425 lines
52 KiB
XML
425 lines
52 KiB
XML
<document id="CD0BA678FC6BB3268193C2BC53B21206" ID-ISSN="1175-5326" ID-ZooBank="E107064F-2E8D-4312-B426-1CFF9E6E5C65" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629831462356" checkinUser="felipe" docAuthor="Nakajima, Jun" docDate="2012" docId="03CBD358FF9EFFACF3D75297FD5AFAA0" docLanguage="en" docName="zt03586p130.pdf" docOrigin="Zootaxa 3586" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cobitis striata subsp. hakataensis Nakajima 2012, subsp. nov." docType="treatment" docVersion="3" lastPageNumber="116" masterDocId="FFF2AB20FF92FFA1F340511BFFF2FFD7" masterDocTitle="Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan" masterLastPageNumber="130" masterPageNumber="103" pageNumber="115" updateTime="1699118331228" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="555164B67BDE60C89D8CB9059EB9737D">Taxonomic study of the Cobitis striata complex (Cypriniformes, Cobitidae) in Japan</mods:title>
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<mods:namePart id="77C3D1A812165A635B44522BA939413D">Nakajima, Jun</mods:namePart>
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<mods:title id="E92071E44D2E0236E61A61C3E81309FB">Zootaxa</mods:title>
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<mods:date id="D367EDB02B999BB5AD62ED6470518E8C">2012</mods:date>
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<mods:number id="81582A60A39902F554B1C946877F4740">2012-12-14</mods:number>
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<mods:number id="384E54643523D3428B0FBA8AA65DE56E">3586</mods:number>
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<treatment id="03CBD358FF9EFFACF3D75297FD5AFAA0" ID-DOI="http://doi.org/10.5281/zenodo.5259137" ID-Zenodo-Dep="5259137" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0" httpUri="http://treatment.plazi.org/id/03CBD358FF9EFFACF3D75297FD5AFAA0" lastPageId="13" lastPageNumber="116" pageId="12" pageNumber="115">
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<subSubSection id="C37831C5FF9EFFADF3D75297FD02FC72" box="[151,752,907,934]" pageId="12" pageNumber="115" type="nomenclature">
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<paragraph id="8BDD624EFF9EFFADF3D75297FD02FC72" blockId="12.[151,752,907,966]" box="[151,752,907,934]" pageId="12" pageNumber="115">
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<heading id="D095D522FF9EFFADF3D75297FD02FC72" bold="true" box="[151,752,907,934]" fontSize="11" level="1" pageId="12" pageNumber="115" reason="1">
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<emphasis id="B916BE5CFF9EFFADF3D75297FD02FC72" bold="true" box="[151,752,907,934]" pageId="12" pageNumber="115">
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<taxonomicName id="4C6219CDFF9EFFADF3D75297FDABFC72" ID-CoL="7JMR3" authority="Nakajima" authorityName="Nakajima" authorityYear="2012" box="[151,601,907,934]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="115" phylum="Chordata" rank="subSpecies" species="striata" status="subsp. nov." subSpecies="hakataensis">
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<emphasis id="B916BE5CFF9EFFADF3D75297FE27FC71" bold="true" box="[151,469,908,934]" italics="true" pageId="12" pageNumber="115">Cobitis striata hakataensis</emphasis>
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Nakajima
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2250327FF9EFFADF1275290FD02FC72" box="[615,752,907,933]" pageId="12" pageNumber="115" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37831C5FF9EFFADF3D752B5FB21FBFC" pageId="12" pageNumber="115" type="description">
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<paragraph id="8BDD624EFF9EFFADF3D752B5FE5FFC11" blockId="12.[151,752,907,966]" box="[151,429,942,966]" pageId="12" pageNumber="115">
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(
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<figureCitation id="13597ECBFF9EFFADF3DF52B5FEF3FC11" box="[159,257,942,966]" captionStart="FIGURE 3" captionStartId="5.[151,250,1820,1843]" captionTargetBox="[156,1378,671,1773]" captionTargetId="figure-375@5.[151,1437,653,1806]" captionTargetPageId="5" captionText="FIGURE 3. Lateral views of 10 species/subspecies of Japanese striated spined loach: (A) C. striata striata Ikeda, 1936, 55.6 mm SL, TKPM-P17340; (B) C. striata fuchigamii subsp. nov., 64.8 mm SL, Holotype, TKPM-P17341; (C) C. striata hakataensis subsp. nov., 58.0 mm SL, Holotype, TKPM-P17342; (D) C. kaibarai sp. nov., 54.5 mm SL, Holotype, TKPM- P17343; (E) C. magnostriata sp. nov., 62.4 mm SL, Holotype, TKPM-P17344; (F) C. minamorii minamorii sp. nov., 48.9 mm SL, Holotype, TKPM-P17345; (G) C. minamorii oumiensis subsp. nov., 51.7 mm SL, Holotype, TKPM-P17346; (H) C. minamorii tokaiensis subsp. nov., 47.9 mm SL, Holotype, MPM-FI1507; (I) C. minamorii saninensis subsp. nov., 62.1 mm SL, Holotype, TKPM-P17348; (J) C. minamorii yodoensis subsp. nov., 45.1 mm SL, Holotype, FAKU55719." pageId="12" pageNumber="115">Figs. 3C</figureCitation>
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,
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<figureCitation id="13597ECBFF9EFFADF24D52B5FEB8FC11" box="[269,330,942,966]" captionStart="FIGURE 4" captionStartId="8.[151,250,1868,1891]" captionTargetBox="[151,1436,1201,1847]" captionTargetId="figure-591@8.[151,1436,1201,1847]" captionTargetPageId="8" captionText="FIGURE 4. Color in life of adult male of 3 subspecies of Cobitis striata. (A–B): C. s. striata; (A) non-spawning season, Ima River system, Fukuoka Pref., 14 XII 2010; (B) spawning season, Aasahi River system, Okayama Pref., 17 VI 2007, collected by Koji Tominaga; (C–D): C. s. fuchigamii subsp. nov.; (C) non-spawning season, Onga River system, Fukuoka Pref., 14 XII 2010; (D) spawning season, same locality, 13 VI 2007; (E–F): C. s. hakataensis subsp. nov.; (E) non-spawning season, Tatara River system, Fukuoka Pref., 14 XII 2010; (F) spawning season, same locality, 23 V 2005." pageId="12" pageNumber="115">4E, F</figureCitation>
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,
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<figureCitation id="13597ECBFF9EFFADF21652B5FE8BFC11" box="[342,377,942,966]" captionStart="FIGURE 5" captionStartId="9.[151,250,1191,1214]" captionTargetBox="[151,1436,195,1170]" captionTargetId="figure-415@9.[151,1436,193,1170]" captionTargetPageId="9" captionText="FIGURE 5. Color in life of adult females: (A) C. striata striata, Yoshii River system, Okayama Pref., 7 VI 2010; (B) C. striata fuchigamii subsp. nov., Onga River system, Fukuoka Pref., 12 VI 2007; (C) C. striata hakataensis subsp. nov., Tatara River system, Fukuoka Pref., 13 VI 2007; (D) C. kaibarai sp. nov., Rokkaku River system, Saga Pref., 28 VI 2007; (E) C. magnostriata sp. nov., near Biwa Lake, Shiga Pref., 13 VI 2007; (F) C. minamorii minamorii sp. nov., Yoshii River system, Okayama Pref., 7 VI 2007; (G) C. minamorii oumiensis subsp. nov., near Biwa Lake, Shiga Pref., 17 VI 2007; (H) C. minamorii tokaiensis subsp. nov., Iride-ota River system, Shizuoka Pref., 13 VI 2007; (I) C. minamorii saninensis subsp. nov., Hii River system, Shimane Pref., 1. VI 2008; (J) C. minamorii yodoensis subsp. nov., Yodo River system, Kyoto Pref., VIII 1996, collected by Fumitoshi Mori, photographed by Ryu Uchiyama." pageId="12" pageNumber="115">5C</figureCitation>
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,
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<figureCitation id="13597ECBFF9EFFADF2C452B5FE54FC11" box="[388,422,942,966]" captionStart="FIGURE 6" captionStartId="11.[151,250,1835,1858]" captionTargetBox="[151,1436,445,1788]" captionTargetId="figure-243@11.[151,1436,445,1815]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal views of lamina circularis and upper segment of the first branched ray at the base of the pectoral fin of adult males: (A) C. striata striata Ikeda, 1936; (B) C. striata fuchigamii subsp. nov.; (C) C. striata hakataensis subsp. nov.; (D) C. kaibarai sp. nov.; (E) C. magnostriata sp. nov.; (F) C. minamorii minamorii sp. nov.; (G) C. minamorii oumiensis subsp. nov.; (H) C. minamorii tokaiensis subsp. nov.; (I) C. minamorii saninensis subsp. nov.; (J) C. minamorii yodoensis subsp. nov.. Bars equal to 1.0 mm." pageId="12" pageNumber="115">6C</figureCitation>
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)
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</paragraph>
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||
<paragraph id="8BDD624EFF9EFFADF3D752EDFB21FBFC" blockId="12.[151,1436,1014,1067]" pageId="12" pageNumber="115">
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Hakata form of
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<taxonomicName id="4C6219CDFF9EFFADF27A52ECFE3EFBDB" authorityName="Ikeda" authorityYear="1936" box="[314,460,1015,1037]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="striata">
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<emphasis id="B916BE5CFF9EFFADF27A52ECFE3EFBDB" box="[314,460,1015,1037]" italics="true" pageId="12" pageNumber="115">Cobitis striata</emphasis>
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</taxonomicName>
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(middle race):
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<bibRefCitation id="EFF31FBFFF9EFFADF12A52EDFC9BFBDB" author="Nakajima, J. & Onikura, N. & Kitagawa, E. & Kitagawa, T. & Oikawa, S." box="[618,873,1014,1037]" pageId="12" pageNumber="115" pagination="10 - 15" refId="ref24943" refString="Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. (2008) Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57, 10 - 15." type="journal article" year="2008">
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||
Nakajima
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||
<emphasis id="B916BE5CFF9EFFADF19352ECFCF5FBDA" box="[723,775,1015,1037]" italics="true" pageId="12" pageNumber="115">et al.</emphasis>
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2008: 13
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||
</bibRefCitation>
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||
, fig. 2G; Hakata form of middle race of
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||
<taxonomicName id="4C6219CDFF9EFFADF64A52ECFA6EFBDB" authorityName="Ikeda" authorityYear="1936" box="[1290,1436,1015,1037]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="striata">
|
||
<emphasis id="B916BE5CFF9EFFADF64A52ECFA6EFBDB" box="[1290,1436,1015,1037]" italics="true" pageId="12" pageNumber="115">Cobitis striata</emphasis>
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||
</taxonomicName>
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||
complex:
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||
<bibRefCitation id="EFF31FBFFF9EFFADF26B550EFDD4FBFC" author="Kitagawa, E. & Nakajima, J. & Hoshino, K. & Kitagawa, T." box="[299,550,1045,1067]" pageId="12" pageNumber="115" pagination="7 - 19" refId="ref24313" refString="Kitagawa, E., Nakajima, J., Hoshino, K. & Kitagawa, T. (2009) Geographic distribution and biogeographical origin of the spined loach genus Cobitis in north-eastern Kyushu Island, Japan. Japanese Journal of Ichthyology, 56, 7 - 19." type="journal article" year="2009">
|
||
Kitagawa
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||
<emphasis id="B916BE5CFF9EFFADF2D3550DFE34FBFC" box="[403,454,1045,1067]" italics="true" pageId="12" pageNumber="115">et al.</emphasis>
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||
2009: 12
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||
</bibRefCitation>
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||
, fig. 2E, F;
|
||
<taxonomicName id="4C6219CDFF9EFFADF1DE550EFCECFBFC" box="[670,798,1045,1067]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="undefined-3">
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||
<emphasis id="B916BE5CFF9EFFADF1DE550EFD15FBFC" box="[670,743,1045,1067]" italics="true" pageId="12" pageNumber="115">Cobitis</emphasis>
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||
sp. 3
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||
</taxonomicName>
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subsp. 3:
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<bibRefCitation id="EFF31FBFFF9EFFADF0C4550EFB70FBFC" author="Nakajima, J. & Suzawa, Y. & Shimizu, T. & Saitoh, K." box="[900,1154,1045,1067]" pageId="12" pageNumber="115" pagination="86 - 95" refId="ref24995" refString="Nakajima, J., Suzawa, Y., Shimizu, T. & Saitoh, K. (2012) Proposal of standard Japanese names for Japanese loaches in the genus Cobitis (Cobitidae). Japanese Journal of Ichthyology, 59, 86 - 95." type="journal article" year="2012">
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||
Nakajima
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||
<emphasis id="B916BE5CFF9EFFADF0AD550DFBD3FBFC" box="[1005,1057,1045,1067]" italics="true" pageId="12" pageNumber="115">et al.</emphasis>
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2012: 92
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||
</bibRefCitation>
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||
, fig. 3c.
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C37831C5FF9EFFADF3D75543FBC6FA00" pageId="12" pageNumber="115" type="materials_examined">
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<paragraph id="8BDD624EFF9EFFADF3D75543FED6FB43" blockId="12.[151,1436,1111,2036]" pageId="12" pageNumber="115">
|
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<materialsCitation id="3B0A6813FF9EFFADF3D75543FED6FB43" collectingDate="2010-12-12" collectorName="J. Nakajima." country="Japan" county="Tatara River" location="Kyushu" municipality="Kasuya" pageId="12" pageNumber="115" specimenCode="TKPM-P17342" specimenCount="1" stateProvince="Fukuoka" typeStatus="holotype">
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<emphasis id="B916BE5CFF9EFFADF3D75543FEF9FBA7" bold="true" box="[151,267,1112,1136]" inLineHeading="true" pageId="12" pageNumber="115" reason="1">
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<typeStatus id="54D9DCECFF9EFFADF3D75543FEF4FBA7" box="[151,262,1112,1136]" pageId="12" pageNumber="115" type="holotype">Holotype</typeStatus>
|
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.
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||
</emphasis>
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||
<specimenCode id="DBC4CA35FF9EFFADF251554CFE32FBB8" box="[273,448,1111,1135]" collectionCode="TKPM-P" pageId="12" pageNumber="115">TKPM-P17342</specimenCode>
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||
, male, 58.0 mm SL,
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<collectingCountry id="F37522DEFF9EFFADF1E85543FD1EFBB8" box="[680,748,1112,1135]" name="Japan" pageId="12" pageNumber="115">Japan</collectingCountry>
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:
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<collectingCounty id="62BC1AC2FF9EFFADF1B65543FC70FBB8" box="[758,898,1111,1135]" pageId="12" pageNumber="115">Tatara River</collectingCounty>
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,
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<collectingMunicipality id="6BB9F834FF9EFFADF0CD5543FC16FBA7" box="[909,996,1112,1136]" pageId="12" pageNumber="115">Kasuya</collectingMunicipality>
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,
|
||
<collectingRegion id="49A6ACACFF9EFFADF0AE554CFB62FBB8" box="[1006,1168,1111,1135]" country="Japan" name="Fukuoka" pageId="12" pageNumber="115">Fukuoka Pref.</collectingRegion>
|
||
,
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<location id="8EBD3495FF9EFFADF7DA554CFB06FBB8" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0:8EBD3495FF9EFFADF7DA554CFB06FBB8" box="[1178,1268,1111,1135]" country="Japan" county="Tatara River" municipality="Kasuya" name="Kyushu" pageId="12" pageNumber="115" stateProvince="Fukuoka">Kyushu</location>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF7BF554CFA6AFBB8" box="[1279,1432,1111,1135]" pageId="12" pageNumber="115" value="2010-12-12">
|
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<collectingDate id="EF98BD66FF9EFFADF7BF554CFA6AFBB8" box="[1279,1432,1111,1135]" pageId="12" pageNumber="115" value="2010-12-12">12. XII. 2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF3D75567FED6FB43" box="[151,292,1148,1172]" pageId="12" pageNumber="115">J. Nakajima.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9EFFADF38755BBFAE8FA90" blockId="12.[151,1436,1111,2036]" pageId="12" pageNumber="115">
|
||
<materialsCitation id="3B0A6813FF9EFFADF38755BBFAE4FA90" collectingDate="2005-04-20" collectingDateMax="2010-05-13" collectingDateMin="2005-04-20" collectionCode="R" collectorName="J. Nakajima & Naka & Minami-ku & Kyushu & Muromi & Nishi-ku & E. Miyamura" country="Japan" county="Tatara" location="Kyushu" municipality="Kasuya" pageId="12" pageNumber="115" specimenCode="KPM-NI29504, FKUN33756" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Fukuoka" typeStatus="paratype">
|
||
<emphasis id="B916BE5CFF9EFFADF38755BBFEB7FB6F" bold="true" box="[199,325,1184,1208]" inLineHeading="true" pageId="12" pageNumber="115" reason="1">
|
||
<typeStatus id="54D9DCECFF9EFFADF38755BBFEB3FB6F" box="[199,321,1184,1208]" pageId="12" pageNumber="115" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
JNC005,
|
||
<specimenCount id="9D64A9C7FF9EFFADF2FA5584FDF8FB60" box="[442,522,1183,1207]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF1585584FD8CFB60" box="[536,638,1183,1207]" metricMagnitude="-2" metricUnit="m" metricValue="5.4399999999999995" pageId="12" pageNumber="115" unit="mm" value="54.4">54.4 mm</quantity>
|
||
SL, same data as holotype; JNC041,
|
||
<specimenCount id="9D64A9C7FF9EFFADF7705584FB72FB60" box="[1072,1152,1183,1207]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF7CE5584FB01FB60" box="[1166,1267,1183,1207]" metricMagnitude="-2" metricUnit="m" metricValue="6.04" pageId="12" pageNumber="115" unit="mm" value="60.4">60.4 mm</quantity>
|
||
SL,
|
||
<collectingCounty id="62BC1AC2FF9EFFADF66D55BBFA80FB60" box="[1325,1394,1184,1207]" pageId="12" pageNumber="115">Tatara</collectingCounty>
|
||
<collectionCode id="ED73FA8BFF9EFFADF63B55BBFA60FB60" box="[1403,1426,1184,1207]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectingMunicipality id="6BB9F834FF9EFFADF3D755DFFF1CFB0B" box="[151,238,1220,1244]" pageId="12" pageNumber="115">Kasuya</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF3B955DFFE6EFB0B" box="[249,412,1220,1244]" country="Japan" name="Fukuoka" pageId="12" pageNumber="115">Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<location id="8EBD3495FF9EFFADF2E755DFFDF3FB0B" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0:8EBD3495FF9EFFADF2E755DFFDF3FB0B" box="[423,513,1220,1244]" county="Tatara" municipality="Kasuya" name="Kyushu" pageId="12" pageNumber="115" stateProvince="Fukuoka">Kyushu</location>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF14D55DFFD60FB0B" box="[525,658,1220,1244]" pageId="12" pageNumber="115" value="2005-05-23">
|
||
<collectingDate id="EF98BD66FF9EFFADF14D55DFFD60FB0B" box="[525,658,1220,1244]" pageId="12" pageNumber="115" value="2005-05-23">23. V. 2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF1DD55DFFCD5FB0B" box="[669,807,1220,1244]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
;
|
||
<specimenCode id="DBC4CA35FF9EFFADF07355DFFC12FB0B" box="[819,992,1220,1244]" collectionCode="KPM-NI" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" lsid="urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="12" pageNumber="115">KPM-NI29504</specimenCode>
|
||
, male, 55.0 mm SL, Tatara
|
||
<collectionCode id="ED73FA8BFF9EFFADF65955DFFAC2FB0C" box="[1305,1328,1220,1243]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
., Kasuya,
|
||
<collectingRegion id="49A6ACACFF9EFFADF3D755FCFECBFB28" box="[151,313,1255,1279]" country="Japan" name="Fukuoka" pageId="12" pageNumber="115">Fukuoka Pref.</collectingRegion>
|
||
, Kyushu,
|
||
<date id="FFDC448EFF9EFFADF2EA55FCFDDDFB28" box="[426,559,1255,1279]" pageId="12" pageNumber="115" value="2008-05-18">
|
||
<collectingDate id="EF98BD66FF9EFFADF2EA55FCFDDDFB28" box="[426,559,1255,1279]" pageId="12" pageNumber="115" value="2008-05-18">18. V. 2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF17A55F3FD36FB28" box="[570,708,1255,1279]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
; MPM-FI1502,
|
||
<specimenCount id="9D64A9C7FF9EFFADF03955FCFC3AFB28" box="[889,968,1255,1279]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF09455FCFBC5FB28" box="[980,1079,1255,1279]" metricMagnitude="-2" metricUnit="m" metricValue="4.88" pageId="12" pageNumber="115" unit="mm" value="48.8">48.8 mm</quantity>
|
||
SL, same data;
|
||
<specimenCode id="DBC4CA35FF9EFFADF7AD55FCFA71FB28" box="[1261,1411,1255,1279]" collectionCode="FKUN" pageId="12" pageNumber="115">FKUN33756</specimenCode>
|
||
,
|
||
<specimenCount id="9D64A9C7FF9EFFADF6CE55FCFF14FAF3" pageId="12" pageNumber="115" type="female">1 female</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF3B05417FEA1FAF4" box="[240,339,1292,1316]" metricMagnitude="-2" metricUnit="m" metricValue="8.74" pageId="12" pageNumber="115" unit="mm" value="87.4">87.4 mm</quantity>
|
||
SL,
|
||
<collectorName id="26970798FF9EFFADF2C85417FE31FAF3" box="[392,451,1292,1316]" pageId="12" pageNumber="115">Naka</collectorName>
|
||
<collectionCode id="ED73FA8BFF9EFFADF2895417FE12FAF4" box="[457,480,1292,1315]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectorName id="26970798FF9EFFADF2B15417FD80FAF3" box="[497,626,1292,1316]" pageId="12" pageNumber="115">Minami-ku</collectorName>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF13C5417FC7EFAF3" box="[636,908,1292,1316]" isEnumeration="true" pageId="12" pageNumber="115">Fukuoka, Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF0D75417FC03FAF3" box="[919,1009,1292,1316]" pageId="12" pageNumber="115">Kyushu</collectorName>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF0BB5417FB75FAF3" box="[1019,1159,1292,1316]" pageId="12" pageNumber="115" value="2005-04-20">
|
||
<collectingDate id="EF98BD66FF9EFFADF0BB5417FB75FAF3" box="[1019,1159,1292,1316]" pageId="12" pageNumber="115" value="2005-04-20">20. IV. 2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF7D15417FAE9FAF3" box="[1169,1307,1292,1316]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
; JNC006,
|
||
<specimenCount id="9D64A9C7FF9EFFADF6CE5417FF22FA90" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF39B5434FECCFA90" box="[219,318,1327,1351]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="12" pageNumber="115" unit="mm" value="59.1">59.1 mm</quantity>
|
||
SL,
|
||
<collectorName id="26970798FF9EFFADF2355434FE23FA90" box="[373,465,1327,1351]" pageId="12" pageNumber="115">Muromi</collectorName>
|
||
<collectionCode id="ED73FA8BFF9EFFADF297542BFE1CFA90" box="[471,494,1328,1351]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectorName id="26970798FF9EFFADF2BF5434FD96FA90" box="[511,612,1327,1351]" pageId="12" pageNumber="115">Nishi-ku</collectorName>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF12F5434FC72FA90" box="[623,896,1327,1351]" isEnumeration="true" pageId="12" pageNumber="115">Fukuoka, Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF0CB5434FC17FA90" box="[907,997,1327,1351]" pageId="12" pageNumber="115">Kyushu</collectorName>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF0B15434FB86FA90" box="[1009,1140,1327,1351]" pageId="12" pageNumber="115" value="2010-05-13">
|
||
<collectingDate id="EF98BD66FF9EFFADF0B15434FB86FA90" box="[1009,1140,1327,1351]" pageId="12" pageNumber="115" value="2010-05-13">13. V. 2010</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="26970798FF9EFFADF73F542BFAE4FA90" box="[1151,1302,1327,1351]" pageId="12" pageNumber="115">E. Miyamura</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9EFFADF387544FFBC6FA00" blockId="12.[151,1436,1111,2036]" pageId="12" pageNumber="115">
|
||
<materialsCitation id="3B0A6813FF9EFFADF387544FFB6BFABB" box="[199,1177,1363,1388]" collectingDate="2010-12-12" collectorName="J. Nakajima." country="Japan" county="Tatara River" location="Kyushu" municipality="Kasuya" pageId="12" pageNumber="115" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fukuoka">
|
||
<emphasis id="B916BE5CFF9EFFADF387544FFE4FFABB" bold="true" box="[199,445,1363,1388]" pageId="12" pageNumber="115">Non-type specimens.</emphasis>
|
||
<specimenCount id="9D64A9C7FF9EFFADF284544FFDE2FABB" box="[452,528,1364,1388]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D64A9C7FF9EFFADF108544FFD4AFABB" box="[584,696,1364,1388]" pageId="12" pageNumber="115" type="female">2 females</specimenCount>
|
||
,
|
||
<quantity id="4C9ACFABFF9EFFADF183544FFC94FABC" box="[707,870,1364,1388]" metricMagnitude="-2" metricUnit="m" metricValue="5.99" metricValueMax="6.34" metricValueMin="5.64" pageId="12" pageNumber="115" unit="mm" value="59.9" valueMax="63.4" valueMin="56.4">56.4–63.4 mm</quantity>
|
||
SL, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0A6813FF9EFFADF7E5544FFADFFA58" collectingDate="2006-06-05" collectionCode="R, VI" collectorName="J. Nakajima" country="Japan" county="Muromi" location="Kyushu" municipality="Nishi-ku" pageId="12" pageNumber="115" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fukuoka, Fukuoka Pref.">
|
||
<specimenCount id="9D64A9C7FF9EFFADF7E5544FFB02FABB" box="[1189,1264,1364,1388]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D64A9C7FF9EFFADF668544FFA6AFABB" box="[1320,1432,1364,1388]" pageId="12" pageNumber="115" type="female">2 females</specimenCount>
|
||
, 64.0–
|
||
<quantity id="4C9ACFABFF9EFFADF396546CFECEFA58" box="[214,316,1399,1423]" metricMagnitude="-2" metricUnit="m" metricValue="6.97" pageId="12" pageNumber="115" unit="mm" value="69.7">69.7 mm</quantity>
|
||
SL,
|
||
<collectingCounty id="62BC1AC2FF9EFFADF238546CFE26FA58" box="[376,468,1399,1423]" pageId="12" pageNumber="115">Muromi</collectingCounty>
|
||
<collectionCode id="ED73FA8BFF9EFFADF29E5463FE07FA58" box="[478,501,1400,1423]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectingMunicipality id="6BB9F834FF9EFFADF149546CFD9CFA58" box="[521,622,1399,1423]" pageId="12" pageNumber="115">Nishi-ku</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF13C546CFC66FA58" box="[636,916,1399,1423]" isEnumeration="true" pageId="12" pageNumber="115">Fukuoka, Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<location id="8EBD3495FF9EFFADF0E2546CFC0EFA58" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0:8EBD3495FF9EFFADF0E2546CFC0EFA58" box="[930,1020,1399,1423]" country="Japan" county="Muromi" municipality="Nishi-ku" name="Kyushu" pageId="12" pageNumber="115" stateProvince="Fukuoka, Fukuoka Pref.">Kyushu</location>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF74A546CFB61FA58" box="[1034,1171,1399,1423]" pageId="12" pageNumber="115" value="2006-06-05">
|
||
<collectingDate id="EF98BD66FF9EFFADF74A546CFB61FA58" box="[1034,1171,1399,1423]" pageId="12" pageNumber="115" value="2006-06-05">
|
||
5.
|
||
<collectionCode id="ED73FA8BFF9EFFADF7695463FBB8FA58" box="[1065,1098,1400,1423]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="115">VI</collectionCode>
|
||
. 2006
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF7E05463FADFFA58" box="[1184,1325,1399,1423]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0A6813FF9EFFADF67B546CFB99FA63" collectingDate="2005-05-23" collectionCode="R, V" collectorName="J. Nakajima" country="Japan" county="Tatara" location="Kyushu" municipality="Kasuya" pageId="12" pageNumber="115" specimenCount="2" specimenCount-male="2" stateProvince="Fukuoka">
|
||
<specimenCount id="9D64A9C7FF9EFFADF67B546CFA65FA58" box="[1339,1431,1399,1423]" pageId="12" pageNumber="115" type="male">2 males</specimenCount>
|
||
, 65.0,
|
||
<quantity id="4C9ACFABFF9EFFADF3955487FECAFA64" box="[213,312,1436,1460]" metricMagnitude="-2" metricUnit="m" metricValue="6.57" pageId="12" pageNumber="115" unit="mm" value="65.7">65.7 mm</quantity>
|
||
SL,
|
||
<collectingCounty id="62BC1AC2FF9EFFADF22E5487FE46FA64" box="[366,436,1436,1459]" pageId="12" pageNumber="115">Tatara</collectingCounty>
|
||
<collectionCode id="ED73FA8BFF9EFFADF2FA5487FE23FA64" box="[442,465,1436,1459]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectingMunicipality id="6BB9F834FF9EFFADF2A15487FDCAFA63" box="[481,568,1436,1460]" pageId="12" pageNumber="115">Kasuya</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF1035487FD16FA63" box="[579,740,1436,1460]" country="Japan" name="Fukuoka" pageId="12" pageNumber="115">Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<location id="8EBD3495FF9EFFADF1AF5487FCBBFA63" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0:8EBD3495FF9EFFADF1AF5487FCBBFA63" box="[751,841,1436,1460]" country="Japan" county="Tatara" municipality="Kasuya" name="Kyushu" pageId="12" pageNumber="115" stateProvince="Fukuoka">Kyushu</location>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF0145487FC25FA63" box="[852,983,1436,1460]" pageId="12" pageNumber="115" value="2005-05-23">
|
||
<collectingDate id="EF98BD66FF9EFFADF0145487FC25FA63" box="[852,983,1436,1460]" pageId="12" pageNumber="115" value="2005-05-23">
|
||
23.
|
||
<collectionCode id="ED73FA8BFF9EFFADF03D5487FC63FA64" box="[893,913,1436,1459]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="115" type="Museum">V</collectionCode>
|
||
. 2005
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF0A15487FB99FA63" box="[993,1131,1436,1460]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0A6813FF9EFFADF7355487FBC2FA00" collectingDate="2008-05-18" collectionCode="R, V" collectorName="J. Nakajima" country="Japan" county="Tatara" location="Kyushu" municipality="Kasuya" pageId="12" pageNumber="115" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fukuoka">
|
||
<specimenCount id="9D64A9C7FF9EFFADF7355487FB32FA63" box="[1141,1216,1436,1460]" pageId="12" pageNumber="115" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D64A9C7FF9EFFADF7B65487FAABFA63" box="[1270,1369,1436,1460]" pageId="12" pageNumber="115" type="female">1 female</specimenCount>
|
||
, 52.6,
|
||
<quantity id="4C9ACFABFF9EFFADF3D754A4FF08FA00" box="[151,250,1471,1495]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" pageId="12" pageNumber="115" unit="mm" value="55.5">55.5 mm</quantity>
|
||
SL,
|
||
<collectingCounty id="62BC1AC2FF9EFFADF27054DBFE87FA00" box="[304,373,1472,1495]" pageId="12" pageNumber="115">Tatara</collectingCounty>
|
||
<collectionCode id="ED73FA8BFF9EFFADF23C54DBFE61FA00" box="[380,403,1472,1495]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="115">R</collectionCode>
|
||
.,
|
||
<collectingMunicipality id="6BB9F834FF9EFFADF2E454DBFE09FA0F" box="[420,507,1472,1496]" pageId="12" pageNumber="115">Kasuya</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49A6ACACFF9EFFADF14654A4FD5AFA00" box="[518,680,1471,1495]" country="Japan" name="Fukuoka" pageId="12" pageNumber="115">Fukuoka Pref.</collectingRegion>
|
||
,
|
||
<location id="8EBD3495FF9EFFADF1F354A4FCFFFA00" LSID="urn:lsid:plazi:treatment:03CBD358FF9EFFACF3D75297FD5AFAA0:8EBD3495FF9EFFADF1F354A4FCFFFA00" box="[691,781,1471,1495]" country="Japan" county="Tatara" municipality="Kasuya" name="Kyushu" pageId="12" pageNumber="115" stateProvince="Fukuoka">Kyushu</location>
|
||
,
|
||
<date id="FFDC448EFF9EFFADF05854A4FC68FA00" box="[792,922,1471,1495]" pageId="12" pageNumber="115" value="2008-05-18">
|
||
<collectingDate id="EF98BD66FF9EFFADF05854A4FC68FA00" box="[792,922,1471,1495]" pageId="12" pageNumber="115" value="2008-05-18">
|
||
18.
|
||
<collectionCode id="ED73FA8BFF9EFFADF00254DBFCA4FA00" box="[834,854,1472,1495]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="115" type="Museum">V</collectionCode>
|
||
. 2008
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26970798FF9EFFADF0E654DBFBC2FA00" box="[934,1072,1471,1495]" pageId="12" pageNumber="115">J. Nakajima</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37831C5FF9EFFADF38754F8FF1CF8CB" pageId="12" pageNumber="115" type="diagnosis">
|
||
<paragraph id="8BDD624EFF9EFFADF38754F8FF1CF8CB" blockId="12.[151,1436,1111,2036]" pageId="12" pageNumber="115">
|
||
<emphasis id="B916BE5CFF9EFFADF38754F8FEB3FA2B" bold="true" box="[199,321,1507,1532]" inLineHeading="true" pageId="12" pageNumber="115" reason="1">Diagnosis.</emphasis>
|
||
This subspecies is distinguishable from other Japanese striated spined loaches by the following characteristics: body size moderate, the mature size about
|
||
<quantity id="4C9ACFABFF9EFFADF002571CFC49F9C8" box="[834,955,1543,1567]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="12" pageNumber="115" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50–60 mm</quantity>
|
||
SL in males,
|
||
<quantity id="4C9ACFABFF9EFFADF729571CFB17F9C8" box="[1129,1253,1543,1567]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" pageId="12" pageNumber="115" unit="mm" value="67.5" valueMax="80.0" valueMin="55.0">55–80 mm</quantity>
|
||
SL in females; lamina circularis at the base of the pectoral fin of adult male simple roundish plate, the upper segments of the first branched soft ray narrow and weak (
|
||
<figureCitation id="13597ECBFF9EFFADF16F5754FD77F9B0" box="[559,645,1615,1639]" captionStart="FIGURE 6" captionStartId="11.[151,250,1835,1858]" captionTargetBox="[151,1436,445,1788]" captionTargetId="figure-243@11.[151,1436,445,1815]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal views of lamina circularis and upper segment of the first branched ray at the base of the pectoral fin of adult males: (A) C. striata striata Ikeda, 1936; (B) C. striata fuchigamii subsp. nov.; (C) C. striata hakataensis subsp. nov.; (D) C. kaibarai sp. nov.; (E) C. magnostriata sp. nov.; (F) C. minamorii minamorii sp. nov.; (G) C. minamorii oumiensis subsp. nov.; (H) C. minamorii tokaiensis subsp. nov.; (I) C. minamorii saninensis subsp. nov.; (J) C. minamorii yodoensis subsp. nov.. Bars equal to 1.0 mm." pageId="12" pageNumber="115">Fig. 6C</figureCitation>
|
||
); PMN commonly 13; line L3 formed by incomplete longitudinal line, reaching to caudal base; line L4 formed by longitudinal jagged weblike line, reaching to postanal body, broader than L
|
||
<quantity id="4C9ACFABFF9EFFADF3A1578CFEFEF978" box="[225,268,1687,1711]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="115" unit="in" value="3.0">3 in</quantity>
|
||
male of non-spawning season; line L5 organized in 11–14 roundish or ovoid blotches in non-spawning season; caudal fin and dorsal fin with 3–4 arcuate bars; upper spot at the caudal base jet-black comparable in size to eye diameter; lower spot at caudal base faint or missing; egg yolk diameter approximately 1.0mm; karyotype diploid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37831C5FF9EFFACF387563CFD46FBAB" lastPageId="13" lastPageNumber="116" pageId="12" pageNumber="115" type="description">
|
||
<paragraph id="8BDD624EFF9EFFACF387563CFDA1FE18" blockId="12.[151,1436,1111,2036]" lastBlockId="13.[151,1436,151,1400]" lastPageId="13" lastPageNumber="116" pageId="12" pageNumber="115">
|
||
<emphasis id="B916BE5CFF9EFFADF387563CFEAAF897" bold="true" box="[199,344,1831,1856]" inLineHeading="true" pageId="12" pageNumber="115" reason="1">Description.</emphasis>
|
||
Lateral view in
|
||
<figureCitation id="13597ECBFF9EFFADF15A563CFD7CF8E8" box="[538,654,1831,1855]" captionStart="FIGURE 3" captionStartId="5.[151,250,1820,1843]" captionTargetBox="[156,1378,671,1773]" captionTargetId="figure-375@5.[151,1437,653,1806]" captionTargetPageId="5" captionText="FIGURE 3. Lateral views of 10 species/subspecies of Japanese striated spined loach: (A) C. striata striata Ikeda, 1936, 55.6 mm SL, TKPM-P17340; (B) C. striata fuchigamii subsp. nov., 64.8 mm SL, Holotype, TKPM-P17341; (C) C. striata hakataensis subsp. nov., 58.0 mm SL, Holotype, TKPM-P17342; (D) C. kaibarai sp. nov., 54.5 mm SL, Holotype, TKPM- P17343; (E) C. magnostriata sp. nov., 62.4 mm SL, Holotype, TKPM-P17344; (F) C. minamorii minamorii sp. nov., 48.9 mm SL, Holotype, TKPM-P17345; (G) C. minamorii oumiensis subsp. nov., 51.7 mm SL, Holotype, TKPM-P17346; (H) C. minamorii tokaiensis subsp. nov., 47.9 mm SL, Holotype, MPM-FI1507; (I) C. minamorii saninensis subsp. nov., 62.1 mm SL, Holotype, TKPM-P17348; (J) C. minamorii yodoensis subsp. nov., 45.1 mm SL, Holotype, FAKU55719." pageId="12" pageNumber="115">Figure 3C</figureCitation>
|
||
illustrate body shape, form and position of fins. Morphometric and meristic data for
|
||
<specimenCount id="9D64A9C7FF9EFFADF21F5657FE34F8B3" box="[351,454,1868,1892]" pageId="12" pageNumber="115" type="male">11 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D64A9C7FF9EFFADF1435657FD80F8B3" box="[515,626,1868,1892]" pageId="12" pageNumber="115" type="female">5 females</specimenCount>
|
||
are summarized in
|
||
<tableCitation id="C6E057F5FF9EFFADF01D5657FC44F8B3" box="[861,950,1868,1892]" captionStart="TABLE 2" captionStartId="6.[147,224,526,546]" captionText="TABLE 2. Morphometric and meristic characters of 10 species and subspecies of the Cobitis striata complex in Japan." pageId="12" pageNumber="115">Table 2</tableCitation>
|
||
. Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7–8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with two well-developed lobes; upper lip with transverse wrinkles on surface. Barbels, 3 pairs, first on rostora, second on maxillae, third on maxillomandibula; each barbel well developed, length of maxillary barbel same as eye diameter; length of rostral and mandibular barbels shorter than that of maxillary barbel. Lateral line short, reaching the central region between the pectoral-fin base and the tip of the fin. PMN commonly 13 (range, 13–14). Very small cycloid scales on the trunk. Lamina circularis at the base of the pectoral fin of adult male simple roundish plate (
|
||
<figureCitation id="13597ECBFF9FFFACF19F501FFCC5FECB" box="[735,823,260,284]" captionStart="FIGURE 6" captionStartId="11.[151,250,1835,1858]" captionTargetBox="[151,1436,445,1788]" captionTargetId="figure-243@11.[151,1436,445,1815]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal views of lamina circularis and upper segment of the first branched ray at the base of the pectoral fin of adult males: (A) C. striata striata Ikeda, 1936; (B) C. striata fuchigamii subsp. nov.; (C) C. striata hakataensis subsp. nov.; (D) C. kaibarai sp. nov.; (E) C. magnostriata sp. nov.; (F) C. minamorii minamorii sp. nov.; (G) C. minamorii oumiensis subsp. nov.; (H) C. minamorii tokaiensis subsp. nov.; (I) C. minamorii saninensis subsp. nov.; (J) C. minamorii yodoensis subsp. nov.. Bars equal to 1.0 mm." pageId="13" pageNumber="116">Fig. 6C</figureCitation>
|
||
). The first branched soft ray of pectoral fin longer than the others; pectoral fin of the male relatively longer than that of the female. The upper segments of the first branched soft ray of pectoral fin narrow and weak. Dorsal-fin base equidistant from the base of the caudal fin and the tip of the snout. Pelvic-fin origin below first or second branched dorsal-fin ray. Anal fin not reaching caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Largest recorded specimens:
|
||
<quantity id="4C9ACFABFF9FFFACF3D750ACFF08FE18" box="[151,250,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="6.57" pageId="13" pageNumber="116" unit="mm" value="65.7">65.7 mm</quantity>
|
||
SL male,
|
||
<quantity id="4C9ACFABFF9FFFACF22D50ACFE22FE18" box="[365,464,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="6.97" pageId="13" pageNumber="116" unit="mm" value="69.7">69.7 mm</quantity>
|
||
SL female.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF38750C0FCA3FCE0" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF38750C0FEBDFE23" bold="true" box="[199,335,475,500]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Coloration.</emphasis>
|
||
<emphasis id="B916BE5CFF9FFFACF21D50C7FD01FE23" box="[349,755,476,500]" italics="true" pageId="13" pageNumber="116">Male in the non-spawning season</emphasis>
|
||
(
|
||
<figureCitation id="13597ECBFF9FFFACF04950C7FC80FE23" box="[777,882,476,500]" captionStart="FIGURE 3" captionStartId="5.[151,250,1820,1843]" captionTargetBox="[156,1378,671,1773]" captionTargetId="figure-375@5.[151,1437,653,1806]" captionTargetPageId="5" captionText="FIGURE 3. Lateral views of 10 species/subspecies of Japanese striated spined loach: (A) C. striata striata Ikeda, 1936, 55.6 mm SL, TKPM-P17340; (B) C. striata fuchigamii subsp. nov., 64.8 mm SL, Holotype, TKPM-P17341; (C) C. striata hakataensis subsp. nov., 58.0 mm SL, Holotype, TKPM-P17342; (D) C. kaibarai sp. nov., 54.5 mm SL, Holotype, TKPM- P17343; (E) C. magnostriata sp. nov., 62.4 mm SL, Holotype, TKPM-P17344; (F) C. minamorii minamorii sp. nov., 48.9 mm SL, Holotype, TKPM-P17345; (G) C. minamorii oumiensis subsp. nov., 51.7 mm SL, Holotype, TKPM-P17346; (H) C. minamorii tokaiensis subsp. nov., 47.9 mm SL, Holotype, MPM-FI1507; (I) C. minamorii saninensis subsp. nov., 62.1 mm SL, Holotype, TKPM-P17348; (J) C. minamorii yodoensis subsp. nov., 45.1 mm SL, Holotype, FAKU55719." pageId="13" pageNumber="116">Figs. 3C</figureCitation>
|
||
,
|
||
<figureCitation id="13597ECBFF9FFFACF0C550C7FC5AFE23" box="[901,936,476,500]" captionStart="FIGURE 4" captionStartId="8.[151,250,1868,1891]" captionTargetBox="[151,1436,1201,1847]" captionTargetId="figure-591@8.[151,1436,1201,1847]" captionTargetPageId="8" captionText="FIGURE 4. Color in life of adult male of 3 subspecies of Cobitis striata. (A–B): C. s. striata; (A) non-spawning season, Ima River system, Fukuoka Pref., 14 XII 2010; (B) spawning season, Aasahi River system, Okayama Pref., 17 VI 2007, collected by Koji Tominaga; (C–D): C. s. fuchigamii subsp. nov.; (C) non-spawning season, Onga River system, Fukuoka Pref., 14 XII 2010; (D) spawning season, same locality, 13 VI 2007; (E–F): C. s. hakataensis subsp. nov.; (E) non-spawning season, Tatara River system, Fukuoka Pref., 14 XII 2010; (F) spawning season, same locality, 23 V 2005." pageId="13" pageNumber="116">4E</figureCitation>
|
||
). Body yellowish white with dark brown pigmentation in fresh specimens. Clear streak running from the tip of snout to the occiput, crossing to the eye. Upper part of head, opercle and snout covered with oval or amorphous shape spots. Body pigmentation organized in one middorsal and four lateral zones. Line L1 consisting of a series of 14–16, saddles or oval-shaped blotches, irregularly chained to each other. Line L2 formed by longitudinal jagged line, reaching to middorsal region, often fused with L1. Line L3 formed by incomplete longitudinal line, reaching to caudal base. Line L4 formed by longitudinal jagged weblike line, reaching to postanal body, broader than L3. Line L5 organized in 11–14 blotches from upper part of the pectoral fin to the caudal-fin base; blotches roundish or ovoid. Caudal fin and dorsal fin with 3–4 arcuate bars. Anal fin pigmented along the fin rays. Upper spot at the caudal base jet-black comparable in size to eye diameter, lower spot at the caudal base faint or missing.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF387525FFE35FC73" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF387525FFDF9FC8B" box="[199,523,836,860]" italics="true" pageId="13" pageNumber="116">Male in the spawning season</emphasis>
|
||
(
|
||
<figureCitation id="13597ECBFF9FFFACF15A525FFD9DFC8B" box="[538,623,836,860]" captionStart="FIGURE 4" captionStartId="8.[151,250,1868,1891]" captionTargetBox="[151,1436,1201,1847]" captionTargetId="figure-591@8.[151,1436,1201,1847]" captionTargetPageId="8" captionText="FIGURE 4. Color in life of adult male of 3 subspecies of Cobitis striata. (A–B): C. s. striata; (A) non-spawning season, Ima River system, Fukuoka Pref., 14 XII 2010; (B) spawning season, Aasahi River system, Okayama Pref., 17 VI 2007, collected by Koji Tominaga; (C–D): C. s. fuchigamii subsp. nov.; (C) non-spawning season, Onga River system, Fukuoka Pref., 14 XII 2010; (D) spawning season, same locality, 13 VI 2007; (E–F): C. s. hakataensis subsp. nov.; (E) non-spawning season, Tatara River system, Fukuoka Pref., 14 XII 2010; (F) spawning season, same locality, 23 V 2005." pageId="13" pageNumber="116">Fig. 4F</figureCitation>
|
||
). Line L4 not visible or formed by faint longitudinal line, present only in anterior half of body. Lines L3 and L5 well developed with broad stripes from the upper part of the pectoral-fin base to the caudal-fin base.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF38752ABFBD6FC3B" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF38752ABFEE9FC1F" box="[199,283,944,968]" italics="true" pageId="13" pageNumber="116">Female</emphasis>
|
||
(
|
||
<figureCitation id="13597ECBFF9FFFACF26A52B4FE70FC10" box="[298,386,943,967]" captionStart="FIGURE 5" captionStartId="9.[151,250,1191,1214]" captionTargetBox="[151,1436,195,1170]" captionTargetId="figure-415@9.[151,1436,193,1170]" captionTargetPageId="9" captionText="FIGURE 5. Color in life of adult females: (A) C. striata striata, Yoshii River system, Okayama Pref., 7 VI 2010; (B) C. striata fuchigamii subsp. nov., Onga River system, Fukuoka Pref., 12 VI 2007; (C) C. striata hakataensis subsp. nov., Tatara River system, Fukuoka Pref., 13 VI 2007; (D) C. kaibarai sp. nov., Rokkaku River system, Saga Pref., 28 VI 2007; (E) C. magnostriata sp. nov., near Biwa Lake, Shiga Pref., 13 VI 2007; (F) C. minamorii minamorii sp. nov., Yoshii River system, Okayama Pref., 7 VI 2007; (G) C. minamorii oumiensis subsp. nov., near Biwa Lake, Shiga Pref., 17 VI 2007; (H) C. minamorii tokaiensis subsp. nov., Iride-ota River system, Shizuoka Pref., 13 VI 2007; (I) C. minamorii saninensis subsp. nov., Hii River system, Shimane Pref., 1. VI 2008; (J) C. minamorii yodoensis subsp. nov., Yodo River system, Kyoto Pref., VIII 1996, collected by Fumitoshi Mori, photographed by Ryu Uchiyama." pageId="13" pageNumber="116">Fig. 5C</figureCitation>
|
||
). Appearance similar to males in the non-spawning season, but number of blotches of line L5 tends to be more than in the male, line L5 of female organized in 11–17 blotches.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF38752ECFCABFBE3" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF38752ECFE44FBC7" bold="true" box="[199,438,1015,1040]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Sexual dimorphism.</emphasis>
|
||
Males have roundish lamina circularis at the base of the pectoral fin, but females do not. Generally, the body size of females is larger than that of males.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF387555BFA64FB80" blockId="13.[151,1436,151,1400]" box="[199,1430,1087,1112]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF387555BFE9EFB8F" bold="true" box="[199,364,1087,1112]" pageId="13" pageNumber="116">Egg diameter.</emphasis>
|
||
0.98 ±
|
||
<quantity id="4C9ACFABFF9FFFACF2825524FDD6FB80" box="[450,548,1087,1111]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="116" unit="mm" value="0.05">0.05 mm</quantity>
|
||
(females, N = 3; collected from the Tatara River system,
|
||
<collectingRegion id="49A6ACACFF9FFFACF7EA5524FA79FB80" box="[1194,1419,1087,1111]" country="Japan" name="Fukuoka" pageId="13" pageNumber="116">Fukuoka Prefecture</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF387557FFD46FBAB" blockId="13.[151,1436,151,1400]" box="[199,692,1124,1148]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF387557FFEBFFBAB" bold="true" box="[199,333,1124,1148]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Karyotype.</emphasis>
|
||
Diploid (
|
||
<bibRefCitation id="EFF31FBFFF9FFFACF2F8557FFD5BFBAB" author="Kitagawa, E. & Nakajima, J. & Hoshino, K. & Kitagawa, T." box="[440,681,1124,1148]" pageId="13" pageNumber="116" pagination="7 - 19" refId="ref24313" refString="Kitagawa, E., Nakajima, J., Hoshino, K. & Kitagawa, T. (2009) Geographic distribution and biogeographical origin of the spined loach genus Cobitis in north-eastern Kyushu Island, Japan. Japanese Journal of Ichthyology, 56, 7 - 19." type="journal article" year="2009">
|
||
Kitagawa
|
||
<emphasis id="B916BE5CFF9FFFACF16C557EFD97FBAB" box="[556,613,1124,1148]" italics="true" pageId="13" pageNumber="116">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37831C5FF9FFFACF387559CFE64FB30" pageId="13" pageNumber="116" type="distribution">
|
||
<paragraph id="8BDD624EFF9FFFACF387559CFA79FB48" blockId="13.[151,1436,151,1400]" box="[199,1419,1159,1184]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF387559CFE92FB77" bold="true" box="[199,352,1159,1184]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Distribution.</emphasis>
|
||
Rivers flowing into Hakata Bay, northern Kyushu:
|
||
<collectingRegion id="49A6ACACFF9FFFACF0E3559CFB8DFB48" box="[931,1151,1159,1183]" country="Japan" name="Fukuoka" pageId="13" pageNumber="116">Fukuoka Prefecture</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFF31FBFFF9FFFACF7CD559CFA8DFB48" author="Nakajima, J. & Onikura, N. & Kitagawa, E. & Kitagawa, T. & Oikawa, S." box="[1165,1407,1159,1184]" pageId="13" pageNumber="116" pagination="10 - 15" refId="ref24943" refString="Nakajima, J., Onikura, N., Kitagawa, E., Kitagawa, T. & Oikawa, S. (2008) Distribution pattern of Cobitis (Telostei: Cobitidae) in northern Kyushu Island, Japan. Folia Zoologica, 57, 10 - 15." type="journal article" year="2008">
|
||
Nakajima
|
||
<emphasis id="B916BE5CFF9FFFACF6435592FAC9FB77" box="[1283,1339,1160,1184]" italics="true" pageId="13" pageNumber="116">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF38755B0FE64FB30" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF38755B0FE4FFB13" bold="true" box="[199,445,1195,1220]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Habitat and biology.</emphasis>
|
||
This species inhabits sandy-mud bottoms of the middle and lower reach of rivers. Life histories are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37831C5FF9FFFACF38755EFFDE4FAF8" pageId="13" pageNumber="116" type="etymology">
|
||
<paragraph id="8BDD624EFF9FFFACF38755EFFDE4FAF8" blockId="13.[151,1436,151,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF38755EFFEBCFADB" bold="true" box="[199,334,1268,1292]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Etymology.</emphasis>
|
||
The subspecific name is derived from the popular common name of the
|
||
<collectingRegion id="49A6ACACFF9FFFACF7E655EFFAFAFADB" box="[1190,1288,1268,1292]" country="Japan" name="Fukuoka" pageId="13" pageNumber="116">Fukuoka</collectingRegion>
|
||
City area in which the
|
||
<typeStatus id="54D9DCECFF9FFFACF24B5403FEC9FAE7" box="[267,315,1304,1328]" pageId="13" pageNumber="116">type</typeStatus>
|
||
locality is situated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37831C5FF9FFFACF3875427FD5AFAA0" pageId="13" pageNumber="116" type="discussion">
|
||
<paragraph id="8BDD624EFF9FFFACF3875427FBDCFA83" blockId="13.[151,1436,151,1400]" box="[199,1070,1340,1364]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF3875427FEC9FA83" bold="true" box="[199,315,1340,1364]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Remarks.</emphasis>
|
||
The genetic features have been reported by
|
||
<bibRefCitation id="EFF31FBFFF9FFFACF06B5427FBD8FA83" author="Kitagawa, E. & Nakajima, J. & Hoshino, K. & Kitagawa, T." box="[811,1066,1340,1364]" pageId="13" pageNumber="116" pagination="7 - 19" refId="ref24313" refString="Kitagawa, E., Nakajima, J., Hoshino, K. & Kitagawa, T. (2009) Geographic distribution and biogeographical origin of the spined loach genus Cobitis in north-eastern Kyushu Island, Japan. Japanese Journal of Ichthyology, 56, 7 - 19." type="journal article" year="2009">
|
||
Kitagawa
|
||
<emphasis id="B916BE5CFF9FFFACF0DD5426FC27FA83" box="[925,981,1340,1364]" italics="true" pageId="13" pageNumber="116">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDD624EFF9FFFACF387547BFD5AFAA0" blockId="13.[151,1436,151,1400]" box="[199,680,1375,1400]" pageId="13" pageNumber="116">
|
||
<emphasis id="B916BE5CFF9FFFACF387547BFE71FAAF" bold="true" box="[199,387,1376,1400]" inLineHeading="true" pageId="13" pageNumber="116" reason="1">Japanese name.</emphasis>
|
||
Hakata-suji-shima-dojyô.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |