314 lines
41 KiB
XML
314 lines
41 KiB
XML
<document id="F0A2477ABA06541179A32C772BE7B24F" ID-DOI="10.11646/zootaxa.3946.3.3" ID-GBIF-Dataset="fcec1ba5-a256-4419-a228-ca2b529948bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="245390" ID-ZooBank="8D7575DB-636C-4473-91C3-A58C1E5F9E73" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034695781" checkinUser="plazi" docAuthor="Zakrzewska, Marta & Giłka, Wojciech" docDate="2015" docId="03CD3E7A7111A71BFF60A89CD0CCF0D5" docLanguage="en" docName="zt03946p360.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus glaesarius Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="11" lastPageNumber="356" masterDocId="FFF446027117A712FFF7AC04D275F355" masterDocTitle="The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk" masterLastPageNumber="360" masterPageNumber="347" pageNumber="353" updateTime="1698574398835" updateUser="plazi">
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<mods:titleInfo id="88CA2FD00DE9C071236C4146B2DA457F">
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<mods:title id="385D37FF147CBC7E0959C7DF1940FA5C">The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk</mods:title>
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<mods:name id="E03485BDB59B7F6B73716322811C2C78" type="personal">
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<mods:namePart id="4EA2FBF76227EA67FB6186B152CF0509">Zakrzewska, Marta</mods:namePart>
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<mods:name id="9B517B7527C7C2B0908B4AC9B99D6C12" type="personal">
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<mods:roleTerm id="6210631F946B4517EF498E6A1A73E3E4">Author</mods:roleTerm>
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<mods:namePart id="1C6D3693B6ED91E7BB0739C34336EA0B">Giłka, Wojciech</mods:namePart>
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<mods:typeOfResource id="5106BE5C4267030E820442C7948C9732">text</mods:typeOfResource>
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<mods:title id="38087FAAB08CC1A08E73E2A22DF565DF">Zootaxa</mods:title>
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<mods:part id="4755CDB7C05FE99BCC945E23966BACBB">
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<mods:date id="EF69EB3B7A1BC48C07611932E8D42542">2015</mods:date>
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<mods:number id="3F5D4CEFB579B088B311AE6FF35C5B92">3946</mods:number>
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<mods:number id="374094FFA600875EF8450A1A56B3806C">3</mods:number>
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<mods:identifier id="596B566FCB1EFBAB018098F650F1F0AB" type="DOI">10.11646/zootaxa.3946.3.3</mods:identifier>
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<mods:identifier id="47D0ABF0122038F5834C15CC6D92F5B8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="15302A4DA3CE58E7776E0B2B98292341" type="Zenodo-Dep">245390</mods:identifier>
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<treatment id="03CD3E7A7111A71BFF60A89CD0CCF0D5" ID-DOI="http://doi.org/10.5281/zenodo.5616328" ID-GBIF-Taxon="119608926" ID-Zenodo-Dep="5616328" LSID="urn:lsid:plazi:treatment:03CD3E7A7111A71BFF60A89CD0CCF0D5" httpUri="http://treatment.plazi.org/id/03CD3E7A7111A71BFF60A89CD0CCF0D5" lastPageId="9" lastPageNumber="356" pageId="6" pageNumber="353">
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<subSubSection id="C37EDCE77111A714FF60A89CD161F7E7" box="[151,788,1176,1202]" pageId="6" pageNumber="353" type="nomenclature">
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<paragraph id="8BDB8F6C7111A714FF60A89CD161F7E7" blockId="6.[151,788,1176,1202]" box="[151,788,1176,1202]" pageId="6" pageNumber="353">
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<heading id="D09338007111A714FF60A89CD161F7E7" bold="true" box="[151,788,1176,1202]" fontSize="11" level="1" pageId="6" pageNumber="353" reason="1">
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<emphasis id="B910537E7111A714FF60A89CD161F7E7" bold="true" box="[151,788,1176,1202]" pageId="6" pageNumber="353">
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<taxonomicName id="4C64F4EF7111A714FF60A89CD0DFF7E7" ID-CoL="9HM8M" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,682,1176,1202]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="species" species="glaesarius" status="sp. nov.">
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<emphasis id="B910537E7111A714FF60A89CD3EBF7E7" bold="true" box="[151,414,1176,1202]" italics="true" pageId="6" pageNumber="353">Tanytarsus glaesarius</emphasis>
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Giłka
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<emphasis id="B910537E7111A714FE04A89CD07DF7E7" bold="true" box="[499,520,1176,1202]" italics="true" pageId="6" pageNumber="353">et</emphasis>
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Zakrzewska
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</taxonomicName>
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,
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<taxonomicNameLabel id="A223EE057111A714FD40A89CD161F7E7" box="[695,788,1176,1202]" pageId="6" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37EDCE77111A714FF60A8DAD3E1F636" pageId="6" pageNumber="353" type="materials_examined">
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<paragraph id="8BDB8F6C7111A714FF60A8DAD3E1F636" blockId="6.[151,1437,1246,2036]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF60A8DAD33FF7A2" bold="true" box="[151,330,1246,1271]" pageId="6" pageNumber="353">
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<typeStatus id="54DF31CE7111A714FF60A8DAD2A7F7A2" box="[151,210,1246,1271]" pageId="6" pageNumber="353">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54DF31CE7111A714FEA0A8DAD3B6F7A2" box="[343,451,1246,1271]" pageId="6" pageNumber="353" type="holotype">Holotype</typeStatus>
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. Adult male (tarsi of left fore and hind leg missing, tarsi of mid legs broken and separated) preserved in
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<date id="FFDAA9AC7111A714FE55A907D383F64E" box="[418,502,1283,1307]" pageId="6" pageNumber="353" value="2013-10-21">21 x 13</date>
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x
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<quantity id="4C9C22897111A714FDE4A907D021F64F" box="[531,596,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="353" unit="mm" value="1.0">1 mm</quantity>
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piece of amber (Eocene, ~45–40 Ma, Baltic amber, Gulf of Gdańsk; MAI- 415a;
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<figureCitation id="135F93E97111A714FF2BA922D369F66A" box="[220,284,1318,1343]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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A); animal syninclusions:
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<taxonomicName id="4C64F4EF7111A714FDB5A922D171F66A" box="[578,772,1318,1343]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
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,
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<taxonomicName id="4C64F4EF7111A714FCF9A922D1E4F66A" box="[782,913,1318,1343]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
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,
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<taxonomicName id="4C64F4EF7111A714FC6CA922D726F66B" authority="Thysanoptera, Psocoptera" authorityName="Thysanoptera, Psocoptera" box="[923,1363,1318,1343]" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="family">Mordellidae, Thysanoptera, Psocoptera</taxonomicName>
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(all as single ind., MAI-415).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE77111A714FF30A96AD659F6D2" box="[199,1068,1390,1415]" pageId="6" pageNumber="353" type="etymology">
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<paragraph id="8BDB8F6C7111A714FF30A96AD659F6D2" blockId="6.[151,1437,1246,2036]" box="[199,1068,1390,1415]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30A96AD3D1F6D2" bold="true" box="[199,420,1390,1415]" pageId="6" pageNumber="353">Derivatio nominis.</emphasis>
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Adjective derived from the Latin noun ‘
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<emphasis id="B910537E7111A714FC9CA96AD1BDF6D2" box="[875,968,1390,1415]" italics="true" pageId="6" pageNumber="353">glaesum</emphasis>
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’, amber.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE77111A714FF30A996D7E4F54F" pageId="6" pageNumber="353" type="diagnosis">
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<paragraph id="8BDB8F6C7111A714FF30A996D7E4F54F" blockId="6.[151,1437,1246,2036]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30A996D334F6FE" bold="true" box="[199,321,1426,1451]" pageId="6" pageNumber="353">Diagnosis.</emphasis>
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Frontal tubercles stout, cylindrical. Antenna with flagellomeres 10–11 fused in part and 12–13 completely fused. Wing vein FCu placed far distally of RM (VR Cu 1.52). Anal point wide at base, distinctly narrowed subapically, tapering to pointed apex. Superior volsella with 2 setae on apex. Digitus long, extending beyond superior volsella, curved, with blunt apex. Stem of median volsella bulbous, with slender foliate lamellae.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37EDCE77111A71BFF30AA21D0BAF230" lastPageId="9" lastPageNumber="356" pageId="6" pageNumber="353" type="description">
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||
<paragraph id="8BDB8F6C7111A714FF30AA21D045F56B" blockId="6.[151,1437,1246,2036]" box="[199,560,1573,1598]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30AA21D324F56B" bold="true" box="[199,337,1573,1598]" pageId="6" pageNumber="353">Description</emphasis>
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. Adult male (n = 1)
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</paragraph>
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<paragraph id="8BDB8F6C7111A714FF30AA4DD0A2F534" blockId="6.[151,1437,1246,2036]" box="[199,727,1609,1634]" pageId="6" pageNumber="353">
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Total length c.
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<quantity id="4C9C22897111A714FE99AA4ED3B2F534" box="[366,455,1610,1634]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="353" unit="mm" value="1.2">1.2 mm</quantity>
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; wing length c. 820 Μm.
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</paragraph>
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<paragraph id="8BDB8F6C7111A714FF30AA69D1CAF46F" blockId="6.[151,1437,1246,2036]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30AA69D371F5D3" box="[199,260,1645,1670]" italics="true" pageId="6" pageNumber="353">Head</emphasis>
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(
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<figureCitation id="135F93E97111A714FEEEAA69D315F5D3" box="[281,352,1645,1670]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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C–E). Eyes bare, reniform, with dorsomedian extensions (
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<figureCitation id="135F93E97111A714FBE1AA69D62BF5D3" box="[1046,1118,1645,1670]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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C). Frontal tubercles stout, cylindrical with blunt apices, c.15-20 Μm long (
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<figureCitation id="135F93E97111A714FD50AA95D09CF5FF" box="[679,745,1681,1706]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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D). Antennal flagellum composed of 11 segments of which 10 are well discernible, flagellomeres 10–11 fused in part (borders between remaining flagellomeres not visible, but distribution of setal tubercles indicates 13 segments), AR 0.82 (when flagellum measured as 10-segmented), AR 0.65 (as 11-segmented); plume fully developed (setae separated, but setal tubercles well developed) (
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<figureCitation id="135F93E97111A714FAC3AAF9D70EF443" box="[1332,1403,1789,1814]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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E). Length of palpomeres 2–5 (Μm): c. 30, 60, 76, 129. At least 15 clypeals.
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</paragraph>
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<paragraph id="8BDB8F6C7111A714FF30AB41D774F40B" blockId="6.[151,1437,1246,2036]" box="[199,1281,1861,1886]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30AB41D36DF40B" box="[199,280,1861,1886]" italics="true" pageId="6" pageNumber="353">Thorax</emphasis>
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. Ac and Dc not visible (thorax damaged in dorsal part), Pa 3 on each side, Scts 6 at least.
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</paragraph>
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<paragraph id="8BDB8F6C7111A714FF30AB6ED093F4A1" blockId="6.[151,1437,1246,2036]" pageId="6" pageNumber="353">
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<emphasis id="B910537E7111A714FF30AB6ED374F4D4" box="[199,257,1898,1921]" italics="true" pageId="6" pageNumber="353">Wing</emphasis>
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(
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<figureCitation id="135F93E97111A714FEE6AB6DD320F4D7" box="[273,341,1897,1922]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="6" pageNumber="353">Fig. 5</figureCitation>
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F). Slender, with anal lobe weak, broadest at 2/3 length, width: 260 Μm, length/width ratio 3.15. FCu placed far distally of RM; VR Cu 1.52. Veins ending as in most extant
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<taxonomicName id="4C64F4EF7111A714FC1BAB8AD610F4F3" box="[1004,1125,1934,1958]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="genus">Tanytarsus</taxonomicName>
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(from base to tip): An, Sc, Cu1, R1, R2+3, M3+4, R4+5, M1+2; distances between ends of R1–R2+3 and R2+3–R4+5 unequal (VR C c. 2.15). Almost whole wing (except base) covered with dense macrotrichia.
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</paragraph>
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<caption id="DF1BDFE47110A715FF60AB7DD05CF4BE" httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="7" pageNumber="354" targetBox="[163,1406,193,1874]" targetPageId="7">
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<paragraph id="8BDB8F6C7110A715FF60AB7DD05CF4BE" blockId="7.[151,1436,1913,2027]" pageId="7" pageNumber="354">
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<emphasis id="B910537E7110A715FF60AB7DD363F4DA" bold="true" box="[151,278,1913,1935]" pageId="7" pageNumber="354">FIGURE 5.</emphasis>
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||
<taxonomicName id="4C64F4EF7110A715FED6AB7ED076F4DA" box="[289,515,1913,1935]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="glaesarius" status="sp. nov.">
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<emphasis id="B910537E7110A715FED6AB7ED076F4DA" box="[289,515,1913,1935]" italics="true" pageId="7" pageNumber="354">Tanytarsus glaesarius</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A223EE057110A715FDFAAB7ED017F4DA" box="[525,610,1914,1935]" pageId="7" pageNumber="354" rank="species">
|
||
<emphasis id="B910537E7110A715FDFAAB7ED017F4DA" bold="true" box="[525,610,1914,1935]" pageId="7" pageNumber="354">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
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, adult male, holotype MAI-415a (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk).
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<emphasis id="B910537E7110A715FEF7AB9CD367F4F8" bold="true" box="[256,274,1944,1965]" pageId="7" pageNumber="354">A</emphasis>
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—inclusion;
|
||
<emphasis id="B910537E7110A715FE6BAB9CD3D8F4F8" bold="true" box="[412,429,1944,1965]" pageId="7" pageNumber="354">B</emphasis>
|
||
—habitus;
|
||
<emphasis id="B910537E7110A715FDD4AB9CD040F4FB" bold="true" box="[547,565,1944,1966]" pageId="7" pageNumber="354">C</emphasis>
|
||
—head;
|
||
<emphasis id="B910537E7110A715FD65AB9CD0D1F4F8" bold="true" box="[658,676,1944,1965]" pageId="7" pageNumber="354">D</emphasis>
|
||
—frontal tubercles,
|
||
<emphasis id="B910537E7110A715FC8BAB9CD1F8F4F8" bold="true" box="[892,909,1944,1965]" pageId="7" pageNumber="354">E</emphasis>
|
||
—antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion);
|
||
<emphasis id="B910537E7110A715FA84ABB3D7F7F499" bold="true" box="[1395,1410,1975,1996]" pageId="7" pageNumber="354">F</emphasis>
|
||
— wing photographed in transmitted light.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1BDFE4711FA71AFF60A989D68AF6B5" httpUri="https://zenodo.org/record/245396/files/figure.png" pageId="8" pageNumber="355" targetBox="[158,1412,193,1399]" targetPageId="8">
|
||
<paragraph id="8BDB8F6C711FA71AFF60A989D68AF6B5" blockId="8.[151,1436,1421,1504]" pageId="8" pageNumber="355">
|
||
<emphasis id="B910537E711FA71AFF60A989D363F6F7" bold="true" box="[151,278,1421,1443]" pageId="8" pageNumber="355">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C64F4EF711FA71AFED6A98AD076F6F6" box="[289,515,1421,1443]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="glaesarius" status="sp. nov.">
|
||
<emphasis id="B910537E711FA71AFED6A98AD076F6F6" box="[289,515,1421,1443]" italics="true" pageId="8" pageNumber="355">Tanytarsus glaesarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223EE05711FA71AFDFAA989D017F6F7" box="[525,610,1421,1442]" pageId="8" pageNumber="355" rank="species">
|
||
<emphasis id="B910537E711FA71AFDFAA989D017F6F7" bold="true" box="[525,610,1421,1442]" pageId="8" pageNumber="355">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male, holotype MAI-415a (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk).
|
||
<emphasis id="B910537E711FA71AFF0CA9A8D359F697" bold="true" box="[251,300,1452,1474]" pageId="8" pageNumber="355">A–C</emphasis>
|
||
—hypopygium, ventral aspect, photographed in transmitted light (
|
||
<emphasis id="B910537E711FA71AFC2EA9A8D19EF694" bold="true" box="[985,1003,1452,1473]" pageId="8" pageNumber="355">A</emphasis>
|
||
), reflected light (
|
||
<emphasis id="B910537E711FA71AFB55A9A8D6C6F694" bold="true" box="[1186,1203,1452,1473]" pageId="8" pageNumber="355">B</emphasis>
|
||
), and drawn (
|
||
<emphasis id="B910537E711FA71AFAB1A9A8D72DF697" bold="true" box="[1350,1368,1452,1474]" pageId="8" pageNumber="355">C</emphasis>
|
||
);
|
||
<emphasis id="B910537E711FA71AFA87A9A8D7F7F694" bold="true" box="[1392,1410,1452,1473]" pageId="8" pageNumber="355">D</emphasis>
|
||
— superior volsella and digitus,
|
||
<emphasis id="B910537E711FA71AFE35A9CED3A6F68A" bold="true" box="[450,467,1482,1503]" pageId="8" pageNumber="355">E</emphasis>
|
||
—median volsella; (D, E—magnified c. 3 times relative to hypopygium drawn).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB8F6C711FA71AFF30AA0AD1E9F525" blockId="8.[151,1436,1548,1652]" pageId="8" pageNumber="355">
|
||
<emphasis id="B910537E711FA71AFF30AA0AD289F570" box="[199,252,1550,1573]" italics="true" pageId="8" pageNumber="355">Legs</emphasis>
|
||
. Tibia of fore leg without spur. Tibial combs of mid and hind legs separated, fan-shaped, teeth up to 16 Μm long; each comb with slender spur, 26 Μm long (mid leg) and 20–32 Μm long (hind leg). Sensilla chaetica on ta1 of p2 not observed. Lengths of leg segments and leg ratios in
|
||
<tableCitation id="C6E6BAD7711FA71AFCB5AA53D1EDF525" box="[834,920,1623,1648]" captionStart="TABLE 2" captionStartId="8.[151,239,1694,1717]" captionTargetBox="[159,1344,1745,1902]" captionTargetPageId="8" captionText="TABLE 2. Leg segment lengths (Μm) and leg ratios of male Tanytarsus glaesarius sp. nov." httpUri="http://table.plazi.org/id/DF1BDFE4711FA71AFF60AA9AD616F5E0" pageId="8" pageNumber="355" tableUuid="DF1BDFE4711FA71AFF60AA9AD616F5E0">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF1BDFE4711FA71AFF60AA9AD616F5E0" ID-Table-UUID="DF1BDFE4711FA71AFF60AA9AD616F5E0" box="[151,1123,1694,1717]" httpUri="http://table.plazi.org/id/DF1BDFE4711FA71AFF60AA9AD616F5E0" pageId="8" pageNumber="355" targetBox="[159,1344,1745,1902]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BDB8F6C711FA71AFF60AA9AD616F5E0" blockId="8.[151,1123,1694,1717]" box="[151,1123,1694,1717]" pageId="8" pageNumber="355">
|
||
<emphasis id="B910537E711FA71AFF60AA9AD37CF5E0" bold="true" box="[151,265,1694,1717]" pageId="8" pageNumber="355">TABLE 2.</emphasis>
|
||
Leg segment lengths (Μm) and leg ratios of male
|
||
<taxonomicName id="4C64F4EF711FA71AFCEAAA9AD673F5E0" box="[797,1030,1694,1717]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="glaesarius" status="sp. nov.">
|
||
<emphasis id="B910537E711FA71AFCEAAA9AD673F5E0" box="[797,1030,1694,1717]" italics="true" pageId="8" pageNumber="355">Tanytarsus glaesarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223EE05711FA71AFBFAAA9AD616F5E0" box="[1037,1123,1694,1717]" pageId="8" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB8F6C711FA71AFEDAAAD5D735F43F" pageId="8" pageNumber="355">
|
||
<table id="F9647DCC711F58EDFF68AAD5D735F43B" box="[159,1344,1745,1902]" gridcols="9" gridrows="4" pageId="8" pageNumber="355">
|
||
<tr id="35548D2E711F58EDFF68AAD5D735F5BE" box="[159,1344,1745,1771]" gridrow="0" pageId="8" pageNumber="355" rowspan-0="1">
|
||
<th id="7685E452711F58EDFEDAAAD5D326F5BE" box="[301,339,1745,1771]" gridcol="1" gridrow="0" pageId="8" pageNumber="355">fe</th>
|
||
<th id="7685E452711F58EDFE4BAAD5D397F5BE" box="[444,482,1745,1771]" gridcol="2" gridrow="0" pageId="8" pageNumber="355">ti</th>
|
||
<th id="7685E452711F58EDFDBCAAD5D004F5BE" box="[587,625,1745,1771]" gridcol="3" gridrow="0" pageId="8" pageNumber="355">ta1</th>
|
||
<th id="7685E452711F58EDFD2EAAD5D08AF5BE" box="[729,767,1745,1771]" gridcol="4" gridrow="0" pageId="8" pageNumber="355">ta2</th>
|
||
<th id="7685E452711F58EDFC9FAAD5D1FBF5BE" box="[872,910,1745,1771]" gridcol="5" gridrow="0" pageId="8" pageNumber="355">ta3</th>
|
||
<th id="7685E452711F58EDFC00AAD5D668F5BE" box="[1015,1053,1745,1771]" gridcol="6" gridrow="0" pageId="8" pageNumber="355">ta4</th>
|
||
<th id="7685E452711F58EDFB72AAD5D6D5F5BE" box="[1157,1184,1745,1771]" gridcol="7" gridrow="0" pageId="8" pageNumber="355">ta5</th>
|
||
<th id="7685E452711F58EDFAE3AAD5D735F5BE" box="[1300,1344,1745,1771]" gridcol="8" gridrow="0" pageId="8" pageNumber="355">LR</th>
|
||
</tr>
|
||
<tr id="35548D2E711F58EDFF68AAF9D735F443" box="[159,1344,1789,1814]" gridrow="1" pageId="8" pageNumber="355">
|
||
<th id="7685E452711F58EDFF68AAF9D2C6F443" box="[159,179,1789,1814]" gridcol="0" gridrow="1" pageId="8" pageNumber="355">p1</th>
|
||
<td id="7685E452711F58EDFEDAAAF9D326F443" box="[301,339,1789,1814]" gridcol="1" gridrow="1" pageId="8" pageNumber="355">415</td>
|
||
<td id="7685E452711F58EDFE4BAAF9D397F443" box="[444,482,1789,1814]" gridcol="2" gridrow="1" pageId="8" pageNumber="355">235</td>
|
||
<td id="7685E452711F58EDFDBCAAF9D004F443" box="[587,625,1789,1814]" gridcol="3" gridrow="1" pageId="8" pageNumber="355">440</td>
|
||
<td id="7685E452711F58EDFD2EAAF9D08AF443" box="[729,767,1789,1814]" gridcol="4" gridrow="1" pageId="8" pageNumber="355">250</td>
|
||
<td id="7685E452711F58EDFC9FAAF9D1FBF443" box="[872,910,1789,1814]" gridcol="5" gridrow="1" pageId="8" pageNumber="355">195</td>
|
||
<td id="7685E452711F58EDFC00AAF9D668F443" box="[1015,1053,1789,1814]" gridcol="6" gridrow="1" pageId="8" pageNumber="355">135</td>
|
||
<td id="7685E452711F58EDFB72AAF9D6D5F443" box="[1157,1184,1789,1814]" gridcol="7" gridrow="1" pageId="8" pageNumber="355">60</td>
|
||
<td id="7685E452711F58EDFAE3AAF9D735F443" box="[1300,1344,1789,1814]" gridcol="8" gridrow="1" pageId="8" pageNumber="355">1.87</td>
|
||
</tr>
|
||
<tr id="35548D2E711F58EDFF68AB2DD735F417" box="[159,1344,1833,1858]" gridrow="2" pageId="8" pageNumber="355">
|
||
<th id="7685E452711F58EDFF68AB2DD2C6F417" box="[159,179,1833,1858]" gridcol="0" gridrow="2" pageId="8" pageNumber="355">p2</th>
|
||
<td id="7685E452711F58EDFEDAAB2DD326F417" box="[301,339,1833,1858]" gridcol="1" gridrow="2" pageId="8" pageNumber="355">410</td>
|
||
<td id="7685E452711F58EDFE4BAB2DD397F417" box="[444,482,1833,1858]" gridcol="2" gridrow="2" pageId="8" pageNumber="355">320</td>
|
||
<td id="7685E452711F58EDFDBCAB2DD004F417" box="[587,625,1833,1858]" gridcol="3" gridrow="2" pageId="8" pageNumber="355">230</td>
|
||
<td id="7685E452711F58EDFD2EAB2DD08AF417" box="[729,767,1833,1858]" gridcol="4" gridrow="2" pageId="8" pageNumber="355">90</td>
|
||
<td id="7685E452711F58EDFC9FAB2DD1FBF417" box="[872,910,1833,1858]" gridcol="5" gridrow="2" pageId="8" pageNumber="355">80</td>
|
||
<td id="7685E452711F58EDFC00AB2DD668F417" box="[1015,1053,1833,1858]" gridcol="6" gridrow="2" pageId="8" pageNumber="355">60</td>
|
||
<td id="7685E452711F58EDFB72AB2DD6D5F417" box="[1157,1184,1833,1858]" gridcol="7" gridrow="2" pageId="8" pageNumber="355">45</td>
|
||
<td id="7685E452711F58EDFAE3AB2DD735F417" box="[1300,1344,1833,1858]" gridcol="8" gridrow="2" pageId="8" pageNumber="355">0.72</td>
|
||
</tr>
|
||
<tr id="35548D2E711F58EDFF68AB50D735F43B" box="[159,1344,1876,1902]" gridrow="3" pageId="8" pageNumber="355">
|
||
<th id="7685E452711F58EDFF68AB50D2C6F43B" box="[159,179,1876,1902]" gridcol="0" gridrow="3" pageId="8" pageNumber="355">p3</th>
|
||
<td id="7685E452711F58EDFEDAAB50D326F43B" box="[301,339,1876,1902]" gridcol="1" gridrow="3" pageId="8" pageNumber="355">440</td>
|
||
<td id="7685E452711F58EDFE4BAB50D397F43B" box="[444,482,1876,1902]" gridcol="2" gridrow="3" pageId="8" pageNumber="355">375</td>
|
||
<td id="7685E452711F58EDFDBCAB50D004F43B" box="[587,625,1876,1902]" gridcol="3" gridrow="3" pageId="8" pageNumber="355">265</td>
|
||
<td id="7685E452711F58EDFD2EAB50D08AF43B" box="[729,767,1876,1902]" gridcol="4" gridrow="3" pageId="8" pageNumber="355">140</td>
|
||
<td id="7685E452711F58EDFC9FAB50D1FBF43B" box="[872,910,1876,1902]" gridcol="5" gridrow="3" pageId="8" pageNumber="355">130</td>
|
||
<td id="7685E452711F58EDFC00AB50D668F43B" box="[1015,1053,1876,1902]" gridcol="6" gridrow="3" pageId="8" pageNumber="355">90</td>
|
||
<td id="7685E452711F58EDFB72AB50D6D5F43B" box="[1157,1184,1876,1902]" gridcol="7" gridrow="3" pageId="8" pageNumber="355">50</td>
|
||
<td id="7685E452711F58EDFAE3AB50D735F43B" box="[1300,1344,1876,1902]" gridcol="8" gridrow="3" pageId="8" pageNumber="355">0.71</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C711FA71BFF30ABABD0BAF230" blockId="8.[151,1436,1966,2027]" lastBlockId="9.[151,1437,151,896]" lastPageId="9" lastPageNumber="356" pageId="8" pageNumber="355">
|
||
<emphasis id="B910537E711FA71AFF30ABABD323F493" box="[199,342,1967,1990]" italics="true" pageId="8" pageNumber="355">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="135F93E9711FA71AFE9FABAAD3D8F492" box="[360,429,1966,1991]" captionStart="FIGURE 6" captionStartId="8.[151,250,1421,1443]" captionTargetBox="[158,1412,193,1399]" captionTargetId="figure@8.[158,1428,193,1399]" captionTargetPageId="8" captionText="FIGURE 6. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, ventral aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — superior volsella and digitus, E — median volsella; (D, E—magnified c. 3 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245396/files/figure.png" pageId="8" pageNumber="355">Fig. 6</figureCitation>
|
||
A–E). Gonostylus stout, slightly arcuate, c. 50 Μm (as long as gonocoxite), with sparse setae placed on median margin in distal half. At least 2-3 tubercles bearing posterolateral setae on each side of anal point. Anal point wide at base, distinctly narrowed subapically, tapering to pointed apex; spinulae or crests unobservable from ventral side. Superior volsella (visible only in distal part) with 2 setae on apex; digitus long, extending beyond superior volsella, curved, with blunt apex (
|
||
<figureCitation id="135F93E9711EA71BFCBDACDBD1FBF3AD" box="[842,910,223,248]" captionStart="FIGURE 6" captionStartId="8.[151,250,1421,1443]" captionTargetBox="[158,1412,193,1399]" captionTargetId="figure@8.[158,1428,193,1399]" captionTargetPageId="8" captionText="FIGURE 6. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, ventral aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — superior volsella and digitus, E — median volsella; (D, E—magnified c. 3 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245396/files/figure.png" pageId="9" pageNumber="356">Fig. 6</figureCitation>
|
||
D). Stem of median volsella bulbous, c. 15 Μm long, with 3 slender foliate lamellae (
|
||
<figureCitation id="135F93E9711EA71BFDB8AD00D0E3F248" box="[591,662,260,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1421,1443]" captionTargetBox="[158,1412,193,1399]" captionTargetId="figure@8.[158,1428,193,1399]" captionTargetPageId="8" captionText="FIGURE 6. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, ventral aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — superior volsella and digitus, E — median volsella; (D, E—magnified c. 3 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245396/files/figure.png" pageId="9" pageNumber="356">Fig. 6</figureCitation>
|
||
E). Inferior volsella reaching half length of gonostylus, stout, with slightly swollen apical half posteromedially directed, armed with several curved setae on apex and strong setiform microtrichia on posterolateral margin (
|
||
<figureCitation id="135F93E9711EA71BFDB0AD48D0FFF230" box="[583,650,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1421,1443]" captionTargetBox="[158,1412,193,1399]" captionTargetId="figure@8.[158,1428,193,1399]" captionTargetPageId="8" captionText="FIGURE 6. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A – C — hypopygium, ventral aspect, photographed in transmitted light (A), reflected light (B), and drawn (C); D — superior volsella and digitus, E — median volsella; (D, E—magnified c. 3 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245396/files/figure.png" pageId="9" pageNumber="356">Fig. 6</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37EDCE7711EA71BFF30AD6BD0CCF0D5" pageId="9" pageNumber="356" type="discussion">
|
||
<paragraph id="8BDB8F6C711EA71BFF30AD6BD0CCF0D5" blockId="9.[151,1437,151,896]" pageId="9" pageNumber="356">
|
||
<emphasis id="B910537E711EA71BFF30AD6BD34EF2DD" bold="true" box="[199,315,367,392]" pageId="9" pageNumber="356">Remarks.</emphasis>
|
||
A set of characters given in the diagnosis and description (except the number of antennal flagellomeres, see remarks below), indicate that the new species belongs to the genus
|
||
<taxonomicName id="4C64F4EF711EA71BFB4EAD91D746F2F9" box="[1209,1331,405,428]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E711EA71BFB4EAD91D746F2F9" box="[1209,1331,405,428]" italics="true" pageId="9" pageNumber="356">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
. Several significant characters, i.e. the shape of the hypopygial superior volsella (with 2 setae on apex), the digitus (long, extending far beyond superior volsella), and the median volsella (with stout bulbous stem and slender foliate lamellae) are known from species of different systematic groups, e.g. the
|
||
<emphasis id="B910537E711EA71BFC29ADFBD6DEF14D" box="[990,1195,511,536]" italics="true" pageId="9" pageNumber="356">eminulus- mendax</emphasis>
|
||
- or
|
||
<taxonomicName id="4C64F4EF711EA71BFB2AADFBD727F14D" box="[1245,1362,511,536]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="chinyensis">
|
||
<emphasis id="B910537E711EA71BFB2AADFBD727F14D" box="[1245,1362,511,536]" italics="true" pageId="9" pageNumber="356">chinyensis</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<emphasis id="B910537E711EA71BFF57AE20D2C9F169" box="[160,188,548,572]" italics="true" pageId="9" pageNumber="356">cf.</emphasis>
|
||
<bibRefCitation id="EFF5F29D711EA71BFF33AE20D3CDF169" author="Reiss" box="[196,440,548,573]" pageId="9" pageNumber="356" refString="Reiss, F. & Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus - Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, supplement 40, 75 - 200." type="journal article" year="1971">Reiss & Fittkau 1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5F29D711EA71BFE31AE20D021F169" author="Ekrem" box="[454,596,548,573]" pageId="9" pageNumber="356" refString="Ekrem, T. (2003) Towards a phylogeny of Tanytarsus van der Wulp (Diptera: Chironomidae). Is morphology sufficient to reconstruct the genealogical relationships? Insect Systematics & Evolution, 34, 199 - 219. http: // dx. doi. org / 10.1163 / 187631203788964845" type="journal article" year="2003">Ekrem 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5F29D711EA71BFD97AE20D10DF169" author="Gilka" box="[608,888,548,573]" pageId="9" pageNumber="356" refString="Gilka, W. & Paasivirta L. (2009) Evaluation of diagnostic characters of the Tanytarsus chinyensis group (Diptera: Chironomidae), with description of a new species from Lapland. Zootaxa, 2197, 31 - 42." type="journal article" year="2009">Giłka & Paasivirta 2009</bibRefCitation>
|
||
). However, these characters form a combination unknown from any extant or fossil
|
||
<taxonomicName id="4C64F4EF711EA71BFDD7AE4DD0EFF135" box="[544,666,585,608]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E711EA71BFDD7AE4DD0EFF135" box="[544,666,585,608]" italics="true" pageId="9" pageNumber="356">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
, thus a group membership of
|
||
<taxonomicName id="4C64F4EF711EA71BFC1EAE4DD6AAF135" box="[1001,1247,583,608]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="glaesarius">
|
||
<emphasis id="B910537E711EA71BFC1EAE4DD6AAF135" box="[1001,1247,583,608]" italics="true" pageId="9" pageNumber="356">Tanytarsus glaesarius</emphasis>
|
||
</taxonomicName>
|
||
remains open.
|
||
<taxonomicName id="4C64F4EF711EA71BFF60AE69D3F8F1D0" box="[151,397,620,645]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="glaesarius">
|
||
<emphasis id="B910537E711EA71BFF60AE69D3F8F1D0" box="[151,397,620,645]" italics="true" pageId="9" pageNumber="356">Tanytarsus glaesarius</emphasis>
|
||
</taxonomicName>
|
||
is the first Eocene species of this genus known from the adult male having the antenna with the reduced number of flagellomeres (
|
||
<figureCitation id="135F93E9711EA71BFDBCAE8BD0FAF1FD" box="[587,655,655,680]" captionStart="FIGURE 5" captionStartId="7.[151,250,1913,1935]" captionTargetBox="[163,1406,193,1874]" captionTargetId="figure@7.[163,1407,193,1876]" captionTargetPageId="7" captionText="FIGURE 5. Tanytarsus glaesarius sp. nov., adult male, holotype MAI- 415 a (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A — inclusion; B — habitus; C — head; D — frontal tubercles, E — antenna photographed in reflected light (white arrows: borders between well discernible flagellomeres; grey arrow: incomplete fusion; black arrows: complete fusion); F — wing photographed in transmitted light." httpUri="https://zenodo.org/record/245395/files/figure.png" pageId="9" pageNumber="356">Fig. 5</figureCitation>
|
||
E). This character is similar to that discussed above (see remarks on
|
||
<taxonomicName id="4C64F4EF711EA71BFF60AEB0D3FFF198" box="[151,394,692,717]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="electra">
|
||
<emphasis id="B910537E711EA71BFF60AEB0D3FFF198" box="[151,394,692,717]" italics="true" pageId="9" pageNumber="356">Stempellinella electra</emphasis>
|
||
</taxonomicName>
|
||
), but we propose to weigh it differently on a background of all known
|
||
<taxonomicName id="4C64F4EF711EA71BFB5DAEB1D751F199" box="[1194,1316,693,716]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910537E711EA71BFB5DAEB1D751F199" box="[1194,1316,693,716]" italics="true" pageId="9" pageNumber="356">Tanytarsus</emphasis>
|
||
</taxonomicName>
|
||
, including extant species of this genus (males with 13 flagellomeres). Since we have not observed other characters that might suggest non-typical structure (e.g. deformations, asymmetry, brachyptery or other associated features known from non-flying chironomids;
|
||
<emphasis id="B910537E711EA71BFE58AF24D3B9F06D" box="[431,460,800,824]" italics="true" pageId="9" pageNumber="356">cf.</emphasis>
|
||
<bibRefCitation id="EFF5F29D711EA71BFE25AF1BD02AF06D" author="Gilka" box="[466,607,799,824]" pageId="9" pageNumber="356" refString="Gilka, W. (2011 b) Six unusual Cladotanytarsus Kieffer: towards a systematics of the genus and resurrection of Lenziella Kieffer (Diptera: Chironomidae: Tanytarsini). Zootaxa, 3100, 1 - 34." type="journal article" year="2011" yearSuffix="b">Giłka 2011b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF5F29D711EA71BFD9CAF1BD14DF06D" author="Gilka" box="[619,824,799,824]" pageId="9" pageNumber="356" refString="Gilka, W., Soszynska-Maj, A. & Paasivirta, L. (2013 a) The peculiar winter-active midge Diamesa starmachi (Diptera: Chironomidae). Polish Journal of Entomology, 82, 201 - 211. http: // dx. doi. org / 10.2478 / v 10200 - 012 - 0035 - 6" type="journal article" year="2013" yearSuffix="a">
|
||
Giłka
|
||
<emphasis id="B910537E711EA71BFD47AF25D097F06D" box="[688,738,799,824]" italics="true" pageId="9" pageNumber="356">et al</emphasis>
|
||
. 2013a
|
||
</bibRefCitation>
|
||
), we assume that the reduced number of flagellomeres observed in
|
||
<taxonomicName id="4C64F4EF711EA71BFED2AF41D3CCF008" box="[293,441,836,861]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="glaesarius">
|
||
<emphasis id="B910537E711EA71BFED2AF41D3CCF008" box="[293,441,836,861]" italics="true" pageId="9" pageNumber="356">T. glaesarius</emphasis>
|
||
</taxonomicName>
|
||
should be treated as a species-specific feature rather than a character which might be considered in the context of phylogenetic trends.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |