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<mods:title id="875BFDF7DD700E49E62D67731F3806FA">An Exceptional Large Sample Of The Early Miocene Ctenodactyline Rodent Sayimys Giganteus, Specific Variation And Taxonomic Implications</mods:title>
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<mods:namePart id="0E58B2A2DF4BCC9AEE49B28EE2529601">Hartman, Julian</mods:namePart>
<mods:affiliation id="B3CD6918D9A2AD5A8CE8D146C9378CA0">Oranjepoldererf 36, 2807 NE Gouda, the Netherlands; e-mail: juulhartman @ gmail. com. &amp; Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="AA9C607DD555B77ED7D1CA350F28471F">Van De Weerd, Andrew A.</mods:namePart>
<mods:affiliation id="B77399D6C6C15A3A8588B41F938F2932">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="5B0024DD09C57221021D3E98C69CDDE4">Bruijn, Hans De</mods:namePart>
<mods:affiliation id="F4ED6E48136C402AA41D0E14F9A2118C">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:namePart id="3869687B7B035B2F2B3B57D085A13CCF">Wessels, Wilma</mods:namePart>
<mods:affiliation id="2678D5BA3A3A0B797161500171E8ED8A">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<paragraph id="8BF009018C39617CFF55FAAE7DBCFA5B" blockId="13.[178,717,1383,1408]" box="[178,717,1383,1408]" pageId="13" pageNumber="372">
<heading id="D0B8BE6D8C39617CFF55FAAE7DBCFA5B" box="[178,717,1383,1408]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="372" reason="2">
<emphasis id="B93BD5138C39617CFF55FAAE7EC7FA5B" bold="true" box="[178,438,1383,1408]" pageId="13" pageNumber="372">
<emphasis id="B93BD5138C39617CFF55FAAE7E15FAA4" bold="true" box="[178,356,1383,1408]" italics="true" pageId="13" pageNumber="372">
<taxonomicName id="4C4F72828C39617CFF55FAAE7E7DFA5B" authorityName="WOOD" authorityYear="1937" box="[178,268,1384,1408]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
baskini
</emphasis>
LÓPEZ-
</emphasis>
<emphasis id="B93BD5138C39617CFE51FAAE7DBCFA5B" bold="true" box="[438,717,1384,1408]" pageId="13" pageNumber="372">ANTOÑANZAS et SEN, 2003</emphasis>
</heading>
</paragraph>
<paragraph id="8BF009018C39617CFF41FA5E7D6CFA6B" blockId="13.[166,541,1431,1456]" box="[166,541,1431,1456]" pageId="13" pageNumber="372">H o l o t y p e. An isolated right P4.</paragraph>
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<paragraph id="8BF009018C39617CFF41FA017E29FA24" blockId="13.[129,767,1479,1536]" pageId="13" pageNumber="372">
Ty p e l o c a l i t y. Y747, Potwar Plateau,
<collectingCountry id="F35849918C39617CFD94FA017DA0FA04" box="[627,721,1479,1503]" name="Pakistan" pageId="13" pageNumber="372">Pakistan</collectingCountry>
; age of the site: 18.3 Ma.
</paragraph>
<paragraph id="8BF009018C39617CFF41F9D17D8DF9F4" blockId="13.[129,764,1559,1616]" box="[166,764,1559,1583]" pageId="13" pageNumber="372">
Additional material from the
<typeStatus id="54F4B7A38C39617CFDA1F9D17D0EF9F4" box="[582,639,1559,1583]" pageId="13" pageNumber="372">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BF009018C39617CFF66F9F17D8AF98B" blockId="13.[129,764,1559,1616]" box="[129,763,1591,1616]" pageId="13" pageNumber="372">52 cheek teeth, representing all tooth positions except DP3.</paragraph>
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<materialsCitation id="3B27035C8C39617CFF41F9A17E89F93B" collectionCode="GSP" country="Pakistan" location="According" municipality="Potwar Plateau" pageId="13" pageNumber="372" specimenCode="GSP116" specimenCount="1">
O t h e r l o c a l i t i e s w i t h
<emphasis id="B93BD5138C39617CFE1AF9A17D82F9A4" box="[509,755,1639,1663]" italics="true" pageId="13" pageNumber="372">S a y i m y s b a s k i n i</emphasis>
.
<location id="8E905FDA8C39617CFF66F9417F83F944" LSID="urn:lsid:plazi:treatment:03E6B8178C39617CFF55FAAE7CDDFD4A:8E905FDA8C39617CFF66F9417F83F944" box="[129,242,1671,1695]" country="Pakistan" municipality="Potwar Plateau" name="According" pageId="13" pageNumber="372">According</location>
to
<bibRefCitation id="EFDE74F08C39617CFEF2F9417DFFF944" author="Lopez-Antonanzas, R. &amp; Sen, S." box="[277,654,1671,1695]" pageId="13" pageNumber="372" pagination="521 - 529" refId="ref16639" refString="Lopez-Antonanzas, R., Sen, S. (2003): Systematic revision of Mio-Pliocene Ctenodactylidae (Mammalia, Rodentia) from the Indian subcontinent. - Eclogae Geologicae Helvetiae, 96: 521 - 529." type="journal article" year="2003">López-Antoñanzas and Sen (2003)</bibRefCitation>
: site Y721
<collectingMunicipality id="6B94937B8C39617CFF66F96E7E54F91B" box="[129,293,1704,1728]" pageId="13" pageNumber="372">Potwar Plateau</collectingMunicipality>
,
<collectingCountry id="F35849918C39617CFED6F96E7EFAF91B" box="[305,395,1704,1728]" name="Pakistan" pageId="13" pageNumber="372">Pakistan</collectingCountry>
and site H.-
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(Banda Daud Shah, the
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locality of
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.
<emphasis id="B93BD5138C39617CFE4AF90F7E9CF93B" box="[429,493,1737,1760]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
)
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.
</paragraph>
<paragraph id="8BF009018C39617CFF41F93E7CDDFD4A" blockId="13.[129,767,1784,2100]" lastBlockId="13.[843,1482,151,658]" pageId="13" pageNumber="372">
D i s c u s s i o n. The collections from the type locality Y747 and from Y721 were studied by
<bibRefCitation id="EFDE74F08C39617CFDD3F8DE7DA3F8EB" author="Baskin, J." box="[564,722,1816,1840]" pageId="13" pageNumber="372" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin (1996)</bibRefCitation>
. He concluded that the specimens from these two sites may represent a new species, but refrained from naming it because awaiting additional material of
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<taxonomicName id="4C4F72828C39617CFDCFF8BC7D0EF84A" authorityName="WOOD" authorityYear="1937" box="[552,639,1914,1937]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
minor
</emphasis>
from its type locality in the Murree Formation at Banda Daud Shah. However,
<bibRefCitation id="EFDE74F08C39617CFED3F87C7DDAF809" author="Lopez-Antonanzas, R. &amp; Sen, S." box="[308,683,1978,2002]" pageId="13" pageNumber="372" pagination="521 - 529" refId="ref16639" refString="Lopez-Antonanzas, R., Sen, S. (2003): Systematic revision of Mio-Pliocene Ctenodactylidae (Mammalia, Rodentia) from the Indian subcontinent. - Eclogae Geologicae Helvetiae, 96: 521 - 529." type="journal article" year="2003">López-Antoñanzas and Sen (2003)</bibRefCitation>
did not have this restraint, and described the species
<emphasis id="B93BD5138C39617CFD86F81D7D1FF829" box="[609,622,2011,2034]" italics="true" pageId="13" pageNumber="372">S</emphasis>
.
<emphasis id="B93BD5138C39617CFD9BF81D7DB8F828" box="[636,713,2011,2035]" italics="true" pageId="13" pageNumber="372">baskini</emphasis>
with type locality Y747, and included specimens from Y721. Not having studied the collection from
<collectingCountry id="F35849918C39617CFDFDF7DA7D05F7EF" box="[538,628,2076,2100]" name="Pakistan" pageId="13" pageNumber="372">Pakistan</collectingCountry>
themselves,
<bibRefCitation id="EFDE74F08C39617CFCACFF5E7BB8FF6B" author="Lopez-Antonanzas, R. &amp; Sen, S." box="[843,1225,151,176]" pageId="13" pageNumber="372" pagination="521 - 529" refId="ref16639" refString="Lopez-Antonanzas, R., Sen, S. (2003): Systematic revision of Mio-Pliocene Ctenodactylidae (Mammalia, Rodentia) from the Indian subcontinent. - Eclogae Geologicae Helvetiae, 96: 521 - 529." type="journal article" year="2003">López-Antoñanzas and Sen (2003)</bibRefCitation>
refer to
<bibRefCitation id="EFDE74F08C39617CFAC9FF517ABBFF74" author="Baskin, J." box="[1326,1482,151,175]" pageId="13" pageNumber="372" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin (1996)</bibRefCitation>
for figures, description and measurements. The selection of the P4 as
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is unfortunate, as it is morphologically the most variable element in large
<taxonomicName id="4C4F72828C39617CFB05FF3E7A48FED4" authorityName="WOOD" authorityYear="1937" box="[1250,1337,248,271]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">
<emphasis id="B93BD5138C39617CFB05FF3E7A48FED4" box="[1250,1337,248,271]" italics="true" pageId="13" pageNumber="372">Sayimys</emphasis>
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populations. Regrettably, the published figures in
<bibRefCitation id="EFDE74F08C39617CFB1CFEDE7AEAFEEB" author="Baskin, J." box="[1275,1435,280,304]" pageId="13" pageNumber="372" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin (1996)</bibRefCitation>
are simplified line drawings, inadequate for a good evaluation of dental morphology. Baskin writes that the single unworn dp4 has a short mesolophid, its hypolophid is transversely directed, lower molars lack a mesolophid, the “M12 lack an anteroloph”, thus the paraflexus is absent and proto- and anteroloph are fused. Unfortunately, the only M12 of
<emphasis id="B93BD5138C39617CFC8DFE3C7C06FDCA" box="[874,887,506,529]" italics="true" pageId="13" pageNumber="372">S</emphasis>
.
<emphasis id="B93BD5138C39617CFC61FE3C7CB7FDCA" box="[902,966,506,529]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
is rather worn, so the anteroloph is completely fused with the protoloph. We conclude that without more material from the type locality of
<emphasis id="B93BD5138C39617CFB56FDFC7BCFFD8A" box="[1201,1214,570,593]" italics="true" pageId="13" pageNumber="372">S</emphasis>
.
<emphasis id="B93BD5138C39617CFB2CFDFC7A7AFD8A" box="[1227,1291,570,593]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
that can ascertain the specific difference,
<emphasis id="B93BD5138C39617CFBB7FD9C7B2CFDAA" box="[1104,1117,602,625]" italics="true" pageId="13" pageNumber="372">S</emphasis>
.
<emphasis id="B93BD5138C39617CFB97FD9C7BCCFDA9" box="[1136,1213,602,626]" italics="true" pageId="13" pageNumber="372">baskini</emphasis>
is a junior synonym of
<emphasis id="B93BD5138C39617CFCACFDBC7C29FD4A" box="[843,856,634,657]" italics="true" pageId="13" pageNumber="372">S</emphasis>
.
<emphasis id="B93BD5138C39617CFC82FDBD7CD4FD49" box="[869,933,635,658]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
.
</paragraph>
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