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<document id="1EE498D1C53FAF359490AA830A9152B3" ID-CLB-Dataset="4251" ID-DOI="10.11646/palaeoentomology.4.1.2" ID-GBIF-Dataset="e6da253b-3709-43cb-8ba9-12b1dd02727a" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507667" ID-ZooBank="BBBA32F8-0EA5-4069-A84C-7612D875DB0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631643950535" checkinUser="carolina" docAuthor="WAGNER, RÜDIGER, HOFFEINS, CHRISTEL &amp; HOFFEINS, HANS WERNER" docDate="2021" docId="03E93A18FF804A205099FD5F81B5FBCA" docLanguage="en" docName="palaeoentomology.4.1.2.pdf" docOrigin="Palaeoentomology 4 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.1.2" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Hoffeinsodes inexpectatus WAGNER &amp; HOFFEINS &amp; HOFFEINS 2021, comb. nov." docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFD04260FF814A225011FFEB805AFFDD" masterDocTitle="Re-description of Hoffeinsodes inexpectatus (Wagner, 2012) (Psychodidae, Bruchomyiinae) from Baltic amber" masterLastPageNumber="18" masterPageNumber="15" pageNumber="16" updateTime="1699448706747" updateUser="ExternalLinkService">
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<mods:title id="A84C5FC90642A00DC61AD6857E3A81E6">Re-description of Hoffeinsodes inexpectatus (Wagner, 2012) (Psychodidae, Bruchomyiinae) from Baltic amber</mods:title>
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<mods:namePart id="E8B4FECA7C729DA8BAED08B064828A12">WAGNER, RÜDIGER</mods:namePart>
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<mods:namePart id="B09189F0614A71BBBA036CB0B5EB0F40">HOFFEINS, CHRISTEL</mods:namePart>
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<mods:title id="BED3B3D4ED05A1B0B226F0BBC6C68BB3">Palaeoentomology</mods:title>
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<mods:date id="33B579574BBCCBBE717A7BD702C75286">2021</mods:date>
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<treatment id="03E93A18FF804A205099FD5F81B5FBCA" ID-DOI="http://doi.org/10.5281/zenodo.5515379" ID-GBIF-Taxon="188164750" ID-Zenodo-Dep="5515379" LSID="urn:lsid:plazi:treatment:03E93A18FF804A205099FD5F81B5FBCA" httpUri="http://treatment.plazi.org/id/03E93A18FF804A205099FD5F81B5FBCA" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
<subSubSection id="C35AD885FF804A235099FD5F80EDFD2C" pageId="1" pageNumber="16" type="nomenclature">
<paragraph id="8BFF8B0EFF804A235099FD5F80EDFD2C" blockId="1.[136,778,581,790]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A235099FD5F80EDFD2C" bold="true" pageId="1" pageNumber="16">
Species
<taxonomicName id="4C40F08DFF804A2350F2FD5F82EDFD11" authority="(Wagner 2012)" authorityName="WAGNER &amp; HOFFEINS &amp; HOFFEINS" authorityYear="2021" baseAuthorityName="Wagner" baseAuthorityYear="2012" box="[227,695,692,716]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="inexpectatus" status="comb. nov.">
<emphasis id="B934571CFF804A2350F2FD5F825CFD11" bold="true" box="[227,518,692,716]" italics="true" pageId="1" pageNumber="16">Hoffeinsodes inexpectatus</emphasis>
(
<bibRefCitation id="EFD1F6FFFF804A235205FD5F82EAFD11" author="Wagner, R. &amp; Stuckenberg, B. R." box="[532,688,692,716]" pageId="1" pageNumber="16" pagination="355 - 367" refId="ref2547" refString="Wagner, R. &amp; Stuckenberg, B. R. (2012) New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae, Bruchomyiinae). African Invertebrates, 53, 355 - 367. https: // doi. org / 10.5733 / afin. 053.0119" type="journal article" year="2012">Wagner 2012</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A207EA67FF804A2352D4FD5F80EDFD2C" pageId="1" pageNumber="16" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C35AD885FF804A235099FD158083FCCB" box="[136,217,766,790]" pageId="1" pageNumber="16" type="description">
<paragraph id="8BFF8B0EFF804A235099FD158083FCCB" blockId="1.[136,778,581,790]" box="[136,217,766,790]" pageId="1" pageNumber="16">
(
<figureCitation id="137B978BFF804A235081FD158094FCCB" box="[144,206,766,790]" captionStart="FIGURE 1" captionStartId="1.[136,235,1892,1916]" captionTargetBox="[151,1436,1017,1869]" captionTargetId="figure-293@1.[151,1436,1017,1869]" captionTargetPageId="1" captionText="FIGURE 1. Hoffeinsodes inexpectatus. A, Male, latero-dorsal view. B, Wing. Abbreviations: Scsubcosta; R, radius 1 to 5; M, media 1 to 4; CuA, anterior cubitus; CuP, posterior cubitus. C, Rear part of abdomen with terminalia. D, Drawing of abdomen segment VIII and terminalia. Abbreviations: St VIII, sternite 8; T VIII, Tergite VIII; cla, clasp along epandrium edge; cer, cerci; epa, epandrium; epaa, epandrium appendages; gcxc, connate gonocoxites; gst, gonostylus; lgst, lobe of gonostylus. Scale bars in A and B = 1mm, C and D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5507669" httpUri="https://zenodo.org/record/5507669/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35AD885FF804A235099FCA08269FC59" pageId="1" pageNumber="16" type="reference_group">
<paragraph id="8BFF8B0EFF804A235099FCA08246FC82" blockId="1.[136,563,842,901]" box="[136,540,842,864]" pageId="1" pageNumber="16">
=
<taxonomicName id="4C40F08DFF804A23508DFCA18246FC82" ID-CoL="8CYT7" authority="Wagner, 2012" authorityName="Wagner" authorityYear="2012" box="[156,540,842,864]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B934571CFF804A23508DFCA181C8FCBD" box="[156,402,842,864]" italics="true" pageId="1" pageNumber="16">Nemopalpus inexpectatus</emphasis>
<bibRefCitation id="EFD1F6FFFF804A235186FCA08246FC82" author="Wagner, R. &amp; Stuckenberg, B. R." box="[407,540,842,864]" pageId="1" pageNumber="16" pagination="355 - 367" refId="ref2547" refString="Wagner, R. &amp; Stuckenberg, B. R. (2012) New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae, Bruchomyiinae). African Invertebrates, 53, 355 - 367. https: // doi. org / 10.5733 / afin. 053.0119" type="journal article" year="2012">Wagner, 2012</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFF8B0EFF804A235099FC9B8269FC59" blockId="1.[136,563,842,901]" box="[136,563,879,901]" pageId="1" pageNumber="16">
=
<taxonomicName id="4C40F08DFF804A23508DFC848269FC59" ID-CoL="9HM55" authority="Wagner, 2017" authorityName="Wagner" authorityYear="2017" box="[156,563,879,901]" class="Insecta" family="Psychodidae" genus="Palaeosycorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B934571CFF804A23508DFC8481F2FC58" box="[156,424,879,901]" italics="true" pageId="1" pageNumber="16">Palaeosycorax inexpectatus</emphasis>
<bibRefCitation id="EFD1F6FFFF804A2351BFFC9B8269FC59" author="Wagner, R." box="[430,563,879,901]" pageId="1" pageNumber="16" pagination="100 - 120" refId="ref2646" refString="Wagner, R. (2017) Synopsis of extinct Bruchomyiinae (Diptera, Psychodidae) from Burmese, Baltic and Dominican amber, with descriptions of new genera and species. Zootaxa, 4320 (1), 100 - 120. https: // doi. org / 10.11646 / zootaxa. 4320.1.6" type="journal article" year="2017">Wagner, 2017</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C35AD885FF804A235099FC5C852BFE26" pageId="1" pageNumber="16" type="materials_examined">
<paragraph id="8BFF8B0EFF804A235099FC5C84C8FF22" blockId="1.[136,778,951,975]" lastBlockId="1.[810,1452,159,975]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A235099FC5C80ADFC12" bold="true" box="[136,247,951,975]" pageId="1" pageNumber="16">
<typeStatus id="54FB35ACFF804A235099FC5C80A8FC12" box="[136,242,951,975]" pageId="1" pageNumber="16" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, collection no. GZG.BST. 22886 (old no. Z 3900) Baltic amber—Late Eocene (lodged in the amber collection of the Geowissenschaftliches Zentrum der Universität Göttingen,
<collectingCountry id="F357CB9EFF804A23540EFF0C84DDFF22" box="[1055,1159,231,255]" name="Germany" pageId="1" pageNumber="16">Germany</collectingCountry>
).
</paragraph>
<paragraph id="8BFF8B0EFF804A23534EFEE0852BFE26" blockId="1.[810,1452,159,975]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A23534EFEE08393FEFE" bold="true" box="[863,969,267,291]" pageId="1" pageNumber="16">Material.</emphasis>
<materialsCitation id="3B288153FF804A2353C0FEE08537FE26" ID-GBIF-Occurrence="3353677301" collectionCode="SDEI" pageId="1" pageNumber="16" specimenCode="CCHH 711-2" specimenCount="2" specimenCount-male="2">
An additional
<specimenCount id="9D464087FF804A235460FEE784BAFEFE" box="[1137,1248,267,291]" pageId="1" pageNumber="16" type="male">two males</specimenCount>
in a single amber piece,
<specimenCount id="9D464087FF804A235362FEDB8382FE9A" box="[883,984,303,327]" pageId="1" pageNumber="16" type="male">one male</specimenCount>
complete; the second male lacks parts of the left side of the thorax and the fore abdomen segments, terminalia complete;
<specimenCode id="DBE62375FF804A235401FE9C84C5FE52" box="[1040,1183,375,399]" collectionCode="CCHH" pageId="1" pageNumber="16">CCHH 711-2</specimenCode>
<collectionCode id="ED5113CBFF804A2354B9FE9C84B9FE52" box="[1192,1251,375,399]" pageId="1" pageNumber="16">SDEI</collectionCode>
inventory number Dip 00899 (
<figureCitation id="137B978BFF804A2353BCFE7083B4FE6E" box="[941,1006,411,435]" captionStart="FIGURE 1" captionStartId="1.[136,235,1892,1916]" captionTargetBox="[151,1436,1017,1869]" captionTargetId="figure-293@1.[151,1436,1017,1869]" captionTargetPageId="1" captionText="FIGURE 1. Hoffeinsodes inexpectatus. A, Male, latero-dorsal view. B, Wing. Abbreviations: Scsubcosta; R, radius 1 to 5; M, media 1 to 4; CuA, anterior cubitus; CuP, posterior cubitus. C, Rear part of abdomen with terminalia. D, Drawing of abdomen segment VIII and terminalia. Abbreviations: St VIII, sternite 8; T VIII, Tergite VIII; cla, clasp along epandrium edge; cer, cerci; epa, epandrium; epaa, epandrium appendages; gcxc, connate gonocoxites; gst, gonostylus; lgst, lobe of gonostylus. Scale bars in A and B = 1mm, C and D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5507669" httpUri="https://zenodo.org/record/5507669/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
). The amber piece with the inclusions is embedded in a rectangular block of polyester resin (Voss- Chemie) (
<bibRefCitation id="EFD1F6FFFF804A235382FE088462FE26" author="Hoffeins, H. W." box="[915,1080,483,507]" pageId="1" pageNumber="16" pagination="215 - 219" refId="ref2028" refString="Hoffeins, H. W. (2001) On the preparation and conservation of amber inclusions in artificial resin. Polskie Pismo Entomologiczne, 70, 215 - 219." type="journal article" year="2001">Hoffeins, 2001</bibRefCitation>
), with size 12 × 34 ×
<quantity id="4CB826EBFF804A23550EFE088537FE26" box="[1311,1389,483,507]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="16" unit="mm" value="56.0">56 mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35AD885FF804A23534EFDEC85D2FD72" pageId="1" pageNumber="16" type="diagnosis">
<paragraph id="8BFF8B0EFF804A23534EFDEC85D2FD72" blockId="1.[810,1452,159,975]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A23534EFDEC8417FDC2" bold="true" box="[863,1101,519,543]" pageId="1" pageNumber="16">Emended diagnosis.</emphasis>
A
<taxonomicName id="4C40F08DFF804A235493FDEC857CFDC2" authorityName="Macquart" authorityYear="1838" box="[1154,1318,519,543]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="subFamily" subFamily="Bruchomyiinae">Bruchomyiinae</taxonomicName>
species in genus
<taxonomicName id="4C40F08DFF804A235364FDC0845AFD9E" authorityName="Wagner" authorityYear="2017" box="[885,1024,555,579]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B934571CFF804A235364FDC0845AFD9E" box="[885,1024,555,579]" italics="true" pageId="1" pageNumber="16">Hoffeinsodes</emphasis>
</taxonomicName>
, gonocoxites large, connate with tips turned dorsally, gonostyli crescent shaped, epandrium with a pair of central bilobed appendages, and a pair of sclerites along lateral edge of epandrium, apically with robust tip.
</paragraph>
</subSubSection>
<subSubSection id="C35AD885FF804A20534EFD5081B5FBCA" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="description">
<paragraph id="8BFF8B0EFF804A23534EFD50846DFC76" blockId="1.[810,1452,159,975]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A23534EFD5083B0FD0E" bold="true" box="[863,1002,699,723]" pageId="1" pageNumber="16">Description.</emphasis>
Head is as typical for the subfamily with roundish eyes and without eye bridge. antenna about as long as abdomen. Length of scapus, pedicel and flagellomeres: 0.07-0.10-0.25-0.18-0.19-0.16-0.16-0.16-0.14-0.14-0.11- 0.11-0.12-0.11-0.11-
<quantity id="4CB826EBFF804A235415FCA0843EFCBE" box="[1028,1124,843,867]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="16" unit="mm" value="0.11">0.11 mm</quantity>
.Ascoids not discernable (small or indistinguishable from hairs). Length of palpus segments: 0.12-0.14-
<quantity id="4CB826EBFF804A235389FC788469FC76" box="[920,1075,915,939]" metricMagnitude="-4" metricUnit="m" metricValue="3.45" metricValueMax="5.2" metricValueMin="1.7" pageId="1" pageNumber="16" unit="mm" value="0.345" valueMax="0.52" valueMin="0.17">0.17-0.52 mm</quantity>
.
</paragraph>
<paragraph id="8BFF8B0EFF804A20534EFC5C8098FEB6" blockId="1.[810,1452,159,975]" lastBlockId="2.[136,778,159,1047]" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
Thorax: wing about 3.5 times longer than wide; wing length:
<quantity id="4CB826EBFF834A2050CBFF748164FF6A" box="[218,318,159,183]" metricMagnitude="-3" metricUnit="m" metricValue="3.67" pageId="2" pageNumber="17" unit="mm" value="3.67">3.67 mm</quantity>
. Basis partly hidden by debris. Wing apex is between R
<subScript id="17C4894BFF834A205104FF388147FF3C" attach="left" box="[277,285,211,225]" fontSize="6" pageId="2" pageNumber="17">4</subScript>
and R
<subScript id="17C4894BFF834A205174FF388137FF3C" attach="left" box="[357,365,211,225]" fontSize="6" pageId="2" pageNumber="17">5</subScript>
. Radial fork distad of medial fork, R
<subScript id="17C4894BFF834A205313FF388350FF3C" attach="left" box="[770,778,211,225]" fontSize="6" pageId="2" pageNumber="17">2</subScript>
is approximately 0.73 the length of R
<subScript id="17C4894BFF834A205230FF1C8263FED8" attach="left" box="[545,569,247,261]" fontSize="6" pageId="2" pageNumber="17">2+3</subScript>
, CuA and CuP run parallel to wing margin. Metathorax with an oval sclerite dorsally behind halters with a row of small setae along apical edge.
</paragraph>
<caption id="DF3FDB86FF804A235099F88F81A4F7D1" ID-DOI="http://doi.org/10.5281/zenodo.5507669" ID-Zenodo-Dep="5507669" httpUri="https://zenodo.org/record/5507669/files/figure.png" pageId="1" pageNumber="16" startId="1.[136,235,1892,1916]" targetBox="[151,1436,1017,1869]" targetPageId="1">
<paragraph id="8BFF8B0EFF804A235099F88F81A4F7D1" blockId="1.[136,1452,1892,2061]" pageId="1" pageNumber="16">
<emphasis id="B934571CFF804A235099F88F815CF8A1" bold="true" box="[136,262,1892,1916]" pageId="1" pageNumber="16">FIGURE 1.</emphasis>
<emphasis id="B934571CFF804A23511FF88E827AF8A1" box="[270,544,1893,1917]" italics="true" pageId="1" pageNumber="16">
<taxonomicName id="4C40F08DFF804A23511FF88E8246F8A1" authorityName="WAGNER &amp; HOFFEINS &amp; HOFFEINS" authorityYear="2021" baseAuthorityName="Wagner" baseAuthorityYear="2012" box="[270,540,1893,1917]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="inexpectatus">Hoffeinsodes inexpectatus</taxonomicName>
.
</emphasis>
<emphasis id="B934571CFF804A235236F88F8263F8A1" bold="true" box="[551,569,1892,1916]" pageId="1" pageNumber="16">A</emphasis>
, Male, latero-dorsal view.
<emphasis id="B934571CFF804A235341F88E833BF8A0" bold="true" box="[848,865,1893,1917]" pageId="1" pageNumber="16">B</emphasis>
, Wing. Abbreviations: Scsubcosta; R, radius 1 to 5; M, media 1 to 4; CuA, anterior cubitus; CuP, posterior cubitus.
<emphasis id="B934571CFF804A2352EBF8638356F87D" bold="true" box="[762,780,1928,1952]" pageId="1" pageNumber="16">C</emphasis>
, Rear part of abdomen with terminalia.
<emphasis id="B934571CFF804A2354BEF862849BF87C" bold="true" box="[1199,1217,1929,1953]" pageId="1" pageNumber="16">D</emphasis>
, Drawing of abdomen segment VIII and terminalia. Abbreviations: St VIII, sternite 8; T VIII, Tergite VIII; cla, clasp along epandrium edge; cer, cerci; epa, epandrium; epaa, epandrium appendages; gcxc, connate gonocoxites; gst, gonostylus; lgst, lobe of gonostylus. Scale bars in
<emphasis id="B934571CFF804A235099F81F80C0F7D1" bold="true" box="[136,154,2036,2060]" pageId="1" pageNumber="16">A</emphasis>
and
<emphasis id="B934571CFF804A2350DDF81E8087F7D0" bold="true" box="[204,221,2037,2061]" pageId="1" pageNumber="16">B</emphasis>
= 1mm,
<emphasis id="B934571CFF804A235129F81F8110F7D1" bold="true" box="[312,330,2036,2060]" pageId="1" pageNumber="16">C</emphasis>
and
<emphasis id="B934571CFF804A23516DF81E81D4F7D0" bold="true" box="[380,398,2037,2061]" pageId="1" pageNumber="16">D</emphasis>
= 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BFF8B0EFF834A2050ACFE9C82E5FCE2" blockId="2.[136,778,159,1047]" pageId="2" pageNumber="17">
Abdomen with eight segments and terminalia (
<figureCitation id="137B978BFF834A2052A0FE9C80C7FE6E" captionStart="FIGURE 1" captionStartId="1.[136,235,1892,1916]" captionTargetBox="[151,1436,1017,1869]" captionTargetId="figure-293@1.[151,1436,1017,1869]" captionTargetPageId="1" captionText="FIGURE 1. Hoffeinsodes inexpectatus. A, Male, latero-dorsal view. B, Wing. Abbreviations: Scsubcosta; R, radius 1 to 5; M, media 1 to 4; CuA, anterior cubitus; CuP, posterior cubitus. C, Rear part of abdomen with terminalia. D, Drawing of abdomen segment VIII and terminalia. Abbreviations: St VIII, sternite 8; T VIII, Tergite VIII; cla, clasp along epandrium edge; cer, cerci; epa, epandrium; epaa, epandrium appendages; gcxc, connate gonocoxites; gst, gonostylus; lgst, lobe of gonostylus. Scale bars in A and B = 1mm, C and D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5507669" httpUri="https://zenodo.org/record/5507669/files/figure.png" pageId="2" pageNumber="17">Fig. 1A, C</figureCitation>
); first segment not markedly smaller than the following; inversion of the terminalia by torsion of segments 8 and 9 each by 90°. Terminalia (
<figureCitation id="137B978BFF834A2051D0FE088267FE26" box="[449,573,483,507]" captionStart="FIGURE 1" captionStartId="1.[136,235,1892,1916]" captionTargetBox="[151,1436,1017,1869]" captionTargetId="figure-293@1.[151,1436,1017,1869]" captionTargetPageId="1" captionText="FIGURE 1. Hoffeinsodes inexpectatus. A, Male, latero-dorsal view. B, Wing. Abbreviations: Scsubcosta; R, radius 1 to 5; M, media 1 to 4; CuA, anterior cubitus; CuP, posterior cubitus. C, Rear part of abdomen with terminalia. D, Drawing of abdomen segment VIII and terminalia. Abbreviations: St VIII, sternite 8; T VIII, Tergite VIII; cla, clasp along epandrium edge; cer, cerci; epa, epandrium; epaa, epandrium appendages; gcxc, connate gonocoxites; gst, gonostylus; lgst, lobe of gonostylus. Scale bars in A and B = 1mm, C and D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5507669" httpUri="https://zenodo.org/record/5507669/files/figure.png" pageId="2" pageNumber="17">Fig. 1C, D</figureCitation>
) with voluminous gonocoxites connate in middle and with hypandrium. In lateral view gonocoxites strongly curved, thinner in middle with broad dark appendages turned towards epandrium; along apical edge with dark setae. Broad, light and crescent shaped gonostyli located between central protrusion and lateral arc of gonocoxites; tip rectangular and black. Laterally between gonocoxites and epandrium run thinly sclerotized elongate lobes, apically with few setulae in whorls, probably descending from the gonostyli. Aedeagus not visible.
</paragraph>
<paragraph id="8BFF8B0EFF834A2050ACFCA081B5FBCA" blockId="2.[136,778,159,1047]" pageId="2" pageNumber="17">Epandrium trough shaped, rectangular in lateral view; parallel to lateral edges of epandrium run dark clasps terminating in a robust tip turned upwards. A pair of bilobed balloon-like appendages along apical edge of epandrium between tips of lateral clasps (difficult to identify). Proctiger small; cerci oval, pale and setose.</paragraph>
</subSubSection>
</treatment>
</document>