387 lines
50 KiB
XML
387 lines
50 KiB
XML
<document id="1D8C21B7F5C263DA7EA4862E04795A3C" ID-DOI="10.5281/zenodo.5176567" ID-ISSN="1942-1354" ID-Zenodo-Dep="5176567" ID-ZooBank="845665BE-16F9-4391-A361-FA74183A1ED0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628329558323" checkinUser="felipe" docAuthor="Edmonds, W. D. & Figueroa, Luis" docDate="2013" docId="03D81D59FFB2FFEAC5C97270247A7A01" docLanguage="en" docName="InsectaMundi.0313.1-4.pdf" docOrigin="Insecta Mundi 2013 (313)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Anomiopus pishtaco Edmonds & Figueroa 2013, new species" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="FFE16521FFB0FFEFC509770D2526785A" masterDocTitle="A Remarkable New Anomiopus Westwood from Peru (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1699044458418" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="B135E1EB497D67097AE9B144B4FFDE90" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1BE848B61CBFA4A07FFC61C6E5348FED">
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<mods:title id="06E2F652073A643BEE19B38F5998E27E">A Remarkable New Anomiopus Westwood from Peru (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="1DDAF516F44D8199A1009A0F41EB6ACC" type="personal">
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<mods:namePart id="8FD4C5757E08BC80A6A9B0C13800EC53">Edmonds, W. D.</mods:namePart>
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<mods:roleTerm id="D525DDF36126486C89C959D9F5584FFC">Author</mods:roleTerm>
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<mods:namePart id="C3A2543CDFAE86E9691504626CAF4214">Figueroa, Luis</mods:namePart>
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<mods:typeOfResource id="B1D72EBEC938B9FA9C6C990B9A39633B">text</mods:typeOfResource>
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<mods:title id="8985D9EC099B74EBB8631F0098C9165F">Insecta Mundi</mods:title>
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<mods:part id="DEC80EB2DBB54EBD2AD82AFF5EA972A0">
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<mods:date id="2BF8175902A84C7765150F4817BFCC91">2013</mods:date>
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<mods:number id="856F507ADB677B7315393384309B629C">2013-06-28</mods:number>
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<mods:detail id="B457E0DBA7EB37A2792DF2938BE366C5" type="volume">
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<mods:number id="EF62E4D22B12906A17BBB05EF614F0FB">2013</mods:number>
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<mods:detail id="180730F8849743FB52E7A637E67D9F76" type="issue">
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<mods:number id="8E0EB2D104EEEFF38A8AD792D152832C">313</mods:number>
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<mods:classification id="EFDD840EE9689677D73B7CF0FBB3D4A2">journal article</mods:classification>
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<mods:identifier id="2605AEB2640744D2FEC906069DFE654C" type="DOI">10.5281/zenodo.5176567</mods:identifier>
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<mods:identifier id="1B61500BECBECA61353174BEDF0B8065" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="CB35B12FE498EC299F657F7101946203" type="Zenodo-Dep">5176567</mods:identifier>
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</mods:mods>
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<treatment id="03D81D59FFB2FFEAC5C97270247A7A01" LSID="urn:lsid:plazi:treatment:03D81D59FFB2FFEAC5C97270247A7A01" httpUri="http://treatment.plazi.org/id/03D81D59FFB2FFEAC5C97270247A7A01" lastPageId="5" lastPageNumber="4" pageId="2" pageNumber="1">
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<subSubSection id="C36BFFC4FFB2FFEDC5C9727027527DCF" box="[192,628,1404,1429]" pageId="2" pageNumber="1" type="nomenclature">
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<paragraph id="8BCEAC4FFFB2FFEDC5C9727027527DCF" blockId="2.[192,628,1404,1461]" box="[192,628,1404,1429]" pageId="2" pageNumber="1">
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<heading id="D0861B23FFB2FFEDC5C9727027527DCF" bold="true" box="[192,628,1404,1429]" fontSize="10" level="2" pageId="2" pageNumber="1" reason="3">
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<emphasis id="B905705DFFB2FFEDC5C9727027527DCF" bold="true" box="[192,628,1404,1429]" pageId="2" pageNumber="1">
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<taxonomicName id="4C71D7CCFFB2FFEDC5C9727024EA7DCE" ID-CoL="5V6MM" authority="Edmonds & Figueroa, 2013" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[192,460,1404,1429]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="pishtaco" status="sp. nov.">
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<emphasis id="B905705DFFB2FFEDC5C9727024EA7DCE" bold="true" box="[192,460,1404,1429]" italics="true" pageId="2" pageNumber="1">Anomiopus pishtaco</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A236CD26FFB2FFEDC4D1727027527DCF" box="[472,628,1405,1429]" pageId="2" pageNumber="1" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C36BFFC4FFB2FFEDC5C9729024167DEF" box="[192,304,1437,1461]" pageId="2" pageNumber="1" type="description">
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<paragraph id="8BCEAC4FFFB2FFEDC5C9729024167DEF" blockId="2.[192,628,1404,1461]" box="[192,304,1437,1461]" pageId="2" pageNumber="1">
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(
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<figureCitation id="134AB0CAFFB2FFEDC5C0729024007DEF" box="[201,294,1437,1461]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="2" pageNumber="1">Fig. 1-8</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36BFFC4FFB2FFEDC5C972D021987ECF" pageId="2" pageNumber="1" type="materials_examined">
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<paragraph id="8BCEAC4FFFB2FFEDC5C972D021987ECF" blockId="2.[192,1441,1499,1685]" pageId="2" pageNumber="1">
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<emphasis id="B905705DFFB2FFEDC5C972D024417DAF" bold="true" box="[192,359,1501,1525]" pageId="2" pageNumber="1">Type Series.</emphasis>
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<materialsCitation id="3B19A612FFB2FFEDC46472D026357E4F" collectingDate="2012-03" collectorName="M. Vilchez" country="Peru" elevation="224" latitude="-8.311945" location="Calleria" longLatPrecision="21" longitude="-73.682495" pageId="2" pageNumber="1" specimenCount="1" stateProvince="Ucayali" typeStatus="holotype">
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<typeStatus id="54CA12EDFFB2FFEDC46472D024FD7DAF" box="[365,475,1501,1525]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
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(sex unknown) -
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<emphasis id="B905705DFFB2FFEDC7A072D027CC7DAF" bold="true" box="[681,746,1501,1525]" pageId="2" pageNumber="1">
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<collectingCountry id="F366ECDFFFB2FFEDC7A072D027CC7DAF" box="[681,746,1501,1525]" name="Peru" pageId="2" pageNumber="1">Peru</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="49B562ADFFB2FFEDC7F072D026947DAF" box="[761,946,1501,1525]" country="Peru" name="Ucayali" pageId="2" pageNumber="1">Región Ucayali</collectingRegion>
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, Prov. Coronel Portillo,
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<location id="8EAEFA94FFB2FFEDC1C672D020087DAF" LSID="urn:lsid:plazi:treatment:03D81D59FFB2FFEAC5C97270247A7A01:8EAEFA94FFB2FFEDC1C672D020087DAF" box="[1231,1326,1501,1525]" country="Peru" latitude="-8.311945" longLatPrecision="21" longitude="-73.682495" name="Calleria" pageId="2" pageNumber="1" stateProvince="Ucayali">Callería</location>
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(
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<geoCoordinate id="EE45CA88FFB2FFEDC03672D025F67E4F" degrees="8" direction="south" minutes="18" orientation="latitude" pageId="2" pageNumber="1" precision="15" seconds="43" value="-8.311945">
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8
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<superScript id="7C040107FFB2FFEDC04472D620737DB3" attach="none" box="[1357,1365,1499,1513]" fontSize="6" pageId="2" pageNumber="1">o</superScript>
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||
18’43" S
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</geoCoordinate>
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<geoCoordinate id="EE45CA88FFB2FFEDC5DF72F024417E4F" box="[214,359,1531,1557]" degrees="73" direction="west" minutes="40" orientation="longitude" pageId="2" pageNumber="1" precision="15" seconds="57" value="-73.682495">
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73
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<superScript id="7C040107FFB2FFEDC5FA72F625DD7E53" attach="none" box="[243,251,1531,1545]" fontSize="6" pageId="2" pageNumber="1">o</superScript>
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40’57" W
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</geoCoordinate>
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)
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<quantity id="4C8901AAFFB2FFEDC47D72F024997E4E" box="[372,447,1533,1557]" metricMagnitude="2" metricUnit="m" metricValue="2.2399999999999998" pageId="2" pageNumber="1" unit="m" value="224.0">
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<elevation id="005C4B7CFFB2FFEDC47D72F024997E4E" box="[372,447,1533,1557]" metricMagnitude="2" metricUnit="m" metricValue="2.2399999999999998" pageId="2" pageNumber="1" unit="m" value="224.0">224 m</elevation>
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</quantity>
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,
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<date id="FFCF8A8FFFB2FFEDC4C372F0277D7E4F" box="[458,603,1533,1557]" pageId="2" pageNumber="1" value="2012-03">
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<collectingDate id="EF8B7367FFB2FFEDC4C372F0277D7E4F" box="[458,603,1533,1557]" pageId="2" pageNumber="1" value="2012-03">March 2012</collectingDate>
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</date>
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,
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<collectorName id="2684C999FFB2FFEDC76C72F027C57E4F" box="[613,739,1533,1557]" pageId="2" pageNumber="1">M. Vilchez</collectorName>
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col.
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||
</materialsCitation>
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<typeStatus id="54CA12EDFFB2FFEDC61172F026A37E4F" box="[792,901,1533,1557]" pageId="2" pageNumber="1" type="paratype">Paratype</typeStatus>
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||
(sex unknown) -
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||
<emphasis id="B905705DFFB2FFEDC15972F021B77E4F" bold="true" box="[1104,1169,1533,1557]" pageId="2" pageNumber="1">
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||
<collectingCountry id="F366ECDFFFB2FFEDC15972F021B77E4F" box="[1104,1169,1533,1557]" name="Peru" pageId="2" pageNumber="1">Peru</collectingCountry>
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||
</emphasis>
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||
:
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<collectingRegion id="49B562ADFFB2FFEDC19672F0207E7E4F" box="[1183,1368,1533,1557]" country="Peru" name="Ucayali" pageId="2" pageNumber="1">Región Ucayali</collectingRegion>
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, Prov. Coronel Portillo (
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<geoCoordinate id="EE45CA88FFB2FFEDC49B7110272D7E6F" box="[402,523,1563,1589]" degrees="8" direction="south" minutes="26" orientation="latitude" pageId="2" pageNumber="1" precision="15" seconds="09" value="-8.435833">
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||
8
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<superScript id="7C040107FFB2FFEDC4A87116248F7E73" attach="none" box="[417,425,1563,1577]" fontSize="6" pageId="2" pageNumber="1">o</superScript>
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26’09" S
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</geoCoordinate>
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||
<geoCoordinate id="EE45CA88FFB2FFEDC71B711027837E6F" box="[530,677,1563,1589]" degrees="73" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="1" precision="15" seconds="35" value="-73.70972">
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||
73
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<superScript id="7C040107FFB2FFEDC7397116271E7E73" attach="right" box="[560,568,1563,1577]" fontSize="6" pageId="2" pageNumber="1">o</superScript>
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42’35" W
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</geoCoordinate>
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)
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<quantity id="4C8901AAFFB2FFEDC7B27110262E7E6E" box="[699,776,1565,1589]" metricMagnitude="2" metricUnit="m" metricValue="2.27" pageId="2" pageNumber="1" unit="m" value="227.0">227 m</quantity>
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,
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<date id="FFCF8A8FFFB2FFEDC61D711021227E6F" box="[788,1028,1565,1589]" pageId="2" pageNumber="1" value="2012-10-15" valueMax="2012-10-17" valueMin="2012-10-15">15-17 October 2012</date>
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, B. Medina col. The
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<typeStatus id="54CA12EDFFB2FFEDC1F2711020407E6F" box="[1275,1382,1565,1589]" pageId="2" pageNumber="1" type="paratype">paratype</typeStatus>
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is in three pieces (head, prothorax and pterothorax + abdomen), each pointed and labeled on a separate pin. Both
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<typeStatus id="54CA12EDFFB2FFEDC40B7150244C7E2F" box="[258,362,1629,1653]" pageId="2" pageNumber="1" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="54CA12EDFFB2FFEDC4AD715027297E2F" box="[420,527,1629,1653]" pageId="2" pageNumber="1" type="paratype">paratype</typeStatus>
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are deposited in the collection of the Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos (
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<collectingRegion id="49B562ADFFB2FFEDC12D717021417ECF" box="[1060,1127,1661,1685]" country="Peru" name="Lima" pageId="2" pageNumber="1">Lima</collectingRegion>
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||
,
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<collectingCountry id="F366ECDFFFB2FFEDC17A717021977ECF" box="[1139,1201,1661,1685]" name="Peru" pageId="2" pageNumber="1">Peru</collectingCountry>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36BFFC4FFB2FFEBC5C971B026F57B01" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1" type="description">
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||
<paragraph id="8BCEAC4FFFB2FFEBC5C971B026F57B01" blockId="2.[192,1441,1724,1909]" lastBlockId="4.[192,1441,195,860]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
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||
<emphasis id="B905705DFFB2FFEDC5C971B027F47E8E" bold="true" box="[192,722,1724,1749]" pageId="2" pageNumber="1">
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||
Description.
|
||
<typeStatus id="54CA12EDFFB2FFEDC47D71B024D77E8F" box="[372,497,1725,1749]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
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||
(sex unknown):
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</emphasis>
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<emphasis id="B905705DFFB2FFEDC7D471B026397E8F" box="[733,799,1725,1749]" italics="true" pageId="2" pageNumber="1">Color</emphasis>
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- Head and pronotum dark shining reddish brown, elytra dark shining brown.
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<emphasis id="B905705DFFB2FFEDC71371D0277C7EAF" box="[538,602,1757,1781]" italics="true" pageId="2" pageNumber="1">Head</emphasis>
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||
: Clypeus (
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<figureCitation id="134AB0CAFFB2FFEDC7E871D0260C7EAF" box="[737,810,1757,1781]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="2" pageNumber="1">Fig. 2</figureCitation>
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) grossly excised medially for nearly entire length, emargination broadly rounded posteriorly, angulate on each side and extended anteriorly as blunt, wedgeshaped clypeal teeth; posterior edge of emargination separated by fine suture from ventrally directed, triangular sclerite (
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<figureCitation id="134AB0CAFFB2FFEDC4BC703024D97F0F" box="[437,511,1853,1877]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="2" pageNumber="1">Fig. 3</figureCitation>
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, arrow) here interpreted as labrum (see Comments below). Frontoclypeal sutures (
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<figureCitation id="134AB0CAFFB2FFEDC422705024527F2F" box="[299,372,1885,1909]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="2" pageNumber="1">Fig. 2</figureCitation>
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, arrow) strongly arched anteriorly, together forming a semicircle; surface of head posterior to (and including) emargination strongly concave. Clypeus lacking distinct puncturing; parietals (genae) roughened by coarse punctures; frons smooth with moderate puncturing, punctures separated by at least two puncture diameters. Frons completely margined posteriorly by fine carina.
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<emphasis id="B905705DFFB4FFEBC017760E20BE7941" box="[1310,1432,259,283]" italics="true" pageId="4" pageNumber="3">Prothorax</emphasis>
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: Pronotum evenly, broadly convex, completely punctate, punctures separated by at least two puncture diameters, becoming larger and more clearly umbilicate on disk. Circumnotal ridge effaced along posterior margin, otherwise sharp and distinct. Anterolateral angles acute (
|
||
<figureCitation id="134AB0CAFFB4FFEBC124766E21527921" box="[1069,1140,355,379]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
|
||
). Transverse propleural carina (
|
||
<figureCitation id="134AB0CAFFB4FFEBC412768E244479C1" box="[283,354,387,411]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 4</figureCitation>
|
||
, upper arrow) present and complete, extending from middle of coxal margin to near lateral margin where it bends abruptly and extends anteriorly to merge into circumnotal ridge; pleural surface posterior to transverse carina smooth, anterior to carina finely transversely striate.
|
||
<emphasis id="B905705DFFB4FFEBC1B176CE20637981" box="[1208,1349,451,475]" italics="true" pageId="4" pageNumber="3">Pterothorax</emphasis>
|
||
: Upper portion of metasternum, mesepimeron, metasternum and adjacent surfaces of meso- and metacoxae very finely reticulate.
|
||
<emphasis id="B905705DFFB4FFEBC49A750E24F97A41" box="[403,479,515,539]" italics="true" pageId="4" pageNumber="3">Elytra</emphasis>
|
||
: Striae obscure, indicated by narrow grooves separated by convex interstriae; seventh stria evident only medially, basal and apical thirds effaced.
|
||
<emphasis id="B905705DFFB4FFEBC6F87529210F7A61" box="[1009,1065,548,571]" italics="true" pageId="4" pageNumber="3">Legs</emphasis>
|
||
: Protibia (
|
||
<figureCitation id="134AB0CAFFB4FFEBC1A5752E20287A61" box="[1196,1294,547,571]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 6–7</figureCitation>
|
||
) tridentate; teeth collectively occupying less than one-half length, narrow, acute, separated by small denticles that continue along lateral margin from third tooth to base; protarsus inserted subapically on posterior (ventral) surface, longer than one-half of tibia along inner margin, claws small, only slightly curved. Profemur globose, strong carina marking both upper and lower margins. Protibial spur acute, almost straight. Meso- and metatibiae (
|
||
<figureCitation id="134AB0CAFFB4FFEBC73675CE27A27A81" box="[575,644,707,731]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 8</figureCitation>
|
||
) scarcely widened apically; tarsomeres elongate, scarcely widened apically; claws small, almost straight.
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC78775EE266D7AA6" authority=": Very" authorityName="Very" box="[654,843,739,764]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B905705DFFB4FFEBC78775EE26237AA1" box="[654,773,739,763]" italics="true" pageId="4" pageNumber="3">Pygidium</emphasis>
|
||
: Very
|
||
</taxonomicName>
|
||
large (
|
||
<figureCitation id="134AB0CAFFB4FFEBC6A975EE26C17AA1" box="[928,999,739,763]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 5</figureCitation>
|
||
), strongly convex, its greatest width amounts to about three-fourths of combined elytral apices; dorsal margin strongly margined and angulate medially, received by upwardly curved apical elytral margins.
|
||
<emphasis id="B905705DFFB4FFEBC6B4742E21697B61" box="[957,1103,803,827]" italics="true" pageId="4" pageNumber="3">Body length</emphasis>
|
||
(including head): ~
|
||
<quantity id="4C8901AAFFB4FFEBC05B742E20BB7B61" box="[1362,1437,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="3" unit="mm" value="4.0">4 mm</quantity>
|
||
. (
|
||
<typeStatus id="54CA12EDFFB4FFEBC5C1744E24107B01" box="[200,310,835,859]" pageId="4" pageNumber="3" type="paratype">Paratype</typeStatus>
|
||
does not differ from
|
||
<typeStatus id="54CA12EDFFB4FFEBC73D744E27BA7B01" box="[564,668,835,859]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
in any significant detail.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0EFCC7FFB3FFECC5C971FE27057F26" ID-DOI="http://doi.org/10.5281/zenodo.5176569" ID-Zenodo-Dep="5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="3" pageNumber="2" startId="3.[192,286,1779,1801]" targetBox="[203,1430,205,1752]" targetPageId="3">
|
||
<paragraph id="8BCEAC4FFFB3FFECC5C971FE27057F26" blockId="3.[192,1440,1779,1916]" pageId="3" pageNumber="2">
|
||
<emphasis id="B905705DFFB3FFECC5C971FE24687F52" bold="true" box="[192,334,1779,1801]" pageId="3" pageNumber="2">Figures 1–8</emphasis>
|
||
.
|
||
<taxonomicName id="4C71D7CCFFB3FFECC45471FE271D7F52" authority="Edmonds & Figueroa, 2013" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[349,571,1779,1800]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pishtaco" status="sp. nov.">
|
||
<emphasis id="B905705DFFB3FFECC45471FE271D7F52" box="[349,571,1779,1800]" italics="true" pageId="3" pageNumber="2">Anomiopus pishtaco</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A236CD26FFB3FFECC74A71FE27E17F53" box="[579,711,1779,1801]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
|
||
, holotype.
|
||
<emphasis id="B905705DFFB3FFECC64971FE267E7F53" bold="true" box="[832,856,1779,1801]" pageId="3" pageNumber="2">1)</emphasis>
|
||
Habitus, dorsal view.
|
||
<emphasis id="B905705DFFB3FFECC14671FE21407F53" bold="true" box="[1103,1126,1779,1801]" pageId="3" pageNumber="2">2)</emphasis>
|
||
Head, dorsal view.
|
||
<emphasis id="B905705DFFB3FFECC04971FE207E7F53" bold="true" box="[1344,1368,1779,1801]" pageId="3" pageNumber="2">3)</emphasis>
|
||
Head, partial dorsal view (arrow indicates labrum).
|
||
<emphasis id="B905705DFFB3FFECC7B0700227F77F7F" bold="true" box="[697,721,1807,1829]" pageId="3" pageNumber="2">4)</emphasis>
|
||
Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina).
|
||
<emphasis id="B905705DFFB3FFECC675702126B27F18" bold="true" box="[892,916,1836,1858]" pageId="3" pageNumber="2">5)</emphasis>
|
||
Abdomen, caudal view.
|
||
<emphasis id="B905705DFFB3FFECC196702121917F18" bold="true" box="[1183,1207,1836,1858]" pageId="3" pageNumber="2">6)</emphasis>
|
||
Left proleg, anterior view.
|
||
<emphasis id="B905705DFFB3FFECC409704424317F05" bold="true" box="[256,279,1865,1887]" pageId="3" pageNumber="2">7)</emphasis>
|
||
Same, posterior view.
|
||
<emphasis id="B905705DFFB3FFECC715704427127F05" bold="true" box="[540,564,1865,1887]" pageId="3" pageNumber="2">8)</emphasis>
|
||
Pterothorax and abdomen, ventral view.
|
||
<emphasis id="B905705DFFB3FFECC1077044215F7F04" bold="true" box="[1038,1145,1865,1887]" pageId="3" pageNumber="2">Figure 9</emphasis>
|
||
.
|
||
<taxonomicName id="4C71D7CCFFB3FFECC183704724677F26" authority="(Westwood)" baseAuthorityName="Westwood" baseAuthorityYear="1842" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="smaragdinus">
|
||
<emphasis id="B905705DFFB3FFECC183704720867F04" box="[1162,1440,1865,1887]" italics="true" pageId="3" pageNumber="2">Anomiopus smaragdinus</emphasis>
|
||
(Westwood)
|
||
</taxonomicName>
|
||
, dorsal view of head.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36BFFC4FFB4FFEBC5C9748E26C97B81" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C9748E26C97B81" blockId="4.[192,1440,899,987]" pageId="4" pageNumber="3">
|
||
<emphasis id="B905705DFFB4FFEBC5C9748E24707BC1" bold="true" box="[192,342,899,923]" pageId="4" pageNumber="3">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC73C748E27BF7BC1" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[565,665,899,923]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pishtaco">
|
||
<emphasis id="B905705DFFB4FFEBC73C748E27BF7BC1" box="[565,665,899,923]" italics="true" pageId="4" pageNumber="3">pishtaco</emphasis>
|
||
</taxonomicName>
|
||
(from Quechua “
|
||
<emphasis id="B905705DFFB4FFEBC661748926967BC1" box="[872,944,900,923]" italics="true" pageId="4" pageNumber="3">pistay</emphasis>
|
||
”; masculine noun in apposition) refers to an Andean mythological bogeyman said to terrorize indigenous communities often brandishing a dagger, here a metaphor for the formidable clypeal teeth of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36BFFC4FFB4FFEBC5C9730E24E47D06" pageId="4" pageNumber="3" type="discussion">
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C9730E27267D41" blockId="4.[192,1441,1027,1372]" pageId="4" pageNumber="3">
|
||
<emphasis id="B905705DFFB4FFEBC5C9730E247F7C41" bold="true" box="[192,345,1027,1051]" pageId="4" pageNumber="3">Comments.</emphasis>
|
||
This new species will key to
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC7A4730926137C41" authorityName="Westwood" authorityYear="1842" box="[685,821,1028,1051]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905705DFFB4FFEBC7A4730926137C41" box="[685,821,1028,1051]" italics="true" pageId="4" pageNumber="3">Anomiopus</emphasis>
|
||
</taxonomicName>
|
||
with little difficulty in
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC140730E205A7C41" author="Vaz-de-Mello, F. Z. & W. D. Edmonds & F. C. Ocampo & P. Schoolmeesters" box="[1097,1404,1027,1051]" pageId="4" pageNumber="3" pagination="1 - 73" refId="ref2746" refString="Vaz-de-Mello, F. Z., W. D. Edmonds, F. C. Ocampo, and P. Schoolmeesters. 2011. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854: 1 - 73." type="journal article" year="2011">Vaz-de-Mello et al. (2011)</bibRefCitation>
|
||
. In her revision of the genus,
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC4F3732E279F7C61" author="Canhedo, V. L." box="[506,697,1059,1083]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">Canhedo (2006)</bibRefCitation>
|
||
recognized the virescens, the smaragdinus and the cuprarius species groups.
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC475734927497C01" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[380,623,1091,1115]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pishtaco">
|
||
<emphasis id="B905705DFFB4FFEBC475734927497C01" box="[380,623,1091,1115]" italics="true" pageId="4" pageNumber="3">Anomiopus pishtaco</emphasis>
|
||
</taxonomicName>
|
||
is assignable to the latter group on the basis of two characters: lack of a subapical tubercle on the lateral margin of the metatibia; and the presence of a complete transverse propleural carina (
|
||
<figureCitation id="134AB0CAFFB4FFEBC4AE738E24D47CC1" box="[423,498,1155,1179]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 4</figureCitation>
|
||
, upper arrow). It is immediately distinguished from other known species of
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC5C973A9246E7CE1" authorityName="Westwood" authorityYear="1842" box="[192,328,1188,1211]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905705DFFB4FFEBC5C973A9246E7CE1" box="[192,328,1188,1211]" italics="true" pageId="4" pageNumber="3">Anomiopus</emphasis>
|
||
</taxonomicName>
|
||
by the grotesque shape of the head (
|
||
<figureCitation id="134AB0CAFFB4FFEBC61673AE26A17CE1" box="[799,903,1187,1211]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 1–2</figureCitation>
|
||
). The clypeal shape of its congeners, while variable, is always similar to that depicted in Figure 9. Both
|
||
<typeStatus id="54CA12EDFFB4FFEBC6BC73CE213B7C81" box="[949,1053,1219,1243]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54CA12EDFFB4FFEBC15473C921EE7C86" box="[1117,1224,1220,1244]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
were collected in human-feces-baited pitfall traps in evergreen forest about
|
||
<quantity id="4C8901AAFFB4FFEBC67373EE26E27CA1" box="[890,964,1251,1275]" metricMagnitude="4" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="3" unit="km" value="95.0">95 km</quantity>
|
||
east of Pucallpa; the collecting sites are separated by about
|
||
<quantity id="4C8901AAFFB4FFEBC4A4720E24DB7D41" box="[429,509,1283,1307]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="3" unit="km" value="15.0">15 km</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5F9722E24E47D06" blockId="4.[192,1441,1027,1372]" pageId="4" pageNumber="3">
|
||
The following emendation of
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC75D722E26017D61" author="Canhedo, V. L." box="[596,807,1315,1339]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">Canhedo’s (2006)</bibRefCitation>
|
||
key to species groups will separate
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC1D0722920787D61" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[1241,1374,1315,1339]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pishtaco">
|
||
<emphasis id="B905705DFFB4FFEBC1D0722920787D61" box="[1241,1374,1315,1339]" italics="true" pageId="4" pageNumber="3">A. pishtaco</emphasis>
|
||
</taxonomicName>
|
||
from its known congeners:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36BFFC4FFB4FFEBC5C9728E20877E61" pageId="4" pageNumber="3" type="key">
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C9728E25F07DC1" blockId="4.[192,219,1411,1467]" box="[192,214,1411,1435]" pageId="4" pageNumber="3">
|
||
<keyLead id="30800F7AFFB4FFEBC5C9728E25F07DC1" box="[192,214,1411,1435]" pageId="4" pageNumber="3">
|
||
<keyStep id="3085B4EAFFB4FFEBC5C9728E25F07DC1" box="[192,214,1411,1435]" pageId="4" pageNumber="3">
|
||
<key id="FFE0FB7EFFB4FFEBC5C9728E25F07DC1" box="[192,214,1411,1435]" pageId="4" pageNumber="3">1.</key>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C972A920B17DE1" blockId="4.[192,219,1411,1467]" lastBlockId="4.[288,1441,1411,1499]" pageId="4" pageNumber="3">
|
||
<keyLead id="30800F7AFFB4FFEBC5C972A925FD7DE1" box="[192,219,1444,1467]" pageId="4" pageNumber="3">—</keyLead>
|
||
Lateral margin of metatibia simple, not interrupted by teeth or carina .....................................
|
||
<emphasis id="B905705DFFB4FFEBC098728E20877DC1" bold="true" box="[1425,1441,1411,1435]" pageId="4" pageNumber="3">2</emphasis>
|
||
Metatibia with one or more transverse teeth on lateral margin .....................................................
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC44B72C925F07E41" blockId="4.[288,1441,1411,1499]" lastBlockId="4.[192,214,1539,1563]" pageId="4" pageNumber="3">
|
||
.............................................................................
|
||
<emphasis id="B905705DFFB4FFEBC67772CE21717D81" bold="true" box="[894,1111,1475,1499]" pageId="4" pageNumber="3">virescens group</emphasis>
|
||
(see
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC19D72CE20777D81" author="Canhedo, V. L." box="[1172,1361,1475,1499]" pageId="4" pageNumber="3" pagination="449 - 458" refId="ref2582" refString="Canhedo, V. L. 2004 b. Anomiopus Westwood (Coleoptera: Scarabaeidae): Novas especies do grupo virescens (Coleoptera, Scarabaeidae). Revista Brasileira de Entomologia 48: 449 - 458." type="journal article" year="2004">Canhedo 2004b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC05572CE20BE7D81" author="Canhedo, V. L." box="[1372,1432,1475,1499]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">2006</bibRefCitation>
|
||
)
|
||
<keyLead id="30800F7AFFB4FFEBC5C9710E25F07E41" box="[192,214,1539,1563]" pageId="4" pageNumber="3">
|
||
<keyStep id="3085B4EAFFB4FFEBC5C9710E25F07E41" box="[192,214,1539,1563]" pageId="4" pageNumber="3">
|
||
<key id="FFE0FB7EFFB4FFEBC5C9710E25F07E41" box="[192,214,1539,1563]" pageId="4" pageNumber="3">2.</key>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C9714925FD7E01" blockId="4.[192,219,1604,1627]" box="[192,219,1604,1627]" pageId="4" pageNumber="3">
|
||
<keyLead id="30800F7AFFB4FFEBC5C9714925FD7E01" box="[192,219,1604,1627]" pageId="4" pageNumber="3">—</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC429710920877E61" blockId="4.[288,1441,1539,1723]" pageId="4" pageNumber="3">
|
||
Transverse propleural carina complete, reaching lateral margin and turning anteriorly to border episternum). Apices of metatarsomeres 1–4 transversally truncated ....... (cuprarius group)
|
||
<emphasis id="B905705DFFB4FFEBC098712E20877E61" bold="true" box="[1425,1441,1571,1595]" pageId="4" pageNumber="3">3</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36BFFC4FFB4FFEBC429714920867EE1" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8BCEAC4FFFB4FFEBC429714920867EE1" blockId="4.[288,1441,1539,1723]" pageId="4" pageNumber="3">
|
||
Transverse propleural carina semicomplete (reaching lateral margin but not turning anteriorly to border episternum) or incomplete (not reaching lateral margin). At least basal (often all) metatarsomere emarginate or obliquely truncate apically ........................................................... .....................................................................
|
||
<emphasis id="B905705DFFB4FFEBC64F71AE21697EE1" bold="true" box="[838,1103,1699,1723]" pageId="4" pageNumber="3">smaragdinus group</emphasis>
|
||
(see
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC18571AE206C7EE1" author="Canhedo, V. L." box="[1164,1354,1699,1723]" pageId="4" pageNumber="3" pagination="449 - 458" refId="ref2582" refString="Canhedo, V. L. 2004 b. Anomiopus Westwood (Coleoptera: Scarabaeidae): Novas especies do grupo virescens (Coleoptera, Scarabaeidae). Revista Brasileira de Entomologia 48: 449 - 458." type="journal article" year="2004">Canhedo 2004b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC05571AE20BE7EE1" author="Canhedo, V. L." box="[1372,1432,1699,1723]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">2006</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36BFFC4FFB4FFEBC5C971EE20867F21" pageId="4" pageNumber="3" type="key">
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C971EE25F07EA1" blockId="4.[192,214,1763,1787]" box="[192,214,1763,1787]" pageId="4" pageNumber="3">
|
||
<keyLead id="30800F7AFFB4FFEBC5C971EE25F07EA1" box="[192,214,1763,1787]" pageId="4" pageNumber="3">
|
||
<keyStep id="3085B4EAFFB4FFEBC5C971EE25F07EA1" box="[192,214,1763,1787]" pageId="4" pageNumber="3">
|
||
<key id="FFE0FB7EFFB4FFEBC5C971EE25F07EA1" box="[192,214,1763,1787]" pageId="4" pageNumber="3">3.</key>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC5C9704920867F61" blockId="4.[192,219,1860,1883]" lastBlockId="4.[288,1441,1763,1915]" pageId="4" pageNumber="3">
|
||
<keyLead id="30800F7AFFB4FFEBC5C9704925FD7F01" box="[192,219,1860,1883]" pageId="4" pageNumber="3">—</keyLead>
|
||
Clypeus deeply excised medially and medial teeth prolonged anteriorly as a pair of knife-like processes (
|
||
<figureCitation id="134AB0CAFFB4FFEBC4B3700E24D97F41" box="[442,511,1795,1819]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
|
||
); labrum exposed as triangular sclerite at base of clypeal emargination (
|
||
<figureCitation id="134AB0CAFFB4FFEBC079700E246E7F61" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
|
||
, arrow) ............................................................................
|
||
<emphasis id="B905705DFFB4FFEBC6EA702E20867F61" bold="true" box="[995,1440,1827,1851]" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C71D7CCFFB4FFEBC6EA702E21D57F61" ID-CoL="5V6MM" authority="Edmonds & Figueroa, 2013" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[995,1267,1827,1851]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pishtaco" status="sp. nov.">
|
||
<emphasis id="B905705DFFB4FFEBC6EA702E21D57F61" bold="true" box="[995,1267,1827,1851]" italics="true" pageId="4" pageNumber="3">Anomiopus pishtaco</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A236CD26FFB4FFEBC008702E20867F61" box="[1281,1440,1827,1851]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB4FFEBC429704E20867F21" blockId="4.[288,1441,1763,1915]" pageId="4" pageNumber="3">
|
||
Clypeal emargination small, medial teeth not prolonged; labrum not exposed ............................. .................................................
|
||
<emphasis id="B905705DFFB4FFEBC7A4706E26AD7F21" bold="true" box="[685,907,1891,1915]" pageId="4" pageNumber="3">cuprarius group</emphasis>
|
||
(remaining species, see
|
||
<bibRefCitation id="EFE0D1BEFFB4FFEBC1B9706E20BE7F21" author="Canhedo, V. L." box="[1200,1432,1891,1915]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">Canhedo 2006: 359</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36BFFC4FFB5FFEAC5F977CE247A7A01" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="8BCEAC4FFFB5FFEAC5F977CE24E77981" blockId="5.[192,1441,195,603]" pageId="5" pageNumber="4">
|
||
The extreme emargination of the clypeus, which reaches the labro-clypeal suture, has had the consequence of exposing the upper surface of the labrum, which in dung beetles is usually tilted upward beneath and against the lower surface of the clypeus. What we here interpret as the labrum appears in
|
||
<figureCitation id="134AB0CAFFB5FFEAC5C9762E24007961" box="[192,294,291,315]" captionStart="Figures 1–8" captionStartId="3.[192,286,1779,1801]" captionTargetBox="[203,1430,205,1752]" captionTargetId="figure-84@3.[203,1430,192,1752]" captionTargetPageId="3" captionText="Figures 1–8. Anomiopus pishtaco new species, holotype. 1) Habitus, dorsal view. 2) Head, dorsal view. 3) Head, partial dorsal view (arrow indicates labrum). 4) Prothorax, ventrolateral view (upper arrow indicates transverse propleural carina; lower arrow indicates pleurosternal carina). 5) Abdomen, caudal view. 6) Left proleg, anterior view. 7) Same, posterior view. 8) Pterothorax and abdomen, ventral view. Figure 9. Anomiopus smaragdinus (Westwood), dorsal view of head." figureDoi="http://doi.org/10.5281/zenodo.5176569" httpUri="https://zenodo.org/record/5176569/files/figure.png" pageId="5" pageNumber="4">Figure 3</figureCitation>
|
||
(arrow) as an obtusely angulate sclerite separated from the clypeus by a faint suture. A similar modification occurs in the Afrotropical deltochiline genus
|
||
<taxonomicName id="4C71D7CCFFB5FFEAC675764920227901" authority="Frolov and Scholtz." authorityName="Frolov and Scholtz." box="[892,1284,323,347]" class="Insecta" family="Scarabaeidae" genus="Dicranocara" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905705DFFB5FFEAC675764921347901" box="[892,1042,324,347]" italics="true" pageId="5" pageNumber="4">Dicranocara</emphasis>
|
||
Frolov and Scholtz.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C71D7CCFFB5FFEAC003764924337921" class="Insecta" family="Scarabaeidae" genus="Dicranocara" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B905705DFFB5FFEAC003764920867901" box="[1290,1440,324,347]" italics="true" pageId="5" pageNumber="4">Dicranocara</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE0D1BEFFB5FFEAC42B766E27617921" author="Frolov, A. V. & C. H. Scholtz" box="[290,583,355,379]" pageId="5" pageNumber="4" pagination="297 - 299" refId="ref2704" refString="Frolov, A. V. and C. H. Scholtz. 2003. A new genus and species of dung beetle from southern Namibia (Coleoptera: Scarabaeidae: Scarabaeinae). African Entomology 11: 297 - 299." type="journal article" year="2003">Frolov and Scholtz 2003</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFE0D1BEFFB5FFEAC78A766E26E87921" author="Deschodt, C. M. & U. Kryger & C. H. Scholtz" box="[643,974,355,379]" pageId="5" pageNumber="4" pagination="361 - 376" refId="ref2653" refString="Deschodt, C. M., U. Kryger and C. H. Scholtz. 2007. New taxa of relictual Canthonini dung beetles (Scarabaeidae: Scarabaeinae) utilizing rock hyrax middens as refuges in South-western Africa. Insect Systematics and Evolution 38: 361 - 376." type="journal article" year="2007">Deschodt and Scholtz, 2007</bibRefCitation>
|
||
) inhabit the Namib Desert, where they utilize rock hyrax middens as refugial habitats. They possess a similarly deeply emarginate clypeus and exposed labrum, along with specializations of the other mouthparts interpreted as adaptations for manipulating hard food.
|
||
</paragraph>
|
||
<paragraph id="8BCEAC4FFFB5FFEAC5F976EE247A7A01" blockId="5.[192,1441,195,603]" pageId="5" pageNumber="4">
|
||
So as not to risk extensive damage to either
|
||
<typeStatus id="54CA12EDFFB5FFEAC7F576E9261679A6" box="[764,816,484,508]" pageId="5" pageNumber="4">type</typeStatus>
|
||
specimen, we have elected not to attempt dissection of genitalia to determine their sex or to examine details of mouthpart structure.
|
||
<bibRefCitation id="EFE0D1BEFFB5FFEAC171750E206F7A41" author="Canhedo, V. L." box="[1144,1353,515,539]" pageId="5" pageNumber="4" pagination="349 - 502" refId="ref2620" refString="Canhedo, V. L. 2006. Revisao taxonomico do genero Anomiopus Westwood, 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Archivos de Zoologia 37: 349 - 502." type="journal article" year="2006">Canhedo’s (2006)</bibRefCitation>
|
||
indicators of sexual dimorphism are generally imprecise and not definitive for the specimens of
|
||
<taxonomicName id="4C71D7CCFFB5FFEAC1F2752920A67A61" authorityName="Edmonds & Figueroa" authorityYear="2013" box="[1275,1408,547,571]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="pishtaco">
|
||
<emphasis id="B905705DFFB5FFEAC1F2752920A67A61" box="[1275,1408,547,571]" italics="true" pageId="5" pageNumber="4">A. pishtaco</emphasis>
|
||
</taxonomicName>
|
||
at our disposal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |