AA39F14C73FC42" pageId="5" pageNumber="256" type="nomenclature">
<paragraph id="8BE836C04242FFF482AA39F14CADFC1E" blockId="5.[151,694,929,1028]" box="[151,694,929,956]" pageId="5" pageNumber="256">
<heading id="D0A081AC4242FFF482AA39F14CADFC1E" bold="true" box="[151,694,929,956]" fontSize="11" level="1" pageId="5" pageNumber="256" reason="1">
<taxonomicName id="4C574D434242FFF482AA39F14CADFC1E" ID-CoL="3V9XJ" authority="Kirschenblatt, 1938" authorityName="Kirschenblatt" authorityYear="1938" box="[151,694,929,956]" class="Insecta" family="Staphylinidae" genus="Liparocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="B923EAD24242FFF482AA39F14CADFC1E" bold="true" box="[151,694,929,956]" pageId="5" pageNumber="256">
<emphasis id="B923EAD24242FFF482AA39F14FA2FC1E" bold="true" box="[151,441,929,956]" italics="true" pageId="5" pageNumber="256">Liparocephalus litoralis</emphasis>
<bibRefCitation id="EFC64B314242FFF483FD39F14CADFC1E" author="Kirschenblatt, J. D." box="[448,694,929,956]" pageId="5" pageNumber="256" pagination="527 - 536" refId="ref21740" refString="Kirschenblatt, J. D. (1938) On some Far Eastern staphylinid beetles. Trudy Gidrobiologichesk. oy Ekspeditsii (Zoo /. Inst. Akad. Nauk SSSR 1934 g. na Yapanskoye More), 1, 527 - 536. [in Russian with English summary]" type="journal article" year="1938">Kirschenblatt, 1938</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C04242FFF482AA39964C73FC42" blockId="5.[151,694,929,1028]" box="[151,616,966,992]" pageId="5" pageNumber="256">[Japanese name: Hotei-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B4242FFF482AA39BA4F8AFBA6" box="[151,401,1002,1028]" pageId="5" pageNumber="256" type="description">
<paragraph id="8BE836C04242FFF482AA39BA4F8AFBA6" blockId="5.[151,694,929,1028]" box="[151,401,1002,1028]" pageId="5" pageNumber="256">
(
<figureCitation id="136C2A454242FFF482A239BA4F19FBA6" box="[159,258,1002,1028]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1386,191,1910]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2 Aedeagal parameres of Liparocephalus, Diaulota and Rotundicephala gen. n. spp. A &amp; D, parameres of aedeagus (A, L. tokunagai; D, D. aokii); B, C, E, F &amp; G, apical lobe of parameres (B, L. litoralis; C, L. tokunagai; E, D. aokii; F, D. uenoi; G, R. koheii sp. n.). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361570" httpUri="https://zenodo.org/record/10361570/files/figure.png" pageId="5" pageNumber="256">Figs. 2B</figureCitation>
;
<figureCitation id="136C2A454242FFF4833339BA4F93FBA6" box="[270,392,1002,1028]" captionStart="FIGURE 3" captionStartId="4.[151,250,1910,1934]" captionTargetBox="[162,1426,192,1875]" captionTargetId="figure-22@4.[151,1436,181,1886]" captionTargetPageId="4" captionText="FIGURE 3 Habitus and genitalia of Liparocephalus. A–C, habitus (A, L. litoralis female; B, L. tokunagai, male; C, L. tokunagai, female); D &amp; F, median lobe of aedeagus (D, L. litoralis; F, L. tokunagai); E &amp; G, female spermatheca (E, L. litoralis; G, L. tokunagai). Scale bars: 1.0 mm for A–C; 0.1 mm for D–G." figureDoi="http://doi.org/10.5281/zenodo.10361572" httpUri="https://zenodo.org/record/10361572/files/figure.png" pageId="5" pageNumber="256">3A, D &amp; E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4242FFF482AA3E7E4ADDFBC7" pageId="5" pageNumber="256" type="reference_group">
<paragraph id="8BE836C04242FFF482AA3E7E4ADDFBC7" blockId="5.[151,1436,1070,1125]" pageId="5" pageNumber="256">
<treatmentCitationGroup id="AB4711EE4242FFF482AA3E7E4ADDFBC7" pageId="5" pageNumber="256">
<treatmentCitation id="0AF610D14242FFF482AA3E7E4CB8FBE4" author="Kirschenblatt, J. D." box="[151,675,1070,1094]" page="532" pageId="5" pageNumber="256" year="1938">
<taxonomicName id="4C574D434242FFF482AA3E7E4CB8FBE4" ID-CoL="3V9XJ" authority="Kirschenblatt, 1938: 532" authorityName="Kirschenblatt" authorityPageNumber="532" authorityYear="1938" box="[151,675,1070,1094]" class="Insecta" family="Staphylinidae" genus="Liparocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="B923EAD24242FFF482AA3E7E4F8AFBE4" box="[151,401,1070,1094]" italics="true" pageId="5" pageNumber="256">Liparocephalus litoralis</emphasis>
<bibRefCitation id="EFC64B314242FFF483A03E7E4CB8FBE4" author="Kirschenblatt, J. D." box="[413,675,1070,1094]" pageId="5" pageNumber="256" pagination="527 - 536" refId="ref21740" refString="Kirschenblatt, J. D. (1938) On some Far Eastern staphylinid beetles. Trudy Gidrobiologichesk. oy Ekspeditsii (Zoo /. Inst. Akad. Nauk SSSR 1934 g. na Yapanskoye More), 1, 527 - 536. [in Russian with English summary]" type="journal article" year="1938">Kirschenblatt, 1938: 532</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="54EC88624242FFF481A93E7F4DDBFBE5" box="[916,960,1071,1095]" pageId="5" pageNumber="256">type</typeStatus>
locality: Bering Island (Komandor Islands); Maruyama &amp;
<treatmentCitation id="0AF610D14242FFF483693E1D4FE9FBC7" author="Ahn, K. - J. &amp; Maruyama, M." box="[340,498,1101,1125]" page="568" pageId="5" pageNumber="256" year="2000">
<bibRefCitation id="EFC64B314242FFF483693E1D4FE9FBC7" author="Ahn, K. - J. &amp; Maruyama, M." box="[340,498,1101,1125]" pageId="5" pageNumber="256" pagination="359 - 364" refId="ref21121" refString="Ahn, K. - J. &amp; Maruyama, M. (2000) Paramblopusa eoa (Coleoptera: Staphylinidae: Aleocharinae), a new intertidal species from Kuril Islands. Coleopterists Bulletin, 54 (2), 359 - 364. https: // doi. org / 10.1649 / 0010 - 065 X (2000) 054 [0359: PEANIS] 2.0. CO; 2" type="journal article" year="2000">Ahn, 2000: 568</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="0AF610D14242FFF480A63E1D4D58FBC7" author="Haga, K." box="[667,835,1101,1125]" page="216" pageId="5" pageNumber="256" year="2022">
<bibRefCitation id="EFC64B314242FFF480A63E1D4D58FBC7" author="Haga, K." box="[667,835,1101,1125]" pageId="5" pageNumber="256" pagination="216 - 218" refId="ref21657" refString="Haga, K. (2022) A new locality of the intertidal rove beetle Liparocaphalus litoralis Kirschenblatt (Coleoptera, Staphylinidae, Aleocharinae) in Nemuro Subprefecture, Hokkaido, Japan. Elytra, New Series, 12 (2), 216 - 218." type="journal article" year="2022">Haga, 2022: 216</bibRefCitation>
</treatmentCitation>
(additional records; biological notes).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4242FFF482AA3EC34A0BFB57" pageId="5" pageNumber="256" type="materials_examined">
<paragraph id="8BE836C04242FFF482AA3EC34A0BFB57" blockId="5.[151,1437,1171,1594]" pageId="5" pageNumber="256">
<emphasis id="B923EAD24242FFF482AA3EC34F9DFB0F" bold="true" box="[151,390,1171,1197]" pageId="5" pageNumber="256">Material examined.</emphasis>
<materialsCitation id="3B3F3C9D4242FFF483AF3EC34F6AFB73" collectingDate="1999-09-12" collectionCode="KUM" collectorName="M. Maruyama" country="Japan" location="Mochirippu" municipality="Hamanaka-cho" pageId="5" pageNumber="256" specimenCount="1" specimenCount-female="1" stateProvince="Hokkaido">
<emphasis id="B923EAD24242FFF483AF3EC34C6BFB0F" bold="true" box="[402,624,1171,1197]" pageId="5" pageNumber="256">
<collectingCountry id="F34076504242FFF483AF3EC34FC4FB0F" box="[402,479,1171,1197]" name="Japan" pageId="5" pageNumber="256">Japan</collectingCountry>
:
<collectingRegion id="4993F8224242FFF483CD3EC34C70FB0F" box="[496,619,1171,1197]" country="Japan" name="Hokkaido" pageId="5" pageNumber="256">Hokkaidô</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494242FFF480413EC34CFEFB0F" box="[636,741,1171,1197]" count="1" pageId="5" pageNumber="256" type="female">1 female</specimenCount>
, 2 unsexed,
<collectingMunicipality id="6B8CACBA4242FFF481433EC34A31FB0F" box="[894,1066,1171,1197]" pageId="5" pageNumber="256">Hamanaka-chô</collectingMunicipality>
,
<location id="8E88601B4242FFF486073EC34ADAFB0F" LSID="urn:lsid:plazi:treatment:03FE87D64242FFF482AA39F14B7BF99B:8E88601B4242FFF486073EC34ADAFB0F" box="[1082,1217,1171,1197]" country="Japan" municipality="Hamanaka-cho" name="Mochirippu" pageId="5" pageNumber="256" stateProvince="Hokkaido">Mochirippu</location>
,
<date id="FFE910004242FFF486EF3EC34B77FB0F" box="[1234,1388,1171,1198]" pageId="5" pageNumber="256" value="1999-09-12">
<collectingDate id="EFADE9E84242FFF486EF3EC34B77FB0F" box="[1234,1388,1171,1198]" pageId="5" pageNumber="256" value="1999-09-12">12. IX. 1999</collectingDate>
</date>
,
<collectorName id="26A253164242FFF487413EC44F0BFB70" pageId="5" pageNumber="256">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE054242FFF4831F3EE74F72FB73" box="[290,361,1207,1233]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="256" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4242FFF483423EE74B39FB73" box="[383,1314,1207,1234]" collectingDate="2000-09-28" collectionCode="KUM" collectorName="M. Maruyama" country="Japan" location="Mochirippu" municipality="Hamanaka-cho" pageId="5" pageNumber="256" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Hokkaido">
<specimenCount id="9D51FD494242FFF483423EE74FC1FB73" box="[383,474,1207,1234]" count="5" pageId="5" pageNumber="256" type="male">5 males</specimenCount>
,
<specimenCount id="9D51FD494242FFF483D53EE74C42FB73" box="[488,601,1207,1234]" count="5" pageId="5" pageNumber="256" type="female">5 females</specimenCount>
, 67 unsexed, ditto,
<date id="FFE910004242FFF481063EE74DCBFB73" box="[827,976,1207,1234]" pageId="5" pageNumber="256" value="2000-09-28">
<collectingDate id="EFADE9E84242FFF481063EE74DCBFB73" box="[827,976,1207,1234]" pageId="5" pageNumber="256" value="2000-09-28">28. IX. 2000</collectingDate>
</date>
,
<collectorName id="26A253164242FFF481E03EE84A9BFB70" box="[989,1152,1208,1234]" pageId="5" pageNumber="256">M. Maruyama</collectorName>
(
<collectionCode id="ED46AE054242FFF486AC3EE74ACCFB73" box="[1169,1239,1207,1233]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="256" type="Herbarium">KUM</collectionCode>
, cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4242FFF4870D3EE74CA4FB54" collectionCode="VII" collectorName="Bettoga &amp; Tengu-iwa" country="Japan" location="Nemuro-shi" pageId="5" pageNumber="256" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Hokkaido">
<specimenCount id="9D51FD494242FFF4870D3EE74B9AFB73" box="[1328,1409,1207,1233]" count="1" pageId="5" pageNumber="256" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD494242FFF487B33EE74EEAFB57" count="4" pageId="5" pageNumber="256" type="female">4 females</specimenCount>
,
<location id="8E88601B4242FFF482C13E8B4F9EFB57" LSID="urn:lsid:plazi:treatment:03FE87D64242FFF482AA39F14B7BF99B:8E88601B4242FFF482C13E8B4F9EFB57" box="[252,389,1243,1269]" country="Japan" name="Nemuro-shi" pageId="5" pageNumber="256" stateProvince="Hokkaido">Nemuro-shi</location>
,
<collectorName id="26A253164242FFF483AD3E8B4FF1FB57" box="[400,490,1243,1269]" pageId="5" pageNumber="256">Bettôga</collectorName>
,
<collectorName id="26A253164242FFF483C83E8B4C74FB57" box="[501,623,1243,1269]" pageId="5" pageNumber="256">Tengu-iwa</collectorName>
,
<date id="FFE910004242FFF480473E8B4CA4FB54" box="[634,703,1243,1270]" pageId="5" pageNumber="256" value="2021-07-02">
2.
<collectionCode id="ED46AE054242FFF480A83E8C4CA4FB54" box="[661,703,1244,1270]" pageId="5" pageNumber="256">VII</collectionCode>
</date>
</materialsCitation>
. 2021, K. Haga (
<collectionCode id="ED46AE054242FFF481413E8B4DD9FB57" box="[892,962,1243,1269]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="5" pageNumber="256" type="Herbarium">KUM</collectionCode>
, cYT).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4242FFF482FA3EAF4FD5F9B7" pageId="5" pageNumber="256" type="diagnosis">
<paragraph id="8BE836C04242FFF482FA3EAF4AE1FA27" blockId="5.[151,1437,1171,1594]" pageId="5" pageNumber="256">
<emphasis id="B923EAD24242FFF482FA3EAF4F59FABB" bold="true" box="[199,322,1279,1305]" pageId="5" pageNumber="256">Diagnosis.</emphasis>
This species can be easily distinguished from all other Japanese species of the tribe
<taxonomicName id="4C574D434242FFF486D63EAF4B87FABB" authorityName="Casey" authorityYear="1894" box="[1259,1436,1279,1305]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="tribe" tribe="Liparocephalini">Liparocephalini</taxonomicName>
by its large, broad, and blackish body (
<figureCitation id="136C2A454242FFF4805C3F734CA5FA9F" box="[609,702,1315,1341]" captionStart="FIGURE 3" captionStartId="4.[151,250,1910,1934]" captionTargetBox="[162,1426,192,1875]" captionTargetId="figure-22@4.[151,1436,181,1886]" captionTargetPageId="4" captionText="FIGURE 3 Habitus and genitalia of Liparocephalus. A–C, habitus (A, L. litoralis female; B, L. tokunagai, male; C, L. tokunagai, female); D &amp; F, median lobe of aedeagus (D, L. litoralis; F, L. tokunagai); E &amp; G, female spermatheca (E, L. litoralis; G, L. tokunagai). Scale bars: 1.0 mm for A–C; 0.1 mm for D–G." figureDoi="http://doi.org/10.5281/zenodo.10361572" httpUri="https://zenodo.org/record/10361572/files/figure.png" pageId="5" pageNumber="256">Fig. 3A</figureCitation>
). This species is similar to
<taxonomicName id="4C574D434242FFF4863F3F744B52FA9C" authority="Maklin, 1853" authorityName="Maklin" authorityYear="1853" box="[1026,1353,1315,1342]" class="Insecta" family="Staphylinidae" genus="Liparocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B923EAD24242FFF4863F3F744A09FA9F" box="[1026,1042,1316,1341]" italics="true" pageId="5" pageNumber="256">L</emphasis>
.
<emphasis id="B923EAD24242FFF4861F3F734ABFFA9F" box="[1058,1188,1315,1341]" italics="true" pageId="5" pageNumber="256">brevipennis</emphasis>
Mäklin, 1853
</taxonomicName>
and
<taxonomicName id="4C574D434242FFF487BB3F744FF8FAC3" authority="Klimaszewski, 2021" authorityName="Klimaszewski" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Liparocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="humblei">
<emphasis id="B923EAD24242FFF487BB3F744B8DFA9F" box="[1414,1430,1316,1341]" italics="true" pageId="5" pageNumber="256">L</emphasis>
.
<emphasis id="B923EAD24242FFF482AA3F174EEAFAC3" box="[151,241,1351,1377]" italics="true" pageId="5" pageNumber="256">humblei</emphasis>
<bibRefCitation id="EFC64B314242FFF482C13F174FF8FAC3" author="Klimaszewski, J. &amp; Brunke, A. &amp; Sikes, D. S. &amp; Pentinsaari, M. &amp; Godin, B. &amp; Webster, R. P. &amp; Davies, A. &amp; Bourdon, C. &amp; Newton, A. F." box="[252,483,1351,1377]" pageId="5" pageNumber="256" pagination="323 - 337" refId="ref21924" refString="Klimaszewski, J., Brunke, A., Sikes, D. S., Pentinsaari, M., Godin, B., Webster, R. P., Davies, A., Bourdon, C. &amp; Newton, A. F. (2021) Tribe Liparocephalini Fenyes, 1918. In: Klimaszewski, J., Brunke, A., Sikes, D. S., Pentinsaari, M., Godin, B., Webster, R. P., Davies, A., Bourdon, C. &amp; Newton, A. F. (Eds.), A Faunal Review of Aleocharine Beetles in the Rapidly Changing Arctic and Subarctic Regions of North America (Coleoptera, Staphylinidae). Springer, Cham, pp. 323 - 337. https: // doi. org / 10.1007 / 978 - 3 - 030 - 68191 - 3 _ 16" type="book chapter" year="2021">Klimaszewski, 2021</bibRefCitation>
</taxonomicName>
known from the eastern Pacific coast, but distinguished from them by the genital shape (see,
<bibRefCitation id="EFC64B314242FFF483243F3B4C5AFA27" author="Klimaszewski, J. &amp; Brunke, A. &amp; Sikes, D. S. &amp; Pentinsaari, M. &amp; Godin, B. &amp; Webster, R. P. &amp; Davies, A. &amp; Bourdon, C. &amp; Newton, A. F." box="[281,577,1387,1414]" pageId="5" pageNumber="256" pagination="323 - 337" refId="ref21924" refString="Klimaszewski, J., Brunke, A., Sikes, D. S., Pentinsaari, M., Godin, B., Webster, R. P., Davies, A., Bourdon, C. &amp; Newton, A. F. (2021) Tribe Liparocephalini Fenyes, 1918. In: Klimaszewski, J., Brunke, A., Sikes, D. S., Pentinsaari, M., Godin, B., Webster, R. P., Davies, A., Bourdon, C. &amp; Newton, A. F. (Eds.), A Faunal Review of Aleocharine Beetles in the Rapidly Changing Arctic and Subarctic Regions of North America (Coleoptera, Staphylinidae). Springer, Cham, pp. 323 - 337. https: // doi. org / 10.1007 / 978 - 3 - 030 - 68191 - 3 _ 16" type="book chapter" year="2021">
Klimaszewski
<emphasis id="B923EAD24242FFF483823F3C4FEBFA27" box="[447,496,1387,1413]" italics="true" pageId="5" pageNumber="256">et al</emphasis>
., 2021
</bibRefCitation>
). These species have no clear sexual dimorphism in head size.
</paragraph>
<paragraph id="8BE836C04242FFF482FA3FDF4FD5F9B7" blockId="5.[151,1437,1171,1594]" pageId="5" pageNumber="256">
<emphasis id="B923EAD24242FFF482FA3FDF4F31FA0B" bold="true" box="[199,298,1423,1449]" pageId="5" pageNumber="256">Habitat.</emphasis>
This species is found inhabiting the seashore under stone and rock crevices (Maruyama &amp;
<bibRefCitation id="EFC64B314242FFF487223FDF4B83FA0B" author="Ahn, K. - J. &amp; Maruyama, M." box="[1311,1432,1423,1449]" pageId="5" pageNumber="256" pagination="359 - 364" refId="ref21121" refString="Ahn, K. - J. &amp; Maruyama, M. (2000) Paramblopusa eoa (Coleoptera: Staphylinidae: Aleocharinae), a new intertidal species from Kuril Islands. Coleopterists Bulletin, 54 (2), 359 - 364. https: // doi. org / 10.1649 / 0010 - 065 X (2000) 054 [0359: PEANIS] 2.0. CO; 2" type="journal article" year="2000">Ahn, 2000</bibRefCitation>
;
<bibRefCitation id="EFC64B314242FFF482AA3FE44F06FA6F" author="Haga, K." box="[151,285,1459,1486]" pageId="5" pageNumber="256" pagination="216 - 218" refId="ref21657" refString="Haga, K. (2022) A new locality of the intertidal rove beetle Liparocaphalus litoralis Kirschenblatt (Coleoptera, Staphylinidae, Aleocharinae) in Nemuro Subprefecture, Hokkaido, Japan. Elytra, New Series, 12 (2), 216 - 218." type="journal article" year="2022">Haga, 2022</bibRefCitation>
). Due to its restricted habitat, confined to a few coastal localities in the eastern
<collectingRegion id="4993F8224242FFF486AB3FE34B1CFA6F" box="[1174,1287,1459,1485]" country="Japan" name="Hokkaido" pageId="5" pageNumber="256">Hokkaidô</collectingRegion>
,
<collectingCountry id="F34076504242FFF4872F3FE44B4EFA6C" box="[1298,1365,1460,1486]" name="Japan" pageId="5" pageNumber="256">Japan</collectingCountry>
, it has been categorized as a “Data Deficient species (DD)” in “The 4
<superScript id="7C229B884242FFF4816D3F874D46FA47" attach="left" box="[848,861,1495,1509]" fontSize="6" pageId="5" pageNumber="256">th</superScript>
Version of the Japanese Red Lists” (
<bibRefCitation id="EFC64B314242FFF486CE3F874FD9F9B7" author="Ministry of the Environment &amp; Japan" pageId="5" pageNumber="256" refId="ref22526" refString="Ministry of the Environment, Japan (2020) The 4 th version of the Japanese red lists (Insects) [online]. Available from: http: // www. env. go. jp / press / 107905. html (accessed on 9 June 2023)." type="url" year="2020">
Ministry of the Environment,
<collectingCountry id="F34076504242FFF4830A3FAC4F61F9B4" box="[311,378,1532,1558]" name="Japan" pageId="5" pageNumber="256">Japan</collectingCountry>
, 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4242FFF482FA3C4F4B7BF99B" box="[199,1376,1567,1594]" pageId="5" pageNumber="256" type="distribution">
<paragraph id="8BE836C04242FFF482FA3C4F4B7BF99B" blockId="5.[151,1437,1171,1594]" box="[199,1376,1567,1594]" pageId="5" pageNumber="256">
<emphasis id="B923EAD24242FFF482FA3C4F4F7AF99B" bold="true" box="[199,353,1567,1593]" pageId="5" pageNumber="256">Distribution.</emphasis>
<collectingCountry id="F34076504242FFF4835A3C704FB0F998" box="[359,427,1568,1594]" name="Japan" pageId="5" pageNumber="256">Japan</collectingCountry>
:
<collectingRegion id="4993F8224242FFF4838B3C4F4C33F99B" box="[438,552,1567,1593]" country="Japan" name="Hokkaido" pageId="5" pageNumber="256">Hokkaidô</collectingRegion>
;
<collectingCountry id="F34076504242FFF4800E3C4F4C99F99B" box="[563,642,1567,1593]" name="Russia" pageId="5" pageNumber="256">Russia</collectingCountry>
: Kuril Islands (Matua Island), Komandor Islands (Bering Island).
</paragraph>
</subSubSection>
</treatment>
</document>
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